INDEX | VETIGASTROPODA | PULMONTA (LYMNAEOIDEA-PUPILLOIDEA) | PULMONATA (ORTHALICOIDEA) |
PULMONATA (ACHATINOIDEA-SAGDOIDEA) | PULMONATA (GASTRODONTOIDEA-HELICOIDEA) | BIBLIOGRAPHY
PART
2: PULMONTA (Lymnaeidae – Vertiginidae)
SUBLASS PULMONATA –
The Pulmonate snails
Superfamily LYMNAEOIDEA
Family LYMNAEIDAE Rafinesque,
1815
Subfamily LYMNAEINAE
Rafinesque, 1915
Genus Fossaria Westerlund, 1885
Fossaria Westerlund, 1885; Fauna der in der
paläartischen Region lebenden Binnenconchylien, V: 49.
TYPE SPECIES.- Buccinum truncatula Müller, 1774
DISTRIBUTION.- Europe, Asia,
Africa, North America, the West Indies, Central America and
TAXONOMY.- Numerous species.
Subgenus Fossaria Westerlund, 1885
DISTRIBUTION.- The Holarctic
Realm.
TAXONOMY.- Numerous species. One species and two subspecies occurs in the
study area.
Fossaria obrussa
abrussa (Say, 1825)
Lymnaeus
obrussus Say, 1825;
Jour. Phila. Acad., 5: 123.
Galba
obrussa (Say). F. C. Baker, 1911; Chicago Acad. Sci. Spec.
Pub.., 3: 270-283; pl. 26, figs. 8-13; pl. 31, figs. 20-37.
Fossaria
obrussa (Say).
Lymnaea
decidiosa
(Gould). Pilsbry, 1904; Proc. Acad. nat.
Sci. Phila., 54: 777.
TYPE LOCALITY.- Harrowgate,
Philadelphia Co.,
DISTRIBUTION RECORDS.- Widely
distributed in the
Fossaria
(Fossarria) obrussa modicella (Say,
1825)
Lymnaeus
modicellus Say,
1825; Jour. Acad. Nat. Sci. Phila., 5: 122.
Galba
humilis modicella
(Say). F. C. Baker, 1911; Chicago Acad.
Sci. Sp. Pub., 3: 259-268; pl. 29, figs. 31-37 (shell); pl. 31, figs. 1-9
(shell).
Fossaria (Fossarria) modicella
(Say). Bequaert & Miller, 1973:
190-191.
Fossarria
obrussa modicella
(Say). Burch, 1980; Walkerana, 1: 172;
fig. 574.
TYPE LOCALITY.- Susquehanna River
at Oswego, Tioga Co., New Youk. Syntypes
ANSP 58790 (2).
DISTRIBUTION RECORDS.- Widespread
in
Subgenus Bakerlymnaea Weyrauch, 1964
Nasonia F. C. Baker, 1928; Bull. Wisconsin
Geol. Nat. Hist. Survey, 70 (1): 264 (non Nasonia
Ashmead, 1904, Hymenoptera).
Bakerlymnaea Weyrauch, 1964; Arch. für Moll.,
93: 169.
TYPE SEPCIES.- Lymnaea cubensis Pfeiffer, 1839.
DISTRIBUTION.- Southern United
States and the West Indies south to
TAXONOMY.- Numerous species and
subspecies are recognized. Three occur
in the study area.
Fossarria (Bakerlymnaea) bulimoides techella
(Haldeman, 1867)
Limnaea
techella Haldeman,
1867; Amer. Jour. Conch., 3: 194; pl. 6, fig. 4 (shell).
Galba
bulimoides techella
(Haldeman). F. C. Baker, 1911; Chicago
Acad. Sci. Sp. Pub., 3: 214-217; pl. 27, figs. 30-35 (shell); pl. 28, figs.
1-3, 8 (shell).
Stagnicola (Bakerlymnaea) bulimoides
techella (Haldeman). Bequaert &
Miller, 1971: 195-196.
Fossarria (Bakerlymnaea) bulimoides
techella (Haldeman). Burch, 1980;
Walkerana, 1: 174; fig. 586 (shell).
TYPE LOCALITY.- Texas. Syntypes ANSP 59604 (4).
DISTRIBUTION
Fossaria (Bakerlymnaea) cubensis (Pfeiffer, 1839)
Limnaea
cubensis Pfeiffer,
1839; in Wiegm. Arch., 1: 354.- Strebel,
1873: 58; pl. 4, fig. 33.- Fischer &
Crosse, 1880: 50.- Pilsbry, 1891; Proc.
Acad. Nat. Sci. Phila., 55: 320.- Von
Martens, 1899; 379.
Lymnaea
cubensis (Pfeiffer). Hinkley, 1907; Nautilus, 21: 78.
