Butterflies of Southeastern Sulawesi

(Papilionidae, Pieridae, Nymphalidae, Riodinidae, Lycaenidae)

An Illustrated Checklist

Southeastern Sulawesi includes mainland, Buton, Kabaena, Muna, Wowoni and Muna islands, and Kep. Tukangbesi

Links "MALE", "FEMALE" are images showing dorsal (left) and ventral (right) wings. Alternatively, click on a family or subfamily name to show thumbnail images of all species, which link to larger images. Images for Lycaenidae are already arranged in plates. Images for Pieridae and Biblidinae should be added soon.

"Endemic" is endemic to Sulawesi mainland and adjacent islands. "S" = Sulawesi mainland; "B" = Buton island; "K" = Kabaena island; "M" = Muna island; "W" = Wowoni island; "T" = Tukangbesi islands; "?" = possible range.

Records are based on field work by Keith Willmott and other volunteers for Operation Wallacea on mainland, Buton, Kabaena and Pulau Hoga (Tukangbesi) during summer 2000 and: Vane-Wright, R. I., & R. de. Jong. 2003. The butterflies of Sulawesi: annotated checklist for a critical island fauna. Zoologische Verhandlingen, 343: 1-267.

PAPILIONIDAE
Papilioninae
Troidini
Troides Hübner, 1819
Troides helena hephaestus C. & R. Felder, 1864 S, B, K, T
Troides hypolitus cellularis Rothschild, 1895 S, B, K
Pachliopta Reakirt, 1865
Pachliopta polyphontes polyphontes (Boisduval, 1836) S, B, K, M, W MALE


Papilionini
Papilio Linnaeus, 1758
Papilio ascalaphus ascalaphus Boisduval, 1836 Endemic S, B
Papilio ascalaphus munascalaphus Hachitani, 1988 Endemic M
Papilio blumei fruhstorferi Röber, 1897 Endemic S
Papilio fuscus metagenes Fruhstorfer, 1904 T
Papilio fuscus pertinax Wallace, 1865 S, K
Papilio gigon gigon C. & R. Felder, 1864 Endemic S, B, K
Papilio peranthus Fabricius, 1787 ssp. 1 S
Papilio peranthus Fabricius, 1787 ssp. 2 K
Papilio peranthus kransi Jurriaanse & Lindemans, 1920 B
Papilio peranthus wangiwangiensis Kitahara, 1989 T
Papilio polytes alcindor Oberthür, 1879 S, B, K
Papilio polytes tucanus Jordan, 1909 T FEMALE
Papilio sataspes sataspes C. & R. Felder, 1864 Endemic S, B


Lampropterini
Graphium Scopoli, 1777
Graphium agamemnon comodus Fruhstorfer, 1903 S, B, K, M, W, T MALE
Graphium androcles androcles (Boisduval, 1836) Endemic S, B, K MALE
Graphium codrus celebensis Wallace, 1865 S, B, K FEMALE
Graphium deucalion deucalion (Boisduval, 1836) S, B MALE
Graphium dorcus butungensis Hanafusa, 1997 Endemic S, B, K MALE
Graphium encelades (Boisduval, 1836) Endemic S, B MALE
Graphium eurypylus pamphylus C. & R. Felder, 1865 S, B, K MALE
Graphium meyeri meyeri Hopffer, 1874 Endemic S, B, K MALE
Graphium milon milon C. & R. Felder, 1864 S, B, K MALE
Graphium rhesus rhapia Jordan, 1908 Endemic T
Graphium rhesus rhesus (Boisduval, 1836) Endemic S, B MALE
Lamproptera Gray, 1832
Lamproptera meges ennius C. & R. Felder, 1865 S, B


PIERIDAE
Coliadinae
Gandaca Moore, 1906
Gandaca butyrosa samanga Fruhstorfer, 1910 S, B
Eurema Hübner, 1819
Eurema alitha C. & R. Felder, 1862 ssp. S, B, K, M, T
Eurema blanda odinia Fruhstorfer, 1910 S, B, K
Eurema brigitta ina Eliot, 1956 S, M
Eurema celebensis celebensis Wallace, 1867 Endemic S, B, K
Eurema hecabe kalidupa Fruhstorfer, 1910 T
Eurema hecabe latimargo Hopffer, 1874 S, B, K, M
Eurema tominia faunia Fruhstorfer, 1910 T
Eurema tominia tominia Vollenhoven, 1865 S, B, K
Catopsilia Hübner, 1819
Catopsilia pomona flava Butler, 1869 S, B, K, M, T
Catopsilia pyranthe pyranthe (Linnaeus, 1758) S, B, K
Catopsilia scylla asema Staudinger, 1885 S, B, K, T


