Publications Books
2003. Willmott, K. R. The Genus Adelpha: its Systematics, Biology and Biogeography (Lepidoptera: Nymphalidae: Limenitidini). Gainesville, Scientific Publishers. viii + 322 pp. Download pre-print [PDF] - Archivos [PDF] Research papers
XXX. Gompert, Z., Willmott, K. R., Elias, M. Heterogeneity in predator micro-habitat use and the maintenance of Müllerian mimicry diversity. Submitted to Evolution B.
XXX. Hall, J. P. W., Willmott, K. R. Discovery of a new Lucillella species (Riodinidae: Symmachiini) in the eastern Andes of Ecuador using the single rope canopy access technique. Journal of the Lepidopterists' Society, in review.
2009. Janzen, D. J., Hallwachs, W., + 39 authors, Willmott, K. R., + 3 authors. Integration of DNA barcoding into an ongoing inventory of complex tropical biodiversity. Molecular Ecology Resources, 9(Suppl. 1), 1-26.
2009. Elias, M., Gompert, Z., Jiggins, C., Willmott, K. R. Phylogenetic community ecology needs to take positive interactions into account: Insights from colourful butterflies. Communicative & Integrative Biology 2(2): 113-116 [Article addendum]. Download reprint [PDF]
2009. Willmott, K. R., Freitas, A. V. L., Hall, J. P. W., Silva-Brandão, K. L., Paluch, M. A new species of Actinote Hübner from the eastern Andes of Ecuador (Lepidoptera: Nymphalidae: Heliconiinae). Proceedings of the Entomological Society of Washington, 111(1): 47-56. Download reprint [PDF] - NOTE: figures in published version are monochrome Book chapters
2006. Gareca, Y., Forno, E., Pyrcz, T., Willmott, K. R., Reichle, S. Lista preliminar de mariposas diurnas de Bolivia. In: Gareca, Y., and S. Reichle (eds.), Mariposas Diurnas de Bolivia, pp. 1-66. Tarija, Bolivia, PROMETA. Popular publications
2005. Willmott, K. R., and J. P. W. Hall. Common Butterflies of the Ecuadorian Amazon. Museo Ecuatoriano de Ciencias Naturales, Quito. 2 pp. [in press] Letters and reviews
2008. Willmott, K. R. Catalogue of the Hostplants of the Neotropical Butterflies/ Catálogo de las Plantas Huésped de las Mariposas Neotropicales. By Beccaloni, G. W., Viloria, A. L., Hall, S. K. & Robinson, G. S. (2008). Tropical Lepidoptera Research 18(2): 124-125. [book review]
2009. Hall, J. P. W., and K. R. Willmott. Two new species of Euselasia (Riodinidae: Euselasiinae) from western Ecuador. Tropical Lepidoptera Research, 19(1): 52-55. Download reprint [PDF]
2009. Elias, M., Joron, M., Willmott, K. R., Kaiser, V., Silva-Brandão, K. L., Freitas, A. V. L., Arias, C. F., Gomez Piñeres, L. M., Uribe, S., Jiggins, C. Out of the Andes: patterns of diversification in clearwing butterflies. Molecular Ecology, 18: 1716-1729. Download page proofs [PDF]
2008. Elias, M., Gompert, Z., Jiggins, C., Willmott, K. R. Mutualistic interations drive ecological niche convergence in a diverse butterfly community. PLoS Biology, 6(12):e300. Definitive online version or Download reprint [PDF], Supporting data S1 [PDF], Supporting data S2 [PDF]
2008. Willmott, K. R., Vitale, F. Taxonomic notes on Napeogenes (Lepidoptera: Ithomiinae) from Ecuador and Colombia, with the description of ten new subspecies. Genus, 19(3): 585-609. Download reprint [PDF]
2008. Willmott, K. R., Lamas, G. A revision of the genus Megoleria (Lepidoptera, Nymphalidae, Ithomiinae). Tropical Lepidoptera Research, 18(1): 46-59. Download reprint [PDF]
