Publications Books
2003. Willmott, K. R. The Genus Adelpha: its Systematics, Biology and Biogeography (Lepidoptera: Nymphalidae: Limenitidini). Gainesville, Scientific Publishers. viii + 322 pp. Download pre-print [PDF] - Archivos [PDF] Research papers
2007. Elias, M., Hill, R., Dasmahapatra, K., Willmott, K. R., Brower, A., Mallet, J., and C. Jiggins. Limited performance of DNA barcoding in a diverse community of tropical butterflies. Proceedings of the Royal Society of London B, in press. Book chapters
2006. Gareca, Y., Forno, E., Pyrcz, T., Willmott, K. R., Reichle, S. Lista preliminar de mariposas diurnas de Bolivia. In: Gareca, Y., and S. Reichle (eds.), Mariposas Diurnas de Bolivia, pp. 1-66. Tarija, Bolivia, PROMETA. Popular publications
2005. Willmott, K. R., and J. P. W. Hall. Common Butterflies of the Ecuadorian Amazon. Museo Ecuatoriano de Ciencias Naturales, Quito. 2 pp. [in press] Letters and reviews
1998. Willmott, K. R. A fair review? News of the Lepidopterist's Society, 40(4): 92-93.
2007. Brehm, G., Hartmann, T., and K. R. Willmott. Pyrrolizidine alkaloids and pharmacophagous Lepidoptera visitors of Prestonia amabilis (Apocynaceae) in a montane rain forest in Ecuador. Annals of the Missouri Botanical Garden, 94(2): 463-473. Download reprint [PDF]
2007. Hall, J. P. W., and K. R. Willmott. Three new species of Symmachiini from Ecuador (Lepidoptera: Riodinidae). Tropical Lepidoptera, 16(1-2): 1-5. Download reprint [PDF]
2007. Willmott, K. R., and G. Lamas. A revision of Pachacutia, a new genus of rare Andean ithomiine butterflies (Nymphalidae: Ithomiinae), with the description of two new species. Annals of the Entomological Society of America, 100: 449-469. Please e-mail one of the authors for a reprint.
2006. Pyrcz, T. W., Willmott, K. R., Hall, J. P. W., and A. L. Viloria. A review of the genus Manerebia Staudinger (Lepidoptera: Nymphalidae: Satyrinae) in the northern Andes. Journal of Research on the Lepidoptera, 39: 37-79 (2000). Download reprint [PDF]
2006. Willmott, K. R., and A. V. L. Freitas. Higher-level phylogeny of the Ithomiinae (Lepidoptera: Nymphalidae): classification, patterns of larval hostplant colonisation and diversification. Cladistics, 22: 297-368. Please e-mail one of the authors for a reprint.
2006. Jiggins, C. D., Mallarino, R., Willmott, K. R., and E. Bermingham. The phylogenetic pattern of speciation and wing pattern change in neotropical Ithomia butterflies (Lepidoptera: Nymphalidae). Evolution, 60: 1454-1466. Download reprint [PDF]
2006. Willmott, K. R., and G. Lamas. A phylogenetic reassessment of Hyalenna Forbes and Dircenna Doubleday, with a revision of Hyalenna (Lepidoptera: Nymphalidae: Ithomiinae). Systematic Entomology, 31(3): 419-468. Download reprint [PDF]
2006. Brower, A. V. Z., Freitas, A. V. L., Lee, M.-M., Silva Brandão, K. L., Whinnett, A., and K. R. Willmott. Phylogenetic relationships among the Ithomiini (Lepidoptera: Nymphalidae) inferred from one mitochondrial and two nuclear gene regions. Systematic Entomology, 31: 288-301. Download reprint [PDF]
2005. Whinnett, A., Willmott, K. R., Brower, A.V.Z., Simpson, F., Zimmermann, M., Lamas, G., and J. Mallet. Mitochondrial DNA provides an insight into the mechanisms driving diversification in the ithomiine butterfly Hyposcada anchiala (Lepidoptera: Nymphalidae, Ithomiinae). European Journal of Entomology, 102: 633-639. Download reprint [PDF]
2005. Whinnett, A., Zimmermann, M., Willmott, K. R., Herrera, N., Mallarino, R., Simpson, F., Joron, M., Lamas, G., and J. Mallet. Strikingly variable divergence times inferred across an Amazonian butterfly 'suture zone'. Proceedings of the Royal Society of London B, 272: 2525-2533. Download reprint [PDF]