Pierinae
Pareronia Bingham, 1907
Pareronia tritaea bargylia Fruhstorfer, 1910 Endemic S
Pareronia tritaea sarasinorum Martin, 1913 Endemic B, K, M
Pareronia tritaea (C. & R. Felder, 1859) ssp. Endemic T
Hebomoia Hübner, 1819
Hebomoia glaucippe celebensis Wallace, 1863 S, B, K
Leptosia Hübner, 1818
Leptosia nina dione Wallace, 1867 S, B, K, M
Leptosia nina (Fabricius, 1793) ssp. T
Delias Hübner, 1819
Delias rosenbergi munaensis Nakano, 1988 Endemic B, M
Delias rosenbergi rosenbergi Vollenhoven, 1865 Endemic S
Delias rosenbergi ssp. Endemic K
Delias zebuda Hewitson, 1862 Endemic S, K, M
Appias Hübner, 1819
Appias albina albina (Boisduval, 1836) S, B, K
Appias aurosa Yata & Vane-Wright, in press. Endemic S, B
Appias hombroni sulanorum Fruhstorfer, 1902 K
Appias hombroni tombugensis Fruhstorfer, 1902 S, B
Appias ithome C. & R. Felder, 1859 Endemic K
Appias lyncida floresiana Butler, 1898 T
Appias lyncida (Cramer, 1777) ssp. 1 K
Appias lyncida (Cramer, 1777) ssp. 2 B
Appias paulina dohertyi Rothschild, 1802/albata Hopffer, 1874 S?
Appias zarinda zarinda Boisduval, 1836 S, B, K, T
Saletera Distant, 1885
Saletera panda nigerrima Holland, 1891 S, B, K
Cepora Billberg, 1820
Cepora celebensis kazuyoe Watanabe, 1987 Endemic B
Cepora eperia Boisduval, 1836 Endemic S, B
Cepora fora milos Watanabe, 1987 Endemic B
Cepora perimale toekangbesiensis Jurriaanse & Lindemans, 1920 T
Cepora perimale (Donovan, 1805) ssp. M
Cepora timnatha Hewitson, 1862 ssp. Endemic S, B, K
Aoa de Nicéville, 1898
Aoa affinis (Vollenhoven, 1865) Endemic S, B


LYCAENIDAE
Miletinae
Miletini
Allotinus C. & R. Felder, 1865
Allotinus albatus C. & R. Felder, 1865 S?
Allotinus macassarensis Holland, 1891 Endemic S, B, K MALE FEMALE
Allotinus major C. & R. Felder, 1865 Endemic S, B MALE FEMALE
Allotinus maximus Staudinger, 1888 Endemic S, B MALE
Allotinus unicolor C. & R. Felder, 1865 S?
Logania Distant, 1884
Logania obscura Röber, 1886 Endemic S, B, T MALE FEMALE
Miletus Hübner, 1819
Miletus boisduvali Moore, 1858 S
Miletus celinus Eliot, 1961 Endemic S, K FEMALE
Miletus leos Guérin-Ménéville, 1830 ssp. Endemic T
Miletus leos maximus Holland, 1890 S, B, K MALE FEMALE
Spalgis Moore, 1879
Spalgis epius substrigatus Snellen, 1878 S, B, T MALE FEMALE


Curetinae
Curetis Hübner, 1819
Curetis tagalica celebensis C. & R. Felder, 1865 S, B, K, T MALE FEMALE
Curetis venata venata Fruhstorfer, 1908 Endemic S?