Galba
cubensis
(Pfeiffer). F. C. Baker, 1911; Chicago
Acad. Sci. Sp. Pub., 3: 204-28; pl. 27, figs. 9-16 (shell).
Fossaria
(Bakerlymnaea) cubensis (Pfeiffer). Burch,
1980: 174; fig. 387 (shell).
TYPE LOCALITY.- Cuba.
DISTRIBUTION RECORDS.-
Fossaria (Bakerlymnaea) viator (Orbigny, 1835)
Limnaeus
viator Orbigny,
1835; Magazin de Zoologie, 1835: 24.
Lymnaea
viator Orbigny. Pilsbry, 1911;
Limnaeus
viatrix Orbigny,
1835; Voyage dans L’Amérique Méridional, Moll.: 340; pl. 43, figs. 1-3 (error
for Limnaeus viator Orbigny).
Lymnaea
viatrix (Orbigny). Paraense, 1982; Memorias do Instituto Oswaldo
Cruz, 77: 181-188.
Fossaria
viatrix
(Orbigny). Rangel-Ruiz, 1994; Walkerana,
7: 29-37; fig. 1 (shell).- Rangel-Ruiz,
1995; Malacological Review, 28: 71-79.
TYPE LOCALITY.- Restricted by
Pilsbry to banks of the Rio Negro, 7 or 8 leagues above the mouth, 41º S lat.,
Argentina.
DISTRIBUTION.- México and
Genus Pseudosuccinea F. C. Baker, 1908
Pseudosuccinea F. C. Baker, 1908; Science, n. s.,
27: 943.
TYPE SPECIES.- Lymnaea columella
Say, 1817.
DISTRIBUTION.- North America
south to
TAXONOMY.- A single species, Pseudosuccinea columlla, is recognized
in North and
Pseudosuccinea
columella (Say, 1817)
Lymnaea
columella Say,
1917; Jour. Acad. Nat. Sci., 1: 44.
Pseudosuccinea
columella (Say,
1817). F. C. Baker, 1911; Sp. Pub.
Chicago Acad. Sci., 3: 163-171; pl. 10, fig. B (reproductive anatomy); pl. 23,
figs. 8-30 (shell); pl. 23, figs. 1-4 (shell).
TYPE LOCALITY.- Not stated.
DISTRIBUTION.- The typical form
is found from
TAXONOMY.- F. C. Baker (1911)
lists four subspecies. He reports only Pseudosuccinea columella championi from
the study area.
Pseudosuccinea
columella championi (Von Martens, 1899)
Limnaea
columella var. championi Von Martens, 1899: 379; pl.
19, fig. 12 (shell).
Pseudosuccinea
columella championi
(Von Martens). F. C. Baker, 1911; Sp.
Pub. Chicago Acad. Sci., 3: 175-177; pl. 24, fig. 20 (shell).
TYPE LOCALITY.- Bigabo,
DISTRIBUTION RECORDS.- Central
Mexico south to
REMARKS.- The taxonomic status
and geographic distribution of this subspecies need to be investigated.
Genus Stagnicola Leach,1830
TYPE SPECIES.- Buccinum paluistris Müller, 1774.
DISTRIBUTION.- Circumboreal;
North America south to central
TAXONOMY.- Two subgenera are
recognized in the Nearctic Region. One
occurs in the study area. The number of
species in Stagnicola is
debatable. Stagnicola elodes is the only recognized in the study area. Lymnaeus
attenuatus is considered a synonym, but further study may reveal it to be a
distinct species.
Stagnicola elodes (Say,
1821)
Lymnaeus
elodes Say, 1821;
Jour. Acad. Nat. Sci. Phila., 2: 169.
Galba
elodes (Say). F. C. Baker, 1911; Chicago Acad. Sci. Sp.
Pub., 3: 322-327; pl. 30, figs. 32-34; pl. 34, figs. 11-19, 21-24 (shell).
Stagnicola
elodes (Say). Burch, 1980; Walkerana, 1: 176; figs.
600-606, 611 (shell).
Lymnaea elodes (Say). Paraense, 1994; Malacological Review, 27:
5-11; figs. 1-3 (shell), 4-7 (reproductive anatomy), 8-11 (digestive system),
12 (pallial organs), 13 (radula).
Lymnaeus decidiosus (Say). Pilsbry, 1904: Proc. Acad. Nat. Sci. Phila.,
55: 777.
Lymnaeus
attenuatus Say,
1829; New Harmony Dis., II: 244.
Lymnaea
attenuata (Say). Strebel, 1873: 57; pl. 5, figs. 32,
32a.- Fischer & Crosse, 1880:
49.- Von Martens, 1899: 375-377.- Pilsbry, 1904; Proc. Acad. Nat. Sci. Phila.,
55: 776.