2005. Whinnett, A., Brower, A. V. Z., Lee, M.-M., Willmott, K. R., and J. Mallet. The phylogenetic utility of tektin, a novel region for inferring systematic relationships amongst Lepidoptera. Annals of the Entomological Society of America, 98: 873-886. Download reprint [PDF]
2005. Mallarino, R., Bermingham, E., Willmott, K. R., Whinnett, A., and C. D. Jiggins. Molecular systematics of the butterfly genus Ithomia (Lepidoptera: Ithomiinae): a composite phylogenetic hypothesis based on seven genes. Molecular Phylogenetics and Evolution, 34(3): 625-644. Download reprint [PDF]
2005. Hall, J. P. W., K. R. Willmott and R. C. Busby. Five new Penaincisalia species (Lepidoptera: Lycaenidae: Eumaeini) from the Andes of southern Ecuador and northern Peru. Zootaxa, 797: 1-20. Download reprint [PDF]
2004. Balcázar-Lara, M., and K. R. Willmott. A new subspecies of Adelpha erymanthis from Mexico, with a key to identification of similar taxa (Lepidoptera: Nymphalidae: Biblidinae). Tropical Lepidoptera, 12(1-2)("2001"): 25-38. Download reprint [PDF]
2004. Willmott, K. R., and J. Mallet. Correlations between adult mimicry and larval hostplants in ithomiine butterflies. Proceedings of the Royal Society of London B (Suppl.) 271: S266-S269. Download reprint [PDF]
2004. Willmott, K. R., and Lamas, G. A new species of Tithorea (Lepidoptera: Nymphalidae: Ithomiinae) from the Chocó region of northwestern South America and Panama. J. New York Ent. Soc., 112(1): 1-17. Download reprint [PDF]
2004. Willmott, K. R., and J. P. W. Hall. Taxonomic notes on the genus Zaretis, with the description of a new species (Lepidoptera: Nymphalidae: Charaxinae). Tropical Lepidoptera, 12(1-2): 25-38. Download reprint [PDF]
2003. Mallet, J. L. B., and K. R. Willmott. Taxonomy: renaissance or Tower of Babel? Trends in Ecology and Evolution, 18(2): 57-59.
2003. Willmott, K. R. Cladistic analysis of the neotropical butterfly genus Adelpha (Nymphalidae: Limenitidini), with comments on the subtribal classification of the tribe Limenitidini. Systematic Entomology, 28: 1-43. Download reprint [PDF]
2002. Robinson-Dean, J. C., K. R. Willmott, M. J. Catterall, D. J. Kelly, A. Whittington, B. Phalan, N. M. Marples, and D. R. S. Boeadi. A new subspecies of Red-backed Thrush Zoothera erythronota kabaena subsp. nov. (Muscicapidae: Turdidae), from Kabaena island, Indonesia. Forktail, 18: 1-10.
2001. Willmott, K. R., J. P. W. Hall, and G. Lamas. Systematics of Hypanartia (Lepidoptera: Nymphalidae: Nymphalinae), with a test for geographical speciation mechanisms in the Andes. Systematic Entomology, 26: 369-399. Download reprint [PDF]
2001. Willmott, K. R., L. M. Constantino, and J. P. W. Hall. A review of Colobura (Lepidoptera: Nymphalidae) with comments on larval and adult ecology and description of a sibling species. Annals of the Entomological Society of America, 94(2): 185-196. Download reprint [PDF]
2000. Hall, J. P. W., and K. R. Willmott. Patterns of feeding behaviour in adult male riodinid butterflies and their relationship to morphology and ecology. Biol. J. Linn. Soc., 69: 1-23. Download reprint [PDF]
1999. Pyrcz, T., K. R. Willmott, and J. P. W. Hall. Contribution to the knowledge of Ecuadorian Pronophilini. Part III. Three new species and five new subspecies of Lymanopoda (Lepidoptera: Nymphalidae: Satyrinae). Genus, 10(3): 497-522. Download reprint [PDF - 1] [PDF - 2]
1999. Willmott, K. R., and J. P. W. Hall. Taxonomic notes on Ecuadorian Adelpha with the description of two new species and seven new subspecies (Lepidoptera: Nymphalidae: Limenitidinae). Tropical Lepidoptera, 10(1): 1-17. Download reprint [PDF - 1] [PDF - 2]