Theclinae
Arhopalini
Arhopala Boisduval, 1832
Arhopala acetes Hewitson, 1862 Endemic S, B, K, T MALE FEMALE
Arhopala alitaeus alitaeus Hewitson, 1862 S MALE FEMALE
Arhopala araxes araxes C. & R. Felder, 1869 S, K MALE FEMALE
Arhopala argentea Staudinger, 1888 Endemic S, B, K MALE FEMALE
Arhopala cleander sostrata Fruhstorfer, 1914 S?
Arhopala dohertyi Bethune-Baker, 1903 Endemic S MALE
Arhopala eridanus lewara Ribbe, 1926 S MALE FEMALE
Arhopala hercules hercules Hewitson, 1862 S MALE
Arhopala irregularis Bethune-Baker, 1903 Endemic S MALE
Arhopala quercoides Röber, 1886 Endemic S MALE FEMALE
Arhopala straatmani Nieuwenhuis, 1969 Endemic S?
Arhopala tephlis bicolora Röber, 1886 S MALE FEMALE
Arhopala sp. Endemic B FEMALE
Flos Doherty, 1889
Flos apidanus (Cramer, 1779) ssp. S, B MALE
Flos arca de Nicéville, 1893 Endemic S?
Flos diardi imperiosa Fruhstorfer, 1914 S?
Flos kuehni (Röber, 1887) Endemic S FEMALE
Surendra Moore, 1879
Surendra vivarna samina Fruhstorfer, 1904 S, B FEMALE


Amblypodini
Amblypodia Horsfield, 1829
Amblypodia narada confusa Riley, 1922 S?
Iraota Moore, 1881
Iraota rochana johnsoniana Holland, 1890 S, B, K FEMALE


Hypotheclini
Hypothecla Semper, 1890
Hypothecla honos de Nicéville, 1898 Endemic S?


Loxurini
Loxura Horsfield, 1829
Loxura atymnus sulawesiensis Takanami, 1986 S?
Loxura atymnus (Stoll, 1780) ssp. T


Horagini
Horaga Moore, 1881
Horaga syrinx permagna Fruhstorfer, 1912 K MALE FEMALE
Horaga selina Grose-Smith, 1895 Endemic S, B FEMALE


Cheritrini
Drupadia Moore, 1884
Drupadia theda namusa Hewitson, 1863 S, B MALE FEMALE


Aphnaeini
Spindasis Wallengren, 1857
Spindasis vulcanus C. & R. Felder, 1862 ssp. B


Iolaini
Tajuria Moore, 1881
Tajuria cyrillus Hewitson, 1865 Endemic S, B, K FEMALE
Tajuria iapyx Hewitson, 1865 ssp. Endemic S, K, T MALE
Tajuria mantra jalysus C. & R. Felder, 1865 S, K MALE
Dacalana Moore, 1884
Dacalana anysides Röber, 1887 Endemic S?
Dacalana anysis Hewitson, 1865 Endemic S?


Remelanini
Remelana Moore, 1884
Remelana jangala orsolina Hewitson, 1865 S, B, K MALE FEMALE


Hypolycaenini
Hypolycaena C. &. R. Felder, 1862
Hypolycaena erylus gamatius Fruhstorfer, 1912 S, B, K, T MALE FEMALE
Hypolycaena sipylus giscon Fruhstorfer, 1912 S, B, K, T MALE FEMALE
Hypolycaena xenia Grose-Smith, 1895 Endemic S?


Deudorigini
Bindahara Moore, 1881
Bindahara phocides fumata Röber, 1887 S, K MALE
Rapala Moore, 1881
Rapala dioetas (Hewitson, 1869) Endemic S, B MALE FEMALE
Rapala enipeus Staudinger, 1888 Endemic S, B, K, T MALE FEMALE
Rapala manea manea (Hewitson, 1863) S, B MALE FEMALE
Rapala ribbei Röber, 1886 Endemic S?
Rapala varuna olivia Druce, 1895 S, M FEMALE
Deudorix Hewitson, 1863
Deudorix epijarbas megakles Fruhstorfer, 1911 S, B MALE FEMALE
Deudorix loxius Hewitson, 1869 Endemic S?
Artipe Boisduval, 1836
Artipe eryx alax Eliot, 1956 S?
Sinthusa Moore, 1884
Sinthusa indrasari Snellen, 1878 Endemic S?
Sinthusa verriculata Snellen, 1892 Endemic S MALE FEMALE


Polyommatinae
Lycaenesthini
Anthene Doubleday, 1847
Anthene licates licates Hewitson, 1874 S?
Anthene lycaenina Felder, 1868 S?
Anthene philo philo Hopffer, 1874 Endemic S, T MALE
Anthene villosa Snellen, 1878 Endemic S?


