Lepus
Mammalia; Lagomorpha; Leporidae; Lepus (Jack Rabbit/Hare)
|
|
Steve Hutchens works on the Lepus mount.
|
| |
Although the names are frequently interchanged, members of the genus
Lepus are commonly called hares, not rabbits. A hare
is typically larger than a (cottontail) rabbit, reaching up
to 21 inches long from head to tail, and up to a foot tall,
and has noticeably longer ears than rabbits. Hares are also
distinct from rabbits in that the young are born in open fields,
and at birth are fully furred with their eyes wide open. Rabbits,
on the other hand, are born in well-constructed nests, hairless
and blind. Unlike rabbits that rely on burrows to escape danger,
hares rely on their running ability to escape would-be predators,
and rarely dig burrows.
Hares are generally brown to grayish brown in color with a white
underbelly. They are typically solitary nocturnal animals
that are active throughout the year. Hares, like rabbits,
eat grasses and other herbaceous matter. They are known, however,
to eat voles and younger rabbits/hares given the opportunity.
This particular specimen was excavated from an early Pleistocene
(2 MA) site in Citrus County, Florida. These sediments preserve
a microcosm of fossil birds, reptiles, and small mammals in
a karst fissure fill. Hares no longer live in Florida except
for the few individuals that managed to escape from their
would-be pet dealers around the Miami airport.
|
Life History
There are 29 known living species of Lepus, most of which live
in open grassland areas all over the world. Mainly nocturnal,
hares will forage at night, spending their inactive hours in
small depressions in the grass, soil or snow and not in true
burrows, like rabbits. Lepus are generally solitary animals
except during their rather lengthy mating season, anywhere from
3-9 months. They have 3-4 litters annually, with 1-6 leverets
(young hares) in each litter. Gestation is between 41-47 days
long.
Evolutionary History
|
Rabbits, hares, pikas, and their fossil relatives make up a group called
Lagomorpha. Lagomorphs are very similar to rodents in that they
both have a pair of enlarged ever-growing re-curved incisor
teeth well adapted for gnawing. Unlike rodents, however rabbits,
hares, and pikas have a second set of small incisors located
directly behind the large incisors.
Current research suggests that lagomorphs split from rodents early in
the Cenozoic (= The Age of Mammals), some 50 million years ago.
Shortly thereafter, rabbits and pikas are recognizably distinct
lineages in the fossil record. There are two families with the
Lagomorpha: Leporidae (hares and rabbits) and Ochotonids (pikas).
Within the Leporidae, there are some 13 genera (31 are extinct)
and 81 living species. The lineage of Lepus first appears
in the early Pliocene (5 MA) in Europe, making its way over
to North America by the late Pliocene.
|

The upper jaw of a lagomorph.
|
|
Biogeographical History
Hares are naturally widespread, and have become even more so since humans introduced them
to Australia and South America. Currently, hares are found on every
continent, except Antarctica. Lepus first appears in the fossil
record in the Early Pliocene (5MA) in Europe. By the late Pliocene,
they were widespread across Europe, Africa and North America.
Conservation Efforts
So far, hares have not been terribly perturbed as a result of human influence, nevertheless
some North American populations are considered threatened: L. americanus
in Virginia, L. callotis in Mexico and SW New Mexico, and
L. flavigularis of southern Mexico. Also considered threatened
elsewhere are: L. hainanus, L. castroviejoi, L. insularis
and L. yarkandensis.
Recently, hares
were introduced to areas where they had not occurred naturally (Australia,
New Zealand, Java, and parts of South America and Great Britain).
Their ability to adapt and flourish in these new environments has
taken its toll on the native fauna that has not been successful in
competing with them for the same resources. In some of these areas,
steps are being taken to eradicate these introduced hares to preserve
the local wildlife.
History of Specimen
|
The Citrus County site.
|
|
The Lepus skeleton, unmounted.
|
|
A detail of the hardware.
|
| |
The bones of this extinct hare skeleton (Lepus sp.) were excavated
from an early Pleistocene site (2 MA) in Citrus County, Florida
by Steven and Suzan Hutchens in 1994-1995. The skeleton was articulated,
mounted, and subsequently donated to the FLMNH by the Hutchenses
for display in The Hall of Florida Fossils: Evolution of Life
and Land. The method of articulation
employed is an innovative system of screws and linchpins that
allow the skeleton to be taken apart easily for research purposes.
This skeleton
is a composite of bones from many individuals found from the Citrus
County site, as well as from the NorthEastern Cottontail found
in Florida (Sylvilagus floridanus). This pictured mount
is composed of approximately 90% fossil material.
What is
so noteworthy about this particular specimen is the preservation
of the skull. Because they are so lightly built, complete intact
rabbit skulls are rarely preserved in the fossil record. Consequently,
this skull is unique in its preservations, and is the only known
skull of this extinct species, Lepus sp.
|
Bibliography
Hulbert, R. (in press) Emslie, S. D. 1998. Avian community, climate, and sea-level
changes in the Plio-Pleistocene of the Florida Peninsula. Ornithological
Monographs 50: pp 113.
McKenna, M.C., and S.K. Bell 1997. Classification of mammals above the species level.
Columbia Univ. Press, NY. 631 pp.
Nowak, R. M. 1999. Walker's Book of Mammals. Johns Hopkins University Press, London.
Pp. 1720-1721, 1733-1738.
Savage, D. E.
and D. E, Russell 1983. Mammalian Paleofaunas of the World. Addison-Wesley
Publishing Company. Reading, MA. P. 432.
|