Galba
attenuata
(Say). F. C. Baker, 1911; Chicago Acad.
Sci. Spec. Publ. 3: 353-355; pl. 37, figs. 33, 34 (shell).
TYPE LOCALITY.- Lymnaeus attenuatus: ditches and ponds
in the vicinity of Mexico City,
Mexico. Syntypes in the ANSP.
DISTRIBUTION RECORDS.- Interior
basin of Canada south to new England, California and New Mexico with outlying
populations in central Mexico. COAHUILA:
Saltillo (Pilsbry, 1904). HIDALGO: Zimapan
(Von Martens, 1899). MEXICO: Tlalpam
(Pilsbry, 1904). Lago de Chalco (Von
Martens, 1899). MORELOS: Laguna de
Quila, nr. Zempoala (Paraense, 1994).
Superfamily PLANORBOIDEA
Family ANCYLIDAE
Rafinesque, 1815
Subfamily
FERRISSIINAE
Genus Ferrissia Walker, 1908
Ferrissia Walker, 1908; Nautilus, 17:
15.- Basch, 1963; Bull. Mus. Comp.
Zool., 129: 126-128.
TYPE SPECIES.- Ancylus rivularis Say, 1817.
DISTRIBUTION.- North America and
the West Indies south to
TAXONOMY.- Numerous species have
been described. Basch (1963) places many
of these in synonymy. The taxonomic
status of the Middle American species remains to be determined. Two species are listed for the study area.
Ferrissia
bayacalifornica Walker, 1924
Ferrissia
bayacalifornica
Walker, 1924; Jour. Wash. Acad. Sci., 14: 431; fig. 1.- Wurtz, 1951; Nautilus, 64: 124.
Ferrissia
walkeri (Pilsbry
& Ferriss). Basch, 1963; Bull. Mus.
Comp. Zool., 129: 433.
TYPE LOCALITY.- A small pool 2.5
miles inland from San Jose del Cabo, Baja California Sur. Holotype in the UMMZ.
DISTRIBUTION
RECORDS.- BAJA CALIFORNIA: known only
from the type locality.
Ferrissia occidentalis
Walker, 1924
Ferrissia
occidentalis
Walker, 1924; Jour. Wash. Acad. Sci., 14: 431; fig. 2.
Ferrissia
walkeri (Pilsbry
& Ferriss). Basch, 1963; Bull. Mus.
Comp. Zool., 129: 433.
TYPE LOCALITY.- A small pool 2.5 miles inland from San Jose
del Cabo, Baja California Sur. Holotype
in the UMMZ.
DISRIBUTION RECORDS.- BAJA
CALIFORNIA SUR: known only from the
type locality.
Genus Gundlachia Pfeiffer, 1849
Gundlachia
Pfeiffer, 1849;
Zeitsch. für Malak., 6: 98.- Wurtz,
1951; Nautilus, 64: 126.
TYPE SEPCIES.- Gundlachia ancyliformis Pfeiffer, 1849.
DISTRIBUTION.- North America south
to Guatemala and Honduras.
TAXONOMY.- Wurtz (1951)
recognizes seven species.
Gundlachia hinkleyi
Walker, 1917
Gundlachia
hinkleyi Walker,
1917; Nautilus, 31: 51-53; pl. 1, figs. 10-16 (shell); pl. 3, fig. 1
(radula).- Hinkley, 1920; Nautilus, 34:
38, 40.- Wurtz, 1951; Nautilus, 64: 126.
TYPE LOCALITY.- Maya Farms,
Quirigua, Dept. Izabal, Guatemala.
Holotype UMMZ 43455.
DISTRIBUTION RECORDS.- GUATEMALA,
Dept. Guatemala: Lago de Amatitlan
(Hinkley, 1920). Dept. Izabal: Quirigua.
Gundlachia hjalmarsoni
Pfeiffer, 1858
Gundlachia
hjalmarsoni
Pfeiffer, 1859; Malak. Blätt., 5: 197.- Von
Martens, 1899: 403.- Wurtz, 1951;
Nautilus, 64: 126.
TYPE LOCALITY.- Santa Rosa,
Honduras.
DISTRIBUTION RECORDS.- HONDURAS: known only from the type locality.
Genus Hebetancylus
Pilsbry, 1914
Hebetancylus Pilsbry, 1914; proc. Acad. nat.
Sci. Phila., 65: 671.- Wurtz, 1951;
Nautilus, 64: 126.- Basch, 1963; Bull.
Mus. Comp. Zool., 129: 422.
TYPE SPECIES.- Ancylus moricandi Orbigny, 1837.
DISTRIBUTION.- Cuba and
Hispaniola in the Wst Indies; Florida and Texas south to Brazil and Paraguay.
