Systematics of Neotropical Butterflies at the McGuire Center for Lepidoptera and Biodiversity, Florida Museum of Natural History

Publications


Books

2003. Willmott, K. R. The Genus Adelpha: its Systematics, Biology and Biogeography (Lepidoptera: Nymphalidae: Limenitidini). Gainesville, Scientific Publishers. viii + 322 pp. Download pre-print [PDF] - Archivos [PDF]

Research papers

XXX. Chazot, N., Willmott, K. R., Paola G., S. E., Toporov, A., Hill, R., Jiggins, C., Elias, M. Filtering by elevation and mutualistic mimicry shape the structure of Andean butterfly communities. American Naturalist, in press.

XXX. Checa, M. F., Rodríguez, J., Liger, B., Willmott, K. R. Microhabitat significantly affects the composition, abundance and phenology of butterfly assemblages in an Ecuadorian dry forest. Florida Entomologist, in press.

2013. Willmott, K. R., Hall, J. P. W. A new species and two new subspecies of Adelpha Hübner, [1819] from the tropical Andes (Lepidoptera: Limenitidinae). Journal of the Lepidopterists’ Society, 67: 241-252. Download reprint [PDF]

2013. Radford, J., Willmott, K. R. A distinctive new subspecies of Catasticta poujadei from the Cordillera del Cóndor in eastern Ecuador (Lepidoptera: Pieridae: Pierinae). Tropical Lepidoptera Research, 23: 86-91. Download reprint [PDF]

2013. Pyrcz, T. W., Willmott, K. R., Garlacz, R., Boyer, P. Latitudinal gradient and spatial covariance in species richness of tropical Lepidoptera in the Andes. Insect Diversity and Conservation, Online version: DOI: 10.1111/icad.12058.

2013. Ortíz-Acevedo, E., Willmott, K. R. Molecular systematics of the butterfly tribe Preponini (Nymphalidae: Charaxinae). Systematic Entomology. Online version: DOI: 10.1111/syen.12008.

2013. Giraldo, C. E., Willmott, K. R., Vila, R. Uribe, S. I. Ithomiini butterflies (Lepidoptera: Nymphalidae) of Antioquia, Colombia. Neotropical Entomology. Online version: DOI: 10.1007/s13744-012-0102-4.

2012. Matos-Maraví, P. F., Peña, C., Willmott, K. R., Freitas, A. V. L., Wahlberg, N. Systematics and evolutionary history of butterflies in the "Taygetis clade" (Nymphalidae: Satyrinae: Euptychiina): towards a better understanding of Neotropical biogeography. Molecular Phylogenetics and Evolution, 66: 54-68. Online version: DOI: 10.1016/j.ympev.2012.09.005

2012. Hill, R. I., Elias, M., Dasmahapatra, K. K., Jiggins, C. D., Koong, V., Willmott, K. R., Mallet, J. Ecologically relevant cryptic species in the highly polymorphic Amazonian butterfly Mechanitis mazaeus s.l. (Lepidoptera: Nymphalidae; Ithomiini). Biological Journal of the Linnean Society, 106: 540-560. Online version: DOI: 10.1111/j.1095-8312.2012.01874.x

2012. Rosser, N., Phillimore, A.B., Huertas, B., Willmott, K.R., Mallet, J. Testing historical explanations for gradients in species richness in heliconiine butterflies of tropical America. Biological Journal of the Linnean Society, 105: 479-497. Online version: DOI: 10.1111/j.1095-8312.2011.01814.x.

2011. Willmott, K. R., Elias, M., Sourakov, A. Two possible caterpillar mimicry complexes in neotropical danaine butterflies (Nymphalidae, Danainae: Danaini, Ithomiini). Annals of the Entomological Society of America, 104: 1108-1118. Abstract.

2011. Gompert, Z., Willmott, K. R., Elias, M. Heterogeneity in predator micro-habitat use and the maintenance of Müllerian mimicry diversity. Journal of Theoretical Biology, 281: 39-46. Online version.