2007. Elias, M., Hill, R., Dasmahapatra, K., Willmott, K. R., Brower, A., Mallet, J., and C. Jiggins. Limited performance of DNA barcoding in a diverse community of tropical butterflies. Proceedings of the Royal Society of London B, 274: 2881-2889. Download reprint [PDF] and suppplementary documentation
2007. Brehm, G., Hartmann, T., and K. R. Willmott. Pyrrolizidine alkaloids and pharmacophagous Lepidoptera visitors of Prestonia amabilis (Apocynaceae) in a montane rain forest in Ecuador. Annals of the Missouri Botanical Garden, 94(2): 463-473. Download reprint [PDF]
2007. Hall, J. P. W., and K. R. Willmott. Four new species of Symmachiini from Ecuador (Lepidoptera: Riodinidae). Tropical Lepidoptera, 16(1-2): 1-5. Download reprint [PDF]
2007. Willmott, K. R., and G. Lamas. A revision of Pachacutia, a new genus of rare Andean ithomiine butterflies (Nymphalidae: Ithomiinae), with the description of two new species. Annals of the Entomological Society of America, 100: 449-469. Download reprint [PDF]
2006. Pyrcz, T. W., Willmott, K. R., Hall, J. P. W., and A. L. Viloria. A review of the genus Manerebia Staudinger (Lepidoptera: Nymphalidae: Satyrinae) in the northern Andes. Journal of Research on the Lepidoptera, 39: 37-79 (2000). Download reprint [PDF]
2006. Willmott, K. R., and A. V. L. Freitas. Higher-level phylogeny of the Ithomiinae (Lepidoptera: Nymphalidae): classification, patterns of larval hostplant colonisation and diversification. Cladistics, 22: 297-368. Please e-mail one of the authors for a reprint.
2006. Jiggins, C. D., Mallarino, R., Willmott, K. R., and E. Bermingham. The phylogenetic pattern of speciation and wing pattern change in neotropical Ithomia butterflies (Lepidoptera: Nymphalidae). Evolution, 60: 1454-1466. Download reprint [PDF]
2006. Willmott, K. R., and G. Lamas. A phylogenetic reassessment of Hyalenna Forbes and Dircenna Doubleday, with a revision of Hyalenna (Lepidoptera: Nymphalidae: Ithomiinae). Systematic Entomology, 31(3): 419-468. Download reprint [PDF]
2006. Brower, A. V. Z., Freitas, A. V. L., Lee, M.-M., Silva Brandão, K. L., Whinnett, A., and K. R. Willmott. Phylogenetic relationships among the Ithomiini (Lepidoptera: Nymphalidae) inferred from one mitochondrial and two nuclear gene regions. Systematic Entomology, 31: 288-301. Download reprint [PDF]
2005. Whinnett, A., Willmott, K. R., Brower, A.V.Z., Simpson, F., Zimmermann, M., Lamas, G., and J. Mallet. Mitochondrial DNA provides an insight into the mechanisms driving diversification in the ithomiine butterfly Hyposcada anchiala (Lepidoptera: Nymphalidae, Ithomiinae). European Journal of Entomology, 102: 633-639. Download reprint [PDF]
2005. Whinnett, A., Zimmermann, M., Willmott, K. R., Herrera, N., Mallarino, R., Simpson, F., Joron, M., Lamas, G., and J. Mallet. Strikingly variable divergence times inferred across an Amazonian butterfly 'suture zone'. Proceedings of the Royal Society of London B, 272: 2525-2533. Download reprint [PDF]
2005. Whinnett, A., Brower, A. V. Z., Lee, M.-M., Willmott, K. R., and J. Mallet. The phylogenetic utility of tektin, a novel region for inferring systematic relationships amongst Lepidoptera. Annals of the Entomological Society of America, 98: 873-886. Download reprint [PDF]
2005. Mallarino, R., Bermingham, E., Willmott, K. R., Whinnett, A., and C. D. Jiggins. Molecular systematics of the butterfly genus Ithomia (Lepidoptera: Ithomiinae): a composite phylogenetic hypothesis based on seven genes. Molecular Phylogenetics and Evolution, 34(3): 625-644. Download reprint [PDF]