TAXONOMY.- Wurtz lists eight
species in the genus. Two occur in the
study area.
Hebetancylus
excentricus (Morelet, 1851)
Ancylus
excentricus Morelet,
1851; Test. Noviss., II: 17.- Fischer
& Crosse, 1886: 37; pl. 30, figs. 16, 16a (shell).- Pilsbry, 1891; Proc. Acad. Nat. Sci. Phila.,
43: 3.- Von Martens, 1899:
402-403.- Pilsbry, 1904; Proc. Acad.
Nat. Sci. Phila., 55: 777.
Velletia
excentrica (Morelet). Tate, 1970; Amer. Jour. Conch., 5: 158.
Laevapex
excentricus
(Morelet). Hinkley, 1920; Nautilus, 34:
39, 40.- Walker, 1923; The Ancylidae of
South Africa: 10.- Bequaert &
Clench, 1936; Pub. Carnegie Inst. Wash., (457): 70.
Ferrissia
excentrica
(Morelet). Wurtz, 1951; Nautilus, 64:
125.- Solem, 1954; Nautilus, 68: 6-7.
Gundlachia
excentrica
(Morelet). Taylor, 1966; Veliger, 9:
206.
Hebetancylus
excentricus (Morelet). Basch, 1963; Bull. Mus. Comp. Zool., 129:
422-426; fig. 9 (shell), fig. 10 (anatomy).-
Burch, 1980; Walkerana, 1: 214; figs. 762, 769 (shell).
Ancylus
excentricus var. biolleyi Von Martens, 1899: 402.
Ancylus
excentricus var. pittieri Von Martens, 1899: 402.
TYPE LOCALITY.- Lago de Itza
[Petén], Dept. Petén, Guatemala.
DISTRIBUTION RECORDS.- Florida
and Texas south to Costa Rica. COSTA
RICA, Prov. San Jose: Rio Torres; San
Jose (Von Martens, 1899). Prov. Puntarenas: Rio de Las Platanales
(Von Martens, 1899). NICARAGUA, Dept. Chontales: San Nicolas, San
Agustin (Tate, 1870). GUATEMALA, Dept. Guatemala: lago de Amatitlan
(Hinkley, 1920). Dept. Izabal: Quirigua (Hinkley, 1920). Dept.
Petén: Lago de Itza. COAHUILA: Rio
Salado de los Nadadores, at El Cariño de la Montaña, 20 mi. E of Cuatro
Cienegas (Taylor, 1966). DURANGO: Laguna
del Progreso (Solem, 1944). MICHOACÁN:
pressa nr. Uruapan (Pilsbry, 1903).
YUCATAN: Shkolak (Pilsbry, 1891); Xanaba Cenote Grande, 8 mi. SW of
Chichen Itza; Chotch Cenote 2 mi. NE of Pista (Bequaret & Clench, 1936).
Hebetancylus
providentialis Wurtz, 1951
Hebetancylus
providentialis
Wurtz, 1951; Nautilus, 64: 127-128; pl. 4, figs. 8, 8a.
TYPE LOCALITY.- HFLMNHfington’s
Creek, Old Providence Island, Colombia.
Holotype ANSP 186643a.
DISTRIBUTION RECORDS.- COLOMBIA, Old Providence Island: known only from
the type locality.
Genus Laevapex Walker, 1908
Laevapex Walker, 1908; Nautilus, 1908, 17:
15.- Wurtz, 1951; Nautilus, 64:
125.- Basch, 1963; Bull. Mus. Comp.
Zool., 129: 418.
TYPE SPECIES.- Ancylus fuscus C. B. Adams, 1841.
DISTRIBUTION.-
North America and the West Indies south to Panama.
TAXONOMY.- Approximately ten
species are recognized in the literature.
Four occur in the study area.
Laevapex aguadae
(Goodrich & van der Schalie, 1937)
Ferrissia
(Laevapex) aguadae Goodrich & van der Schalie, 1937; Misc. Pub. Mus.
Zool. Univ. Mich., (34): 34-35; pl. 1, figs. 7, 7a (shell).
TYPE LOCALITY.- Aguada de Copó, 1
mile south of La Libertad, Dept. Petén, Guatemala. Holotype in the UMMZ.
DISTRIBUTION RECORDS.- GUATEMALA:
Goodrich & van der Schalie report this species from seven unspecified
localities.
Laevapex joseana
(Morrison, 1946)
Ferrissia (Laevapex) joseana
Morrison, 1946; Smiths. Misc. Coll., 106: 39; pl. 1, figs. 5, 6 (shell).- Wurtz, 1951; Nautilus, 64: 129.
TYPE LOCALITY.- Flood-plain pools
along the middle reaches of a stream opening into the northwest mangrove swamp
on San José Island, Archipelago de las Perlas, Panama. Holotype USNM 542179.