2011. Bolaños M., I. A., Zambrano G., G., Willmott, K. R. Descripción de los estados inmaduros de Pteronymia zerlina zerlina, P. zerlina machay, P. medellina y P. veia n. ssp. (Lepidoptera: Nymphalidae: Ithomiini) de Colombia y del Ecuador. Tropical Lepidoptera Research, 21: 27-33. Download reprint [PDF]

2011. Pyrcz, T., Greeney, H. F., Willmott, K. R., Wojtusiak, J. A taxonomic revision of the genus Daedalma Hewitson with the descriptions of twenty new taxa and the immature stages of two species (Lepidoptera: Nymphalidae: Satyrinae). Zootaxa, 2898: 1-68. Online version

2010. Mullen, S. P., Savage, W., Willmott, K. R. Rapid diversification and not clade age explains high diversity in neotropical Adelpha butterflies. Proceedings of the Royal Society of London B doi:10.1098/rspb.2010.2140 Online version

2010. Hall, J. P. W., Willmott, K. R. Description of a new Lucillella species (Riodinidae: Symmachiini) discovered in the eastern Andes of Ecuador using the single rope canopy access technique. Journal of the Lepidopterists' Society, 64: 139-146. Download high (12 Mb) or low (4.7 Mb) resolution reprint [PDF].

2010. Willmott, K. R., Hall, J. P. W. A new species of Dynamine Hübner, [1819] from northwestern Ecuador (Lepidoptera: Nymphalidae: Biblidinae). Tropical Lepidoptera Research, 20: 23-27. Download reprint [PDF]

2010. de-Silva, D. L., Day, J. J., Elias, M., Willmott, K. R., Whinnett, A., Mallet, J. Molecular phylogenetics of the neotropical butterfly subtribe Oleriina (Nymphalidae: Danainae: Ithomiini). Molecular Phylogenetics and Evolution, 55:1032-1041. Download reprint [PDF]

2009. Janzen, D. J., Hallwachs, W., + 39 authors, Willmott, K. R., + 3 authors. Integration of DNA barcoding into an ongoing inventory of complex tropical biodiversity. Molecular Ecology Resources, 9(Suppl. 1), 1-26. Online version

2009. Elias, M., Gompert, Z., Jiggins, C., Willmott, K. R. Phylogenetic community ecology needs to take positive interactions into account: Insights from colourful butterflies. Communicative & Integrative Biology 2(2): 113-116 [Article addendum]. Download reprint [PDF]

2009. Hall, J. P. W., and K. R. Willmott. Two new species of Euselasia (Riodinidae: Euselasiinae) from western Ecuador. Tropical Lepidoptera Research, 19(1): 52-55. Download reprint [PDF]

2009. Elias, M., Joron, M., Willmott, K. R., Kaiser, V., Silva-Brandão, K. L., Freitas, A. V. L., Arias, C. F., Gomez Piñeres, L. M., Uribe, S., Jiggins, C. Out of the Andes: patterns of diversification in clearwing butterflies. Molecular Ecology, 18: 1716-1729. Download page proofs [PDF]

2009. Willmott, K. R., Freitas, A. V. L., Hall, J. P. W., Silva-Brandão, K. L., Paluch, M. A new species of Actinote Hübner from the eastern Andes of Ecuador (Lepidoptera: Nymphalidae: Heliconiinae). Proceedings of the Entomological Society of Washington, 111(1): 47-56. Download reprint [PDF] - NOTE: figures in published version are monochrome

2008. Elias, M., Gompert, Z., Jiggins, C., Willmott, K. R. Mutualistic interations drive ecological niche convergence in a diverse butterfly community. PLoS Biology, 6(12):e300. Definitive online version or Download reprint [PDF], Supporting data S1 [PDF], Supporting data S2 [PDF]

2008. Willmott, K. R., Vitale, F. Taxonomic notes on Napeogenes (Lepidoptera: Ithomiinae) from Ecuador and Colombia, with the description of ten new subspecies. Genus, 19(3): 585-609. Download reprint [PDF]

2008. Willmott, K. R., Lamas, G. A revision of the genus Megoleria (Lepidoptera, Nymphalidae, Ithomiinae). Tropical Lepidoptera Research, 18(1): 46-59. Download reprint [PDF]

2007. Elias, M., Hill, R., Dasmahapatra, K., Willmott, K. R., Brower, A., Mallet, J., and C. Jiggins. Limited performance of DNA barcoding in a diverse community of tropical butterflies. Proceedings of the Royal Society of London B, 274: 2881-2889. Download reprint [PDF] and suppplementary documentation

2007. Brehm, G., Hartmann, T., and K. R. Willmott. Pyrrolizidine alkaloids and pharmacophagous Lepidoptera visitors of Prestonia amabilis (Apocynaceae) in a montane rain forest in Ecuador. Annals of the Missouri Botanical Garden, 94(2): 463-473. Download reprint [PDF]