Polyommatini
Petrelaea Toxopeus, 1929
Petrelaea tombugensis Röber, 1886 S?
Nacaduba Moore, 1881
Nacaduba angusta pamela Grose Smith, 1895 S, B, K
Nacaduba berenice eliana Fruhstorfer, 1916 S, B, K, T MALE FEMALE
Nacaduba beroe (C. &. R. Felder, 1865) ssp. S?
Nacaduba kurava kurava (Moore, 1858) S? MALE FEMALE
Nacaduba pactolus pactolides Fruhstorfer, 1916 S?
Nacaduba pavana visuna Fruhstorfer, 1916 S, K MALE FEMALE
Nacaduba subperusia (Snellen, 1896) S MALE
Psychonotis Toxopeus, 1930
Psychonotis piepersii Snellen, 1878 Endemic S, B, K MALE FEMALE
Prosotas H. H. Druce, 1871
Prosotas aluta alutina Fruhstorfer, 1916 S
Prosotas dubiosa Semper, 1879 ssp. S, B, K MALE
Prosotas gracilis gracilis Röber, 1886 S?
Prosotas nora nora (Felder, 1860) S, B, K MALE FEMALE
Prosotas pia elioti Tite, 1963 S, B, K MALE
Nothodanis Hirowatari, 1992
Nothodanis schaeffera schaeffera Erschoff, 1821 S?
Catopyrops Toxopeus, 1929
Catopyrops ancyra subfestivus Röber, 1886 S, B, K MALE FEMALE
Catopyrops rita bora Eliot, 1956 S?
Ionolyce Toxopeus, 1929
Ionolyce helicon helicon Felder, 1860 S?
Caleta Fruhstorfer, 1922
Caleta rhode rhodana Fruhstorfer, 1918 S, B MALE
Caleta caleta caleta Hewitson, 1876 S, B MALE
Caleta celebensis Staudinger, 1889 Endemic S?
Discolampa Toxopeus, 1929
Discolampa ethion ulyssides Grose Smith, 1895 S MALE
Discolampa ilissus ilissus (Felder, 1859) S, B MALE
Niphanda
Niphanda tessellata Moore, ssp. K MALE FEMALE
Jamides Hübner, 1819
Jamides alecto latimargus Snellen, 1878 S, K MALE
Jamides aratus lunata de Nicéville, 1899 S, B, K MALE FEMALE
Jamides aratus (Stoll, 1781) ssp. T
Jamides bochus phaidon Fruhstorfer, 1915 S, B, K MALE FEMALE
Jamides celeno optimus Röber, 1886 S, B, K, T MALE FEMALE
Jamides cyta zelia Fruhstorfer, 1916 S, B MALE FEMALE
Jamides festivus festivus Röber, 1886 Endemic S, B MALE FEMALE
Jamides fractilinea Tite, 1960 Endemic S?
Jamides philatus philatus Snellen, 1878 S, B, K, T MALE FEMALE
Jamides pseudosias echeilea Fruhstorfer, 1916 S?
Jamides snelleni Röber, 1886 Endemic S, K MALE
Catochrysops Boisduval, 1832
Catochrysops panormus Felder, 1860 ssp. S?
Catochrysops strabobinna Swinhoe, 1916 S?
Catochrysops strabo celebensis Tite, 1959 S, B, K, T MALE FEMALE
Lampides Hübner, 1819
Lampides boeticus (Linnaeus, 1767) S, B, K, T FEMALE
Castalius Hübner, 1819
Castalius clathratus Holland, 1891 Endemic S?
Castalius fluvialis Grose-Smith, 1895 Endemic S?
Castalius rosimon silas Fruhstorfer, 1922 S?
Famegana Eliot, 1973
Famegana alsulus kalawarus Ribber, 1926 S?
Pithecops Horsfield, 1828
Pithecops phoenix Röber, 1886 Endemic S, B, K
Leptotes Scudder, 1876
Leptotes plinius zingis Fruhstorfer, 1922 S, B MALE FEMALE
Zizeeria Chapman, 1910
Zizeeria karsandra karsandra (Moore, 1865) S, T
Zizina Chapman, 1910
Zizina otis tanagra Felder, 1860 T
Zizina otis (Fabricius, 1787) ssp. S, B, K MALE FEMALE
Zizula Chapman, 1910
Zizula hylax hylax (Fabricius, 1775) S, B, K, T FEMALE
Everes Hübner, 1819
Everes lacturnus Godart, 1824 ssp. S, B, K MALE
Megisba Hübner, 1819
Megisba malaya sikkima Moore, 1884 S, B, K, T MALE FEMALE
Sancterila Eliot & Kawazoé, 1983
Sancterila deliciosa (Pagenstecher, 1896) Endemic S, K MALE FEMALE
Udara Toxopeus, 1928
Udara aristius aristius (Fruhstorfer, 1910) Endemic S?
Udara camenae euphon Fruhstorfer, 1910 S?
Udara dilecta thoria Fruhstorfer, 1910 S?
Udara placidula placidula (H. H. Druce, 1895) S?
Udara rona rona (Grose Smith, 1894) S?
Acytolepis Toxopeus, 1927
Acytolepis najara (Fruhstorfer, 1910) Endemic S?
Acytolepis puspa kuehni Röber, 1886 S, B, K MALE
Acytolepis samanga (Fruhstorfer, 1910) Endemic S?
Celastrina Tutt, 1906
Celastrina lavendularis lyce Grose Smith, 1896 S?
Celastrina philippina gradeniga Fruhstorfer, 1910 S?
Uranobothria Toxopeus, 1927
Uranobothria celebica (Fruhstorfer, 1917) Endemic S?
Monodontides Toxopeus, 1927
Monodontides cara (de Nicéville, 1898) Endemic S?
Monodontides kolari (Ribbe, 1926) S, K MALE
Euchrysops Butler, 1900
Euchrysops cnejus (Fabricius, 1798) ssp. S, B, T MALE FEMALE
Chilades Butler, 1900
Chilades boopis boopis Fruhstorfer, 1915 S, K MALE FEMALE
Chilades putli (Kollar, 1848) S?