DISTRIBUTION RECORDS.- PANAMA:
known only from the type locality.
Laevapex papillaris
(Von Martens, 1899)
Ancylus
papillaris Von
Martens, 1899: 402; pl. 21, figs. 11, 11a, 12, 12a (shell).- Pilsbry, 1904; Proc. Acad. Nat. Sci. Phila.,
55: 777.- Wurtz, 1951; Nautilus, 64:
130.
TYPE LOCALITY.- Rio Ameca,
Jalisco.
DISTRIBUTION RECORDS.- JALISCO:
Rio Ameca. MEXICO: Tlalpam. MICHOACÁN: Uruapan. NUEVO LEON: Monterey (Pilsbry, 1904).
Laevapex sallei
(Bourguignat, 1857)
Ancylus
sallei Bourguignat,
1857; Aménités Malac., ii: 32.-
Bourguignat, 1857; Rev. et Mag. Zool., 1857: 16.- Strebel, 1873: 63; pl. 4, fig. 35
(shell).- Fischer & Crosse, 1886:
38; pl. 30, figs. 17, 17a, 17b (shell).-
Von Martens, 1899: 401.
TYPE LOCALITY.- Laguna Larga de
Toxpan, near Cordoba, Veracruz.
DISTRIBUTION RECORDS.- VERACRUZ:
Veracruz (Srebel, 1873).
Genus Uncancylus Pilsbry, 1914
Uncancylus Pilsbry, 1914; Proc. Acad. Nat.
Sci. Phila., 65: 671.- Wurtz, 1951;
Nautilus, 64: 128-129.
TYPE SPECIES.- Ancylus barilensis Moricand, 1845.
DISTRIBUTION.- Guadeloupe,
Jamaica, Costa Rica south to Brazil and Argentina.
TAXONOMY.- Wurtz (1951) lists
seven species and one subspecies. Two
occur in the study area.
Uncancylus ameliae
Pilsbry, 1920
Uncancylus
ameliae Pilsbry,
1920; Proc. Acad. Nat. Sci. Phila., 72: 9; lower and right text-fig. 4 (shell).
TYPE LOCALITY.- Rio Zapote, at
confluence with the Rio Reventazón, Cartago Prov., Costa Rica; 3450 ft.
alt. Holotype ANSP 105260.
DISTRIBUTION RECORDS.- COSTA
RICA: known only from the type locality.
Uncancylus calverti
Pilsbry, 1920
Uncancylus
calverti Pilsbry,
1920; Proc. Acad. Nat. Sci. Phila., 72: 7-9; left and upper text-fig. 4
(shell); text-fig. 5 (radula).
TYPE LOCALITY.- Brook near Rio
Reventazón, Juan Viñas, Cartago Prov., Costa Rica; 2500 ft. alt. Holotype ANSP 105277.
DISTRIBUTION RECORDS.- COSTA
RICA: known only from the type locality.
Undetermined
species of Ancylidae
Ancylus haldemani
Boettger, 1853
Ancylus
depressus Boettger,
1844; Monogr.: 4 (non Ancylus depressus Deshayes, 1824).
Ancylus
haldemani Boettger,
1853; Jour. de Conchyl., 4: 180.-
Binney, 1865; land & Freshwater Shells N. Amer., 2: 141; fig.
236..- Wurtz, 1951; Nautilus, 64: 130.
TYPE LOCALITY.- Mexico.
DISRTRIBUTION RECORDS.- Unknown.
Ancylus sallei
Boettger, 1857
Ancylus
sallei Boettger,
1857; Rev. Mag. Zool., 9: 16.- Wurtz,
1951; Nautilus, 64: 131.
TYPE LOCALITY.- Mexico and Cuba.
DISTRIBUTION RECORDS.- Unknown
Family PLANORBIDAE Rafinesque,
1815
Subfamily
PLANORBINAE Rafinesque, 1815
Genus Gyraulus Charpentier, 1837
Gyraulus Charpentier, 1837; Mem. Soc. Helv.
De Sci. Natur., 1: 21.
TYPE SPECIES.- Planorbis hispidus Draparnaud (= Planorbis albus O. F. Múller, 1774).
DISTRIBUTION.- Europe, Africa,
Asia, the Philippines, Australia, Tasmania, New Caledonia, Fiji and North
America.
TAXONOMY.- Four subgenera are
recognized (Zilch, 1959). One occurs in
the study area.
Subgenus Torquis Dall, 1905
Torquis Dall, 1905; Harriman Alaska Exp.,
XIII: 83, 86.
TYPE SPECIES.- Planorbis parvus Say, 1817.
DISTRIBUTION.- North America,
northern Asia, Europe.