2005. Hall, J. P. W., K. R. Willmott and R. C. Busby. Five new Penaincisalia species (Lepidoptera: Lycaenidae: Eumaeini) from the Andes of southern Ecuador and northern Peru. Zootaxa, 797: 1-20. Download reprint [PDF]
2004. Balcázar-Lara, M., and K. R. Willmott. A new subspecies of Adelpha erymanthis from Mexico, with a key to identification of similar taxa (Lepidoptera: Nymphalidae: Biblidinae). Tropical Lepidoptera, 12(1-2)("2001"): 25-38. Download reprint [PDF]
2004. Willmott, K. R., and J. Mallet. Correlations between adult mimicry and larval hostplants in ithomiine butterflies. Proceedings of the Royal Society of London B (Suppl.) 271: S266-S269. Download reprint [PDF]
2004. Willmott, K. R., and Lamas, G. A new species of Tithorea (Lepidoptera: Nymphalidae: Ithomiinae) from the Chocó region of northwestern South America and Panama. J. New York Ent. Soc., 112(1): 1-17. Download reprint [PDF]
2004. Willmott, K. R., and J. P. W. Hall. Taxonomic notes on the genus Zaretis, with the description of a new species (Lepidoptera: Nymphalidae: Charaxinae). Tropical Lepidoptera, 12(1-2): 25-38. Download reprint [PDF]
2003. Mallet, J. L. B., and K. R. Willmott. Taxonomy: renaissance or Tower of Babel? Trends in Ecology and Evolution, 18(2): 57-59.
2003. Willmott, K. R. Cladistic analysis of the neotropical butterfly genus Adelpha (Nymphalidae: Limenitidini), with comments on the subtribal classification of the tribe Limenitidini. Systematic Entomology, 28: 1-43. Download reprint [PDF]
2002. Robinson-Dean, J. C., K. R. Willmott, M. J. Catterall, D. J. Kelly, A. Whittington, B. Phalan, N. M. Marples, and D. R. S. Boeadi. A new subspecies of Red-backed Thrush Zoothera erythronota kabaena subsp. nov. (Muscicapidae: Turdidae), from Kabaena island, Indonesia. Forktail, 18: 1-10.
2001. Willmott, K. R., J. P. W. Hall, and G. Lamas. Systematics of Hypanartia (Lepidoptera: Nymphalidae: Nymphalinae), with a test for geographical speciation mechanisms in the Andes. Systematic Entomology, 26: 369-399. Download reprint [PDF]
2001. Willmott, K. R., L. M. Constantino, and J. P. W. Hall. A review of Colobura (Lepidoptera: Nymphalidae) with comments on larval and adult ecology and description of a sibling species. Annals of the Entomological Society of America, 94(2): 185-196. Download reprint [PDF]
2000. Hall, J. P. W., and K. R. Willmott. Patterns of feeding behaviour in adult male riodinid butterflies and their relationship to morphology and ecology. Biol. J. Linn. Soc., 69: 1-23. Download reprint [PDF]
1999. Pyrcz, T., K. R. Willmott, and J. P. W. Hall. Contribution to the knowledge of Ecuadorian Pronophilini. Part III. Three new species and five new subspecies of Lymanopoda (Lepidoptera: Nymphalidae: Satyrinae). Genus, 10(3): 497-522. Download reprint [PDF - 1] [PDF - 2]
1999. Willmott, K. R., and J. P. W. Hall. Taxonomic notes on Ecuadorian Adelpha with the description of two new species and seven new subspecies (Lepidoptera: Nymphalidae: Limenitidinae). Tropical Lepidoptera, 10(1): 1-17. Download reprint [PDF - 1] [PDF - 2]
1998. Hall, J. P. W., and K. R. Willmott. Four new riodinid species from eastern Ecuador (Lepidoptera: Riodinidae). Lambillionea, 98: 325-334. Download reprint [PDF]
1998. Hall, J. P. W., and K. R. Willmott. Nine new species and one new subspecies of Euselasia from Ecuador (Lepidoptera: Riodinidae). Tropical Lepidoptera, Suppl. 1, 9(2): 27-35. Download reprint [PDF]