RIODINIDAE
Hamearinae
Zemeros Boisduval, 1836
Zemeros flegyas sosiphanes Fruhstorfer, 1912 S?
Abisara C. & R. Felder, 1860
Abisara echerius celebica Röber, 1886 S, B, K


NYMPHALIDAE
Morphinae
Amathusiini
Faunis Hübner, 1819
Faunis menado pleonasma Röber, 1896 Endemic S, B, K
Amathusia Fabricius, 1807
Amathusia phidippus celebensis Fruhstorfer, ADD S, B
Amathusia virgata thoanthea Fruhstorfer, 1911 Endemic S, B, K
Amathuxidia Staudinger, 1887
Amathuxidia plateni iamos Brooks, 1937 Endemic S, B, K
Discophora Boisduval, 1836
Discophora bambusae celebensis Holland, 1891 Endemic S, B


Satyrinae
Melanitini
Bletogona C. & R. Felder, 1867
Bletogona mycalesis mycalesis C. & R. Felder, 1867 Endemic S, B, K MALE FEMALE
Melanitis Fabricius, 1807
Melanitis leda celebicola Martin, 1929 S, B, K MALE FEMALE
Melanitis phedima linga Fruhstorfer, 1908 S, B, K MALE FEMALE
Melanitis pyrrha hylecoetes Holland, 1890 S, K MALE
Melanitis velutina ribbei Röber, 1886 S, B, K MALE FEMALE


Elymniini
Elymnias Hübner, 1818
Elymnias cumaea C. & R. Felder, 1867 ssp. Endemic
Elymnias hewitsoni atys Fruhstorfer, 1907 Endemic S, B, K MALE FEMALE
Elymnias hicetas butona Fruhstorfer, 1904 B, K MALE FEMALE
Elymnias hicetas hicetina Fruhstorfer, 1904 S
Elymnias mimalon nysa Fruhstorfer, 1907 Endemic S
Zethera Felder, 1861
Zethera incerta tenggara Roos, 1992 Endemic S MALE
Lethe Hübner, 1818
Lethe europa arcuata Butler, 1868 S, B, K MALE FEMALE
Orsotriaena Wallengren, 1858
Orsotriaena jopas jopas Hewitson, 1864 Endemic S, B, K MALE FEMALE
Orsotriaena medus medus (Fabricius, 1775) S, B, K, T MALE
Mycalesis Wallengren, 1858
Mycalesis horsfieldi Moore, 1892 ssp. K MALE FEMALE
Mycalesis itys remulina Fruhstorfer, 1897 Endemic S, B MALE FEMALE
Mycalesis janardana opaculus Fruhstorfer, 1908 S, B, K MALE FEMALE
Mycalesis newayana Fruhstorfer, 1911 Endemic S, B, K, T MALE FEMALE
Mycalesis perseus lalassis Hewitson, 1864 S, K, T MALE FEMALE
Lohora Moore, 1880
Lohora opthalmica Westwood, 1888 Endemic S, B, K MALE FEMALE
Lohora transiens Fruhstorfer, 1908 Endemic S
Lohora umbrosa Roos, 1997 Endemic S MALE FEMALE
Lohora unipupillata Fruhstorfer, 1898 Endemic S