TAXONOMY.- Three species occur in
North America (Burch, 1980). A single
species occurs in the study area. A
second species is placed here provisionally.
Gyraulus
(Torquis) parvus (Say, 1817)
Planorbis
parvus Say, 1817;
in Nicholson’s Encyclop., 2: pl. 1, fig. 5 (shell).- Pilsbry, 1891; Proc. Acad. Nat. Sci. Phila.,
43: 322.
Planorbis
(Gyraulas) parvus Say. Von Martens,
1899: 394.
Gyraulus
parvus (Say). Branson, McCoy & Sisk, 1964; Southwestern
Naturalist, 9: 103-104.- Bequaert &
Miller, 1973: 204-205.- Burch, 1980;
Walkerana, 1: 194; fig. 709 (shell).
TYPE LOCALITY.- Delaware River,
eastern North America.
DISTRIBUTION RECORDS.- North
America from Alaska and northern Canada to the Greater Antilles and south in
Mexico at least to Morelos. MORELOS:
Yautepec (Pilsbry, 1891). SONORA: Rio
Nacuzari, 7 mi. S of Nacuzari (Branson, et al, 1964).
Gyraulus “percarinatus” Paraense, 2000
Gyraulus
percarinatus
Paraense, 2000; Malac. Rev., 33/34: 16-19; fig. 11A (shell), fig. 11B
(reproductive anatomy), fig. 12 (radula).
TYPE LOCALITY.- Grounds of the
Faculty of Agronomy, Panama University of Tocumen, Panama Prov., Panama. No type specimens designated.
DISTRIBUTION RECORDS.- PANAMA:
known only from the type locality.
TAXONOMY.- The name Gyraulus percarinatus Paraense, 2000 is
invalid, because the description does not include the designation of a holotype
as required by ICZN Articles 16.4 and 62.4.
Subfamilly DREPANOTREMINAE
Zilch, 1959
DISTRIBUTION.- American
subtropics and tropics from Texas and the Greater Antilles south to Argentina.
TAXONOMY.- Three are recognized, Antillorbis, Drepanotrema, and a
Brazilian genus, Acrorbis Ohdner,
1937.
Genus Antillorbis Harry & Hubendick 1964
Antillorbis Harry & Hubendick, 1964;
Göeborgs Kungl. Vetenskaps-och Vitterhets-samhölles handlingar, ser. 6, ser. B:
29-30.
TYPE SPECIES.- Planorbis circumlineatus Shuttleworth,
1854 (= Planorbis aerugnosus Morelet,
1851).
DISTRIBUTION.- The west Indies,
Mexico and Central America.
TAXONOMY.- A single species is
recognized.
Antillorbis
aeruginosus (Morelet, 1851)
Planorbis
aeruginosus Morelet,
1851; Test. Noviss., II: 15.
Gyraulus
aeruginosus
(Morelet). Goodrich & van der
Schalie, 1937; Misc. Pub. Mus. Zool. Univ. Mich., (34): 33.
Antillorbis
feruginosus (Morelet). Harry & Hubendick, 1964; Göeborgs Kungl.
Vetenskaps-och Vitterhets-samhölles handlingar, ser. 6, ser. B: 30-33; figs.
90-92, 114-117 (shell), fig. 25(jaw), fig. 26 (radula), figs. 27-30 (anatomy).
Drepanotrema
(Antillorbis) aeruguinosus (Morelet).
Bequaert & Miller, 1973: 209.-
Burch, 1980; Walkerana, 1 (3): 194, 196; fig. 710 (shell).
Planorbis
salleanus Dunker,
1853; Proc. Zool. Soc. London, 54.
Planorbis
circumlineatus Shuttleworth,
1854; Ann. Lyc. Nat. Hist. N. Y., 6: 72.-
Shuttleworth, 1854; Mill. Naturf. Ges. Bern, 1854: 96.
Planorbaria
circumlineatus
(Shuttleworth). Neubert & Gosteli,
2003; Contributions in Natural History, 1: 17; pl. 1, figs. 1.
Planorbis
lineatus sonorensis
Cooper, 1893; Proc. Calif. Acad. Sci. (2), 3: 343; pl. 14, figs. 10a-d (shell).
Planorbis
filocinctus Pilsbry
& Ferriss, 1906; proc. Acad. Nat. Sci. Phila., 58: 156; pl. 9, figs. 1-3
(shell). (not Planorbis filocinctus von Sandberger, 1875).
Planorbis
arizonensis Pilsbry
& Ferriss, 1915; Proc. Acad. Nat. Sci. Phila., 67: 390 (replacement name
for Planorbis filicinctus P. &
F.).
Gyraulus
arizonensis
(Pilsbry & Ferriss). Richards, 1937;
Trans. Amer. Philos. Soc., 77: 255.