1998. Hall, J. P. W., and K. R. Willmott. Three new species of Riodinini from the cloud forests of Ecuador (Lepidoptera: Riodinidae). Tropical Lepidoptera, Suppl. 1, 9(2): 22-26. Download reprint [PDF]
1996. Hall, J. P. W., and K. R. Willmott. The genus Theope: four new species and a new subspecies (Lepidoptera: Riodinidae). Tropical Lepidoptera, 7(1): 63-67. Download reprint [PDF]
1996. Hall, J. P. W., and K. R. Willmott. Notes on the genus Argyrogrammana Part II, with the description of a new species. Tropical Lepidoptera, 7(1): 71-80. Download reprint [PDF]
1996. Hall, J. P. W., and K. R. Willmott. Systematics of the riodinid tribe Symmachiini, with the description of a new genus and five new species from Ecuador, Venezuela and Brazil (Lepidoptera: Riodinidae). Lambillionea, 96: 637-660. Download reprint [PDF]
1996. Torres, R., J. P. W. Hall, K. R. Willmott, and K. Johnson. A new genus of brilliantly colored elfin butterflies from the high Andes (Lepidoptera: Lycaenidae). Tropical Lepidoptera, 7(1): 81-86. Download reprint [PDF]
1995. Hall, J. P. W., and K. R. Willmott. Two new species of Mesene from western Ecuador (Lepidoptera: Riodinidae). Tropical Lepidoptera, 6(2): 110-112. Download reprint [PDF]
1995. Hall, J. P. W., and K. R. Willmott. Five new species and a new genus of riodinid from the cloud forests of eastern Ecuador (Lepidoptera: Riodinidae). Tropical Lepidoptera, 6(2): 131-135. Download reprint [PDF]
1995. Hall, J. P. W., and K. R. Willmott. Notes on the genus Argyrogrammana, with descriptions of five new species (Lepidoptera: Riodinidae). Tropical Lepidoptera, 6(2): 136-143. Download reprint [PDF]
1995. Willmott, K. R., and J. P. W. Hall. Two new species of satyrines from Ecuador (Nymphalidae: Satyrinae). Tropical Lepidoptera, 6(2): 103-105. Download reprint [PDF]
1995. Willmott, K. R., and J. P. W. Hall. Two new species of Adelpha from eastern Ecuador (Lepidoptera: Nymphalidae). Tropical Lepidoptera, 6(2): 106-109. Download reprint [PDF]
1994. Willmott, K. R., and J. P. W. Hall. Four new species of Riodinidae (Lepidoptera) from Western Ecuador. Tropical Lepidoptera, 5(2): 87-91. Download reprint [PDF]
1993. Hall, J. P. W., and K. R. Willmott. Temporal and elevational additions of distribution for Costa Rican butterflies. Tropical Lepidoptera, 4(2): 139-141.
2004. Willmott, K.R. Brush-footed butterflies (Lepidoptera: Nymphalidae). In: Capinera, J. L. (Ed.), Encyclopedia of Entomology, Volume I A-E, pp. 379-384. Dordrecht / Boston / London, Kluwer Academic Publishers. Download reprint [PDF]
2004. Willmott, K. R., and J. P. W. Hall. Common Butterflies of Yuturi and Yarina lodge. Yuturi Lodge, Ecuador. 2 pp.
2001. Hall, J. P. W., and K. R. Willmott. Ecuador. Out on a limb. National Geographic Magazine (Update from the field), 199(2): 86. Download reprint [PDF]
2001. Willmott, K. R., and J. P. W. Hall. Winged gems of Ecuador. Biodiversity paradise. National Geographic Magazine (Geographica) 199(5): xxviii. Download reprint [PDF]
1998. Willmott, K. R. A fair review? News of the Lepidopterist's Society, 40(4): 92-93.
1997. Hall, J. P. W., and K. R. Willmott. The Butterflies of Venezuela. Part I. Nymphalidae I (Limenitidinae, Apaturinae, Charaxinae). A Comprehensive Guide to the Identification of Adult Nymphalidae, Ppailionidae and Pieridae. By Andrew F. E. Neild. Tropical Lepidoptera, 8(1): 46. [book review]