1998. Hall, J. P. W., and K. R. Willmott. Four new riodinid species from eastern Ecuador (Lepidoptera: Riodinidae). Lambillionea, 98: 325-334. Download reprint [PDF]
1998. Hall, J. P. W., and K. R. Willmott. Nine new species and one new subspecies of Euselasia from Ecuador (Lepidoptera: Riodinidae). Tropical Lepidoptera, Suppl. 1, 9(2): 27-35. Download reprint [PDF]
1998. Hall, J. P. W., and K. R. Willmott. Three new species of Riodinini from the cloud forests of Ecuador (Lepidoptera: Riodinidae). Tropical Lepidoptera, Suppl. 1, 9(2): 22-26. Download reprint [PDF]
1996. Hall, J. P. W., and K. R. Willmott. The genus Theope: four new species and a new subspecies (Lepidoptera: Riodinidae). Tropical Lepidoptera, 7(1): 63-67. Download reprint [PDF]
1996. Hall, J. P. W., and K. R. Willmott. Notes on the genus Argyrogrammana Part II, with the description of a new species. Tropical Lepidoptera, 7(1): 71-80. Download reprint [PDF]
1996. Hall, J. P. W., and K. R. Willmott. Systematics of the riodinid tribe Symmachiini, with the description of a new genus and five new species from Ecuador, Venezuela and Brazil (Lepidoptera: Riodinidae). Lambillionea, 96: 637-660. Download reprint [PDF]
1996. Torres, R., J. P. W. Hall, K. R. Willmott, and K. Johnson. A new genus of brilliantly colored elfin butterflies from the high Andes (Lepidoptera: Lycaenidae). Tropical Lepidoptera, 7(1): 81-86. Download reprint [PDF]
1995. Hall, J. P. W., and K. R. Willmott. Two new species of Mesene from western Ecuador (Lepidoptera: Riodinidae). Tropical Lepidoptera, 6(2): 110-112. Download reprint [PDF]
1995. Hall, J. P. W., and K. R. Willmott. Five new species and a new genus of riodinid from the cloud forests of eastern Ecuador (Lepidoptera: Riodinidae). Tropical Lepidoptera, 6(2): 131-135. Download reprint [PDF]
1995. Hall, J. P. W., and K. R. Willmott. Notes on the genus Argyrogrammana, with descriptions of five new species (Lepidoptera: Riodinidae). Tropical Lepidoptera, 6(2): 136-143. Download reprint [PDF]
1995. Willmott, K. R., and J. P. W. Hall. Two new species of satyrines from Ecuador (Nymphalidae: Satyrinae). Tropical Lepidoptera, 6(2): 103-105. Download reprint [PDF]
1995. Willmott, K. R., and J. P. W. Hall. Two new species of Adelpha from eastern Ecuador (Lepidoptera: Nymphalidae). Tropical Lepidoptera, 6(2): 106-109. Download reprint [PDF]
1994. Willmott, K. R., and J. P. W. Hall. Four new species of Riodinidae (Lepidoptera) from Western Ecuador. Tropical Lepidoptera, 5(2): 87-91. Download reprint [PDF]
1993. Hall, J. P. W., and K. R. Willmott. Temporal and elevational additions of distribution for Costa Rican butterflies. Tropical Lepidoptera, 4(2): 139-141.
2004. Willmott, K.R. Brush-footed butterflies (Lepidoptera: Nymphalidae). In: Capinera, J. L. (Ed.), Encyclopedia of Entomology, Volume I A-E, pp. 379-384. Dordrecht / Boston / London, Kluwer Academic Publishers. Download reprint [PDF]
2004. Willmott, K. R., and J. P. W. Hall. Common Butterflies of Yuturi and Yarina lodge. Yuturi Lodge, Ecuador. 2 pp.
2001. Hall, J. P. W., and K. R. Willmott. Ecuador. Out on a limb. National Geographic Magazine (Update from the field), 199(2): 86. Download reprint [PDF]
2001. Willmott, K. R., and J. P. W. Hall. Winged gems of Ecuador. Biodiversity paradise. National Geographic Magazine (Geographica) 199(5): xxviii. Download reprint [PDF]
1997. Hall, J. P. W., and K. R. Willmott. The Butterflies of Venezuela. Part I. Nymphalidae I (Limenitidinae, Apaturinae, Charaxinae). A Comprehensive Guide to the Identification of Adult Nymphalidae, Ppailionidae and Pieridae. by Andrew F. E. Neild. Tropical Lepidoptera, 8(1): 46. [book review]
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