2007. Hall, J. P. W., and K. R. Willmott. Four new species of Symmachiini from Ecuador (Lepidoptera: Riodinidae). Tropical Lepidoptera, 16(1-2): 1-5. Download reprint [PDF]

2007. Willmott, K. R., and G. Lamas. A revision of Pachacutia, a new genus of rare Andean ithomiine butterflies (Nymphalidae: Ithomiinae), with the description of two new species. Annals of the Entomological Society of America, 100: 449-469. Download reprint [PDF]

2006. Pyrcz, T. W., Willmott, K. R., Hall, J. P. W., and A. L. Viloria. A review of the genus Manerebia Staudinger (Lepidoptera: Nymphalidae: Satyrinae) in the northern Andes. Journal of Research on the Lepidoptera, 39: 37-79 (2000). Download reprint [PDF]

2006. Willmott, K. R., and A. V. L. Freitas. Higher-level phylogeny of the Ithomiinae (Lepidoptera: Nymphalidae): classification, patterns of larval hostplant colonisation and diversification. Cladistics, 22: 297-368. Online version, Download PAGE PROOFS [PDF] or e-mail authors for reprint

2006. Jiggins, C. D., Mallarino, R., Willmott, K. R., and E. Bermingham. The phylogenetic pattern of speciation and wing pattern change in neotropical Ithomia butterflies (Lepidoptera: Nymphalidae). Evolution, 60: 1454-1466. Download reprint [PDF]

2006. Willmott, K. R., and G. Lamas. A phylogenetic reassessment of Hyalenna Forbes and Dircenna Doubleday, with a revision of Hyalenna (Lepidoptera: Nymphalidae: Ithomiinae). Systematic Entomology, 31(3): 419-468. Download reprint [PDF]

2006. Brower, A. V. Z., Freitas, A. V. L., Lee, M.-M., Silva Brandão, K. L., Whinnett, A., and K. R. Willmott. Phylogenetic relationships among the Ithomiini (Lepidoptera: Nymphalidae) inferred from one mitochondrial and two nuclear gene regions. Systematic Entomology, 31: 288-301. Download reprint [PDF]

2005. Whinnett, A., Willmott, K. R., Brower, A.V.Z., Simpson, F., Zimmermann, M., Lamas, G., and J. Mallet. Mitochondrial DNA provides an insight into the mechanisms driving diversification in the ithomiine butterfly Hyposcada anchiala (Lepidoptera: Nymphalidae, Ithomiinae). European Journal of Entomology, 102: 633-639. Download reprint [PDF]

2005. Whinnett, A., Zimmermann, M., Willmott, K. R., Herrera, N., Mallarino, R., Simpson, F., Joron, M., Lamas, G., and J. Mallet. Strikingly variable divergence times inferred across an Amazonian butterfly 'suture zone'. Proceedings of the Royal Society of London B, 272: 2525-2533. Download reprint [PDF]

2005. Whinnett, A., Brower, A. V. Z., Lee, M.-M., Willmott, K. R., and J. Mallet. The phylogenetic utility of tektin, a novel region for inferring systematic relationships amongst Lepidoptera. Annals of the Entomological Society of America, 98: 873-886. Download reprint [PDF]

2005. Mallarino, R., Bermingham, E., Willmott, K. R., Whinnett, A., and C. D. Jiggins. Molecular systematics of the butterfly genus Ithomia (Lepidoptera: Ithomiinae): a composite phylogenetic hypothesis based on seven genes. Molecular Phylogenetics and Evolution, 34(3): 625-644. Download reprint [PDF]

2005. Hall, J. P. W., K. R. Willmott and R. C. Busby. Five new Penaincisalia species (Lepidoptera: Lycaenidae: Eumaeini) from the Andes of southern Ecuador and northern Peru. Zootaxa, 797: 1-20. Download reprint [PDF]

2004. Balcázar-Lara, M., and K. R. Willmott. A new subspecies of Adelpha erymanthis from Mexico, with a key to identification of similar taxa (Lepidoptera: Nymphalidae: Biblidinae). Tropical Lepidoptera, 12(1-2)("2001"): 25-38. Download reprint [PDF]

2004. Willmott, K. R., and J. Mallet. Correlations between adult mimicry and larval hostplants in ithomiine butterflies. Proceedings of the Royal Society of London B (Suppl.) 271: S266-S269. Download reprint [PDF]