Ragadiini
Acrophtalmia Felder, 1861
Acrophtalmia leuce C. & R. Felder, 1861 Endemic S, B MALE


Satyrini
Ypthima Hübner, 1818
Ypthima nynias nynias Fruhstorfer, 1911 Endemic S, B, K, M MALE
Ypthima ancus Fruhstorfer, 1911 Endemic S, K
Ypthima kalelonda anana Fruhstorfer, 1911 S
Ypthima loryma Hewitson, 1865 Endemic S, B, K
Ypthima norma pusilla Fruhstorfer, 1911 S, B
Ypthima sp. Endemic S, K MALE FEMALE


Charaxinae
Charaxini
Charaxes Ochsenheimer, 1816
Charaxes affinis affinis Butler, 1865 Endemic S
Charaxes affinis butongensis Tsukada, 1991 Endemic B, K MALE FEMALE
Charaxes mars mars Staudinger, 1885 S?
Charaxes nitebis nitebis Hewitson, 1862 Endemic S, B, K MALE FEMALE
Charaxes solon brevis Tsukada, 1991 B, K MALE FEMALE
Charaxes solon hannibal Butler, 1869 S
Polyura Billberg, 1820
Polyura alphius piepersianus Martin, 1924 S?
Polyura cognata bellona Tsukada, 1991 Endemic T
Polyura cognata cognata Vollenhoven, 1861 Endemic S


Heliconiinae
Acraeini
Acraea Fabricius, 1807
Acraea andromacha indica Röber, 1885 T
Acraea moluccana dohertyi Holland, 1891 S, B, K MALE


Heliconiini
Cethosiiti
Cethosia Fabricius, 1807
Cethosia biblis picta C. & R. Felder, 1867 S, B, K, M MALE FEMALE
Cethosia myrina sarnada Fruhstorfer, 1912 Endemic S MALE
Cethosia myrina vanbemmeleni Juriaanse & Lindemans, 1918 Endemic B MALE FEMALE
Argynniti
Terinos Boisduval, 1836
Terinos taxiles poros Fruhstorfer, 1906 S, B MALE FEMALE
Vindula Hemming, 1934
Vindula dejone (Erichson, 1834) ssp. T
Vindula dejone celebensis Butler, 1883 S, B MALE FEMALE
Vindula dejone bushi Tsukada, 1985 K
Vindula erota banta Eliot, 1956 S
Vindula erota boetonensis Jurriaanse & Lindemans, 1920 B MALE
Cupha Billberg, 1820
Cupha arias celebensis Fruhstorfer, 1899 S
Cupha arias muna Fruhstorfer, 1897 B, K, M
Cupha arias Felder, 1867 ssp. T MALE FEMALE
Cupha maeonides butungensis Tsukada, 1985 Endemic B MALE
Cupha maeonides maeonides Hewitson, 1859 Endemic S, K, M
Algia Herrich-Schäffer, 1864
Algia satyrina C. & R. Felder, 1867 ssp. Endemic S MALE
Cirrochroa Doubleday, 1847
Cirrochroa eremita Tsukada, 1985 Endemic S MALE
Cirrochroa semiramis C. & R. Felder, 1867 Endemic S, B MALE
Cirrochroa thule C. & R. Felder, 1860 Endemic S MALE
Vagrans Hemming, 1934
Vagrans sinha nupta Staudinger, 1889 B MALE
Phalanta Horsfield, 1829
Phalanta alcippe celebensis Wallace, 1869 S, B, W MALE
Phalanta phalantha sabulum Tsukada, 1985 S, B