Planorbis
santacruzensis
Germain, 1932; Rec. Indian Museum, 21: 138; figs. 18-21 (shell).
TYPE LOCALITIES.- Planorbis aeruginosis: Lago de Izabal,
Guatemala. Planorbis salleans: Santo Domingo (= Hispaniola). Planorbis
circumlineatus: Humacoa, Puerto Rico.
Syntypes Naturhistorische Museum Bern 18952/16 (Neubert & Gosteli,
2003: 17). Planorbis lineatus sonorensis: San Miguel, Sonora. Planorbis
filicinctus: drift of San Pedro River, Benson, Cochis Co., Arizona. Planorbis santacruzensis: St. Croix.
DISTRIBUTION RECORDS.- West
Indies, southern Texas and Arizona south to Panama. GUATEMALA, Dept. Guatemala: outlet of Lago Amatitlan, nr. Cd. Guatemala
(Goodrich & van der Schalie, 1937). Dept. Izabal: Lago de Izabal (Morelet,
1851). QUINTANA ROO: Cozumel Island,
swamp nr. San Miguel; swamp nr. San Gerbacio (Richards, 1937). SONORA: San Miguel (Cooper, 1893).
Genus Drepanotrema Fischer & Crosse, 1880
Drepanotrema Fischer & Crosse, 1880: 59,
75.- H. B. Baker, 1930; Occ. Pap. Mus.
Zool. Univ. Mich., (210): 48-51.-
Pilsbry, 1934; Proc. Acad. Nat. Sci. Phila., 86: 58-59.- Harry & Hubendick, 1964; Meddel. Gotenb.
Mus. Zool. Avdel., 136: 26-29.
TYPE SPECIES.- Planorbis yzabelensis Crosse &
Fischer, 1879 (= Planorbis anatinus
Orbigny, 1835).
DISTRIBUTION.- Tropical America
from Texas and the west Indies south to Argentina.
TAXONOMY.- Two subgenera are
recognized.
Subgenus Drepanotrema Fischer & Crosse, 1880
DISTRIBUTION.- Same as for the
genus.
TAXONOMY.- F. C. Baker (1945:
118) lists eight species. Two occur in
the study area.
Drepanotrema (Drepanotrema) anatinum (Orbigny, 1835)
Planorbis
anatinus Orbigny,
1835; Mag. De Zool., 5 (62): 28.
Drepanotrema
anatinum
(Orbigny). Aguayo, 1933; Nautilus, 47:
64-68.- F. C. Baker, 1945; Moll. Fam.
Planorbidae: 118; pl. 9, figs. 1-3 (reproductive anatomy); pl. 68, fig. 3
(radula); pl. 79, figs. 16-18 (shell); pl. 124, figs. 1-13, 30 (shell).- Harry & Hubendick, 1964; Meddel. Gotenb.
Mus. Zool. Avdel., 136: 21, figs. 87-89, 118-120 (shell).- Paraense, 2000; Malac. Rev., 33: 12-13; figs.
7A (shell), 7B (reproductive anatomy}.
Planorbis
haldemani C. B.
Adams, 1849; Contrib. to Conch.: 43.
Planorbis
aracasnsis
Gundlach, 1857; Malak Blätt. 4: 179.
Planorbis
esperanzensis Tryon,
1866; Amer. Jour. Conch., 2: 10; pl. 2, figs. 11-13.
Planorbis
yzabalensis Crosse
& Fischer, 1879; Jour. de Conchyl., 342.-
Fischer & Crosse, 1880: 75; pl. 33, figs. 2, 2a-c (shell).
Planorbis
(Hippeutus) yzabalensis (Crosse &
Fischer). Von Martens, 1899: 397-398.
Anisus (Gyraulus) lenzi Benthem
Jutting, 1943; Archiv für Hydrobiologie, 39: 460.
TYPE LOCALITY.- Planorbis anatinus: Rio Parana,
Argentina; syntype in the BMNH (Aguayo, 1933).
Planorbis haldemani:
Jamaica. Planorbis esperanzensis: Esperanza, Cuba. Planorbis
aracasnsis: Trinidad, Cuba. Planorbis yzabalensis: Lago de Izabal,
Guatemala. Anisus (Gyraulus) lenzi: Rio São Francisco, San Pedro
Dias-Buchr, Pernambuco, Brazil.
DISTRIBUTION RECORDS.- Widely
distributed throughout the West Indies and from Texas south to Argentina and
Brazil. PANAMA, Canal Zone: Gamboa (Aguayo, 1933).