2004. Willmott, K. R., and Lamas, G. A new species of Tithorea (Lepidoptera: Nymphalidae: Ithomiinae) from the Chocó region of northwestern South America and Panama. J. New York Ent. Soc., 112(1): 1-17. Download reprint [PDF]

2004. Willmott, K. R., and J. P. W. Hall. Taxonomic notes on the genus Zaretis, with the description of a new species (Lepidoptera: Nymphalidae: Charaxinae). Tropical Lepidoptera, 12(1-2): 25-38. Download reprint [PDF]

2003. Mallet, J. L. B., and K. R. Willmott. Taxonomy: renaissance or Tower of Babel? Trends in Ecology and Evolution, 18(2): 57-59. Online version

2003. Willmott, K. R. Cladistic analysis of the neotropical butterfly genus Adelpha (Nymphalidae: Limenitidini), with comments on the subtribal classification of the tribe Limenitidini. Systematic Entomology, 28: 1-43. Download reprint [PDF]

2002. Robinson-Dean, J. C., K. R. Willmott, M. J. Catterall, D. J. Kelly, A. Whittington, B. Phalan, N. M. Marples, and D. R. S. Boeadi. A new subspecies of Red-backed Thrush Zoothera erythronota kabaena subsp. nov. (Muscicapidae: Turdidae), from Kabaena island, Indonesia. Forktail, 18: 1-10.

2001. Willmott, K. R., J. P. W. Hall, and G. Lamas. Systematics of Hypanartia (Lepidoptera: Nymphalidae: Nymphalinae), with a test for geographical speciation mechanisms in the Andes. Systematic Entomology, 26: 369-399. Download reprint [PDF]

2001. Willmott, K. R., L. M. Constantino, and J. P. W. Hall. A review of Colobura (Lepidoptera: Nymphalidae) with comments on larval and adult ecology and description of a sibling species. Annals of the Entomological Society of America, 94(2): 185-196. Download reprint [PDF]

2000. Hall, J. P. W., and K. R. Willmott. Patterns of feeding behaviour in adult male riodinid butterflies and their relationship to morphology and ecology. Biol. J. Linn. Soc., 69: 1-23. Download reprint [PDF]

1999. Pyrcz, T., K. R. Willmott, and J. P. W. Hall. Contribution to the knowledge of Ecuadorian Pronophilini. Part III. Three new species and five new subspecies of Lymanopoda (Lepidoptera: Nymphalidae: Satyrinae). Genus, 10(3): 497-522. Download reprint [PDF - 1] [PDF - 2]

1999. Willmott, K. R., and J. P. W. Hall. Taxonomic notes on Ecuadorian Adelpha with the description of two new species and seven new subspecies (Lepidoptera: Nymphalidae: Limenitidinae). Tropical Lepidoptera, 10(1): 1-17. Download reprint [PDF - 1] [PDF - 2]

1998. Hall, J. P. W., and K. R. Willmott. Four new riodinid species from eastern Ecuador (Lepidoptera: Riodinidae). Lambillionea, 98: 325-334. Download reprint [PDF]

1998. Hall, J. P. W., and K. R. Willmott. Nine new species and one new subspecies of Euselasia from Ecuador (Lepidoptera: Riodinidae). Tropical Lepidoptera, Suppl. 1, 9(2): 27-35. Download reprint [PDF]

1998. Hall, J. P. W., and K. R. Willmott. Three new species of Riodinini from the cloud forests of Ecuador (Lepidoptera: Riodinidae). Tropical Lepidoptera, Suppl. 1, 9(2): 22-26. Download reprint [PDF]

1996. Hall, J. P. W., and K. R. Willmott. The genus Theope: four new species and a new subspecies (Lepidoptera: Riodinidae). Tropical Lepidoptera, 7(1): 63-67. Download reprint [PDF]

1996. Hall, J. P. W., and K. R. Willmott. Notes on the genus Argyrogrammana Part II, with the description of a new species. Tropical Lepidoptera, 7(1): 71-80. Download reprint [PDF]

1996. Hall, J. P. W., and K. R. Willmott. Systematics of the riodinid tribe Symmachiini, with the description of a new genus and five new species from Ecuador, Venezuela and Brazil (Lepidoptera: Riodinidae). Lambillionea, 96: 637-660. Download reprint [PDF]