Biblidinae
Limenitidini
Neptiti
Pantoporia Hübner, 1819
Pantoporia antara pytheas Fruhstorfer, 1913 S, B, K, T
Lasippa Moore, 1898
Lasippa neriphus tawayana Fruhstorfer, 1899 Endemic S, B, K, M
Neptis Fabricius, 1807
Neptis celebica celebica Moore, 1899 Endemic S?
Neptis hylas timorensis Röber, 1891 T
Neptis ida kalidupa Eliot, 1969 Endemic T
Neptis ida liliputa Martin, 1924 Endemic B, K, M
Neptis ida sphaerica Fruhstorfer, 1907 Endemic S
Phaedyma Felder, 1861
Phaedyma daria albescens Rothschild, 1892 Endemic S, K
Partheniti
Parthenos Hübner, 1819
Parthenos sylvia salentia Hopffer, 1874 S
Euthaliti
Lexias Boisduval, 1832
Lexias aeetes butongensis Tsukada, 1991 Endemic B, K
Lexias aeetes phasiana Butler, 1870 Endemic S
Euthalia Hübner, 1819
Euthalia amanda amanda Hewitson, 1862 Endemic S, K
Euthalia amanda rubicundus Tsukada, 1991 Endemic B, T
Dophla Moore, 1880
Dophla evelina dermoides Rothschild, 1892 S, B, K
Bassarona Moore, 1897
Bassarona labotas pallesco Tsukada, 1991 Endemic B
Bassarona labotas labotas (Hewitson, 1864) Endemic S
Genus unknown (Tanaecia or Euthalia)
Sp. B
Limenitiditi
Tarattia Moore, 1898
Tarattia lysanias aberrans Hanafusa, 1989 Endemic M
Tarattia lysanias lysanias (Hewitson, 1859) Endemic S, B, K
Athyma Westwood, [1850]
Athyma libnites libnites (Hewitson, 1859) Endemic S
Athyma libnites noctesco Tsukada, 1991 Endemic B, K, M
Moduza Moore, 1881
Moduza lycone lyconides Fruhstorfer, 1913 Endemic B, K
Moduza lymire lymire (Hewitson, 1859) Endemic S
Moduza lymire munaensis Hanafusa, 1989 Endemic B, M
Moduza lymire nectareus Tsukada, 1991 Endemic K
Lamasia Moore, 1898
Lamasia lyncides notus Tsukada, 1991 Endemic S, B, K, M
Tacola Moore, 1898
Tacola eulimene badoura Butler, 1866 S, B, K


Biblidini
Ariadne Horsfield, 1829
Ariadne ariadne gedrosia Fruhstorfer, 1912 B, K, M MALE
Ariadne celebensis Holland, 1891 ssp. Endemic S
Ariadne merionoides merionoides Holland, 1891 Endemic S, B, K MALE FEMALE


Marpesiini
Chersonesia Distant, 1883
Chersonesia rahria celebensis Rothschild, 1892 S, B MALE
Cyrestis Boisduval, 1832
Cyrestis paulinus kransi Jurriaanse & Lindemans, 1920 S, B, W MALE
Cyrestis strigata strigata C. & R. Felder, 1862 Endemic S, B, K, W MALE
Cyrestis thyonneus celebensis Staudinger, 1896 S, B, K MALE


Pseudergolini
Pseudergolis C. & R. Felder, 1867
Pseudergolis avesta nimbus Tsukada, 1991 Endemic B MALE
Pseudergolis avesta toalarum Fruhstorfer, 1912 Endemic S
Dichorragia Butler, 1869
Dichorragia nesimachus pelurius Fruhstorfer, 1897 S, B FEMALE


Nymphalinae
Nymphalini
Symbrenthia Hübner, 1819
Symbrenthia hippalus C. & R. Felder, 1867 Endemic S?
Symbrenthia hippoclus clausus Fruhstorfer, 1904 B, K MALE
Symbrenthia lilaea utakata Tsukada & Nishiyama, 1985 S?
Junonia Hübner, 1819
Junonia almana battana Fruhstorfer, 1906 S, B, K MALE
Junonia atlites acera Fruhstorfer, 1912 S, B, K MALE
Junonia erigone gardineri Fruhstorfer, 1902 S, B, K, T MALE
Junonia hedonia intermedia C. & R. Felder, 1867 S, B, K MALE
Junonia orithya celebensis Staudinger, 1888 S, K, M
Junonia orithya kuehni Fruhstorfer, 1904 T
Yoma Doherty, 1886
Yoma sabina nimbus Tsukada, 1985 S, B, K, M, T MALE
Rhinopalpa C. & R. Felder, 1860
Rhinopalpa polynice megalonice C. & R. Felder, 1867 S, B MALE
Hypolimnas Hübner, 1819
Hypolimnas alimena alimena (Linnaeus, 1758) T MALE
Hypolimnas anomala stellata Fruhstorfer, 1912 S, B, K FEMALE
Hypolimnas bolina bolina (Linnaeus, 1758) S, B, K MALE FEMALE
Hypolimnas diomea diomea Hewitson, 1861 Endemic S, B MALE
Hypolimnas diomea serica Tsukada, 1985 Endemic K
Hypolimnas misippus (Linnaeus, 1758) (CHECK not 1764) S, M
Doleschallia C. & R. Felder, 1860
Doleschallia polibete celebensis Fruhstorfer, 1899 S, B, K, T MALE