1996. Torres, R., J. P. W. Hall, K. R. Willmott, and K. Johnson. A new genus of brilliantly colored elfin butterflies from the high Andes (Lepidoptera: Lycaenidae). Tropical Lepidoptera, 7(1): 81-86. Download reprint [PDF]

1995. Hall, J. P. W., and K. R. Willmott. Two new species of Mesene from western Ecuador (Lepidoptera: Riodinidae). Tropical Lepidoptera, 6(2): 110-112. Download reprint [PDF]

1995. Hall, J. P. W., and K. R. Willmott. Five new species and a new genus of riodinid from the cloud forests of eastern Ecuador (Lepidoptera: Riodinidae). Tropical Lepidoptera, 6(2): 131-135. Download reprint [PDF]

1995. Hall, J. P. W., and K. R. Willmott. Notes on the genus Argyrogrammana, with descriptions of five new species (Lepidoptera: Riodinidae). Tropical Lepidoptera, 6(2): 136-143. Download reprint [PDF]

1995. Willmott, K. R., and J. P. W. Hall. Two new species of satyrines from Ecuador (Nymphalidae: Satyrinae). Tropical Lepidoptera, 6(2): 103-105. Download reprint [PDF]

1995. Willmott, K. R., and J. P. W. Hall. Two new species of Adelpha from eastern Ecuador (Lepidoptera: Nymphalidae). Tropical Lepidoptera, 6(2): 106-109. Download reprint [PDF]

1994. Willmott, K. R., and J. P. W. Hall. Four new species of Riodinidae (Lepidoptera) from Western Ecuador. Tropical Lepidoptera, 5(2): 87-91. Download reprint [PDF]

1993. Hall, J. P. W., and K. R. Willmott. Temporal and elevational additions of distribution for Costa Rican butterflies. Tropical Lepidoptera, 4(2): 139-141.

Book chapters

2006. Gareca, Y., Forno, E., Pyrcz, T., Willmott, K. R., Reichle, S. Lista preliminar de mariposas diurnas de Bolivia. In: Gareca, Y., and S. Reichle (eds.), Mariposas Diurnas de Bolivia, pp. 1-66. Tarija, Bolivia, PROMETA.

2004. Willmott, K.R. Brush-footed butterflies (Lepidoptera: Nymphalidae). In: Capinera, J. L. (Ed.), Encyclopedia of Entomology, Volume I A-E, pp. 379-384. Dordrecht / Boston / London, Kluwer Academic Publishers. Download reprint [PDF]

Popular publications

2004. Willmott, K. R., and J. P. W. Hall. Common Butterflies of Yuturi and Yarina lodge. Yuturi Lodge, Ecuador. 2 pp.

2001. Hall, J. P. W., and K. R. Willmott. Ecuador. Out on a limb. National Geographic Magazine (Update from the field), 199(2): 86. Download reprint [PDF]

2001. Willmott, K. R., and J. P. W. Hall. Winged gems of Ecuador. Biodiversity paradise. National Geographic Magazine (Geographica) 199(5): xxviii. Download reprint [PDF]

Letters and reviews

2012. Willmott, K. R. Lepidoptera. Borboletas e Mariposas do Brasil. [Lepidoptera. Butterflies and Moths of Brazil.] By de Almeida, A. C., Freitas, A. V. L. (2012). Tropical Lepidoptera Research 22(2): 120. [book review]

2009. Willmott, K. R. Mariposas de Canandé. Sus Amenazas, Potencial y Futuro. [The Butterflies of Canandé. Their Threats, Potential and Future.] By Checa V., M. F. (2009). Tropical Lepidoptera Research 19(1): 60. [book review]

2008. Willmott, K. R. Catalogue of the Hostplants of the Neotropical Butterflies/ Catálogo de las Plantas Huésped de las Mariposas Neotropicales. By Beccaloni, G. W., Viloria, A. L., Hall, S. K. & Robinson, G. S. (2008). Tropical Lepidoptera Research 18(2): 124-125. [book review]

1998. Willmott, K. R. A fair review? News of the Lepidopterist's Society, 40(4): 92-93.

1997. Hall, J. P. W., and K. R. Willmott. The Butterflies of Venezuela. Part I. Nymphalidae I (Limenitidinae, Apaturinae, Charaxinae). A Comprehensive Guide to the Identification of Adult Nymphalidae, Ppailionidae and Pieridae. By Andrew F. E. Neild. Tropical Lepidoptera, 8(1): 46. [book review]