Apaturinae
Rohana Moore, 1880
Rohana macar butongensis Tsukada, 1991 Endemic B MALE
Rohana macar macar (Wallace, 1869) Endemic S, K, M
Helcyra Felder, 1860
Helcyra celebensis celebensis Martin, 1913 Endemic S
Helcyra celebensis semifusca Masui, 1999 Endemic B FEMALE
Euripus Doubleday, 1848
Euripus robustus Wallace, 1869 Endemic S?


Libytheinae
Libythea Fabricius, 1807
Libythea geoffroy celebensis Staudinger, 1889 S?
Libythea narina canuleia Fruhstorfer, 1909 S?


Danainae
Danaini
Parantica Moore, 1880
Parantica cleona luciplena Fruhstorfer, 1892 S, B, K, W MALE
Parantica menadensis Moore, 1883 ssp. Endemic S, B MALE
Ideopsis Horsfield, 1858
Ideopsis juventa ishma Butler, 1869 S, B, K, M FEMALE
Ideopsis juventa sequana Fruhstorfer, 1910 T FEMALE
Ideopsis vitrea arachosia Fruhstorfer, 1910 S, B, K FEMALE
Tirumala Moore, 1880
Tirumala choaspes kroeseni Martin, 1910 Endemic B MALE
Tirumala choaspes (Butler, 1886) ssp. Endemic S
Tirumala hamata goana Martin, 1910 S?
Tirumala ishmoides ishmoides Moore, 1883 S, M
Tirumala limniace makassara Martin, 1910 S?
Danaus Kluk, 1802
Danaus affinis fulgurata Butler, 1866 S
Danaus affinis (Fabricius, 1775) ssp. 1 K
Danaus affinis (Fabricius, 1775) ssp. 2 T FEMALE
Danaus chrysippus gelderi Snellen, 1891 S
Danaus chrysippus (Linnaeus, 1758) ssp. 1 T
Danaus chrysippus (Linnaeus, 1758) ssp. 2 K
Danaus genutia leucoglene C. & R. Felder, 1865 S
Danaus genutia telemissus Fruhstorfer, 1910 B, K, M MALE
Danaus genutia (Cramer, 1779) ssp. T
Danaus ismare fulvus Ribbe, 1890 S, K MALE


Euploeini
Euploea Fabricius, 1807
Euploea algea horsfieldi C. & R. Felder, 1865 S
Euploea algea tombugensis Fruhstorfer, 1899 B, K, M, W MALE FEMALE
Euploea configurata (C. & R. Felder, 1865) Endemic S, K MALE FEMALE
Euploea eleusina vollenhovii C. & R. Felder, 1865 S, K MALE FEMALE
Euploea eupator vanoorti Jurriaanse & Lindemans, 1920 Endemic K MALE
Euploea eupator Hewitson, 1858 ssp. Endemic B MALE FEMALE
Euploea hewitsonii hewitsonii C. & R. Felder, 1865 Endemic S
Euploea hewitsonii reducta Jurriaanse, 1919 Endemic B, K, M MALE
Euploea latifasciata latifasciata Weymer, 1885 S?
Euploea phaenareta (Schaller, 1785) ssp. B
Euploea redtenbacheri redtenbacheri C. & R. Felder, 1865 S, B MALE FEMALE
Euploea sylvester schlegelii C. & R. Felder, 1865 S, B
Euploea sylvester (Fabricius, 1793) ssp. K, M MALE
Euploea westwoodii meyeri Hopffer, 1874 Endemic S MALE
Euploea westwoodii C. & R. Felder, 1865 ssp. Endemic B, K FEMALE
Idea Fabricius, 1807
Idea blanchardii munaensis Fruhstorfer, 1899 Endemic B, K, M MALE
Idea blanchardii phlegeton Fruhstorfer, 1904 Endemic T
Idea blanchardii Marchal, 1845 ssp. Endemic S