Prov. Panama: ditch at the National University, Tocumán (Paraense,
2000). COSTA RICA, Prov. Guanacaste: Catalina (Paraense, 2000). NICARAGUA, Dept. Rivas: Rivas (Paraense,
2000). GUATEMALA, Dept. Izabal: Lago de Izabal, Mariscos; El Prado (Paraense,
2000). Dept. Santa Rosa: Rio Helado, Taxisco. BELIZE, Belize Dist.: Boston Village (Paraense, 2000). TABASCO: Rio Usumacinta; Balancan (Von Martens,
1899).
Drepanotrema (Drepanotrema) lucidum (Pfeiffer, 1839)
Planorbis
lucidus Pfeiffer,
1839; in Weigmanns, Arch. für
Naturgesch.: 354.
Planorbis
(Dreopanotrema) lucidus Pfeiffer. Bequaert
& Clench, 1936; Pub. Canegie Inst. Wash., (457): 67-68.
Drepanotrema
lucidum
(Pfeiffer). H. B. Baker, 1930; Occ. Pap.
Mus. Zool. Univ. Mich., (210): 48-49; pl. 28, fig. 9 (radula).- F. C. Baker, 1945; Moll. Fam. Planorbidae:
118; pl. 11, figs. 1-7 (reproductive anatomy); pl. 46, fig. 16 (kidney); pl.
48, fig. 11 (stomach); pl. 50, fig. 27 (jaw); pl. 68, fig. 2 (radula); pl. 78,
figs. 11-13 (shell); pl. 124, figs. 29, 30, 32 (shell); pl. 125, figs. 1-17
(shell).- Harry & Hubendick, 1964;
Meddel. Gotenb. Mus. Zool. Avdel., 136: 22-23; figs. 84-86 (shell), figs. 121-123
(shell).- Paraense, 2000; Malac. Rev.,
33/34: 16; fig. 9A (shell), fig. 9B (reproductive anatomy).
TYPE LOCALITY.- Cuba.
DISTRIBUTION RECORDS.- The West
Indies and Mexico south to Brazil and Peru.
NICARAGUA, Dept. Grenada:
Grenada. Dept. Rivas: Rivas (Paraense, 2000). GUATEMALA, Dept. Izabal: Mariscos (Paraense, 2000). BELIZE, Belize
Dist.: pond by rd. to International Airport (Paraense, 2000). YUCATAN: Aguado Halal nr. Merida; aguado 14
km N, 2 km E of Merida; Aguado Daadz, 6.5 mi. SW of Chichen Itza (Bequaert
& Clench, 1936).
Subgenus Fossulorbis Pilsbry, 1934
Fossulorbis Pilsbry, 1934; Proc. Acad. Nat.
Sci. Phila., 86: 59.- F. C. Baker, 1945;
Moll. Fam. Planorbidae: 118.
TYPE SPECIES.- Planorbis cultratus Orbigny, 1841.
DISTRIBUTION.- The West Indies
and southern Texas south to Argentina.
TAXONOMY.- Six species and four
subspecies occur in the study area.
Drepanotrema
(Fossulorbis) cimex (Moricand, 1839)
Planorbis
cimex Moricand,
1839; Mèm. Soc. Phys. Hist. Nat., Genève, 8: 143; pl. 3, figs. 8-9 (shell).
Drepanotrema (Fossulorbis) cimex
(Moricand). F. C. Baker, 1945; Moll.
Fam. Planorbidae: 148; pl. 127, figs. 1-3, 7, 8, 16 (shell); pl. 128, figs.
17-22 (shell). - Harry & Hubendick,
1964; Meddel. Gotenb. Mus. Zool. Avdel., 136: 24-28; figs. 81-83, 14-16, 130
(shell).- Burch, 1980; Walkerana, 1:
198; fig. 715 (shell).
TYPE LOCALITY.- Bahia, Brazil.
DISTRIBUTION RECORDS.- Known from
southern Texas, Mexico, Central America, the West Indies, Venezuela and Brazil.
Drepanotrema
(Fossulorbis) cultratum cultratum (Orbigny, 1841)
Planorbis
cultratus Orbigny,
1841; in Sagra, Hist. Fis. Polit. y nat. de Cuba, Moluscos: 196; pl. 14, figs.
5-8.- Fischer & Crosse, 1880: 68;
pl. 32, figs. 7, 7a-c (shell).- Von Martens,
1899: 395-396.- Hinkley, 1920; Nautilus,
34: 38.
Planorbis
(Spiralina) cultratus Orbigny. Bequaert
& Clench, 1936; Pub. Canegie Inst. Wash., (457): 67.
Drepanotrema
cultratum
(Orbigny). H. B. Baker, 1930; Occ. Pap.
Mus. Zool. Univ. Mich., (210): 51; pl. 29, fig. 8 (radula).- Pilsbry, 1934; Proc. Acad. Nat. Sci. Phila.,
86: 60; pl. 11, figs. 7-8b (shell).-
Richards, 1937; Trans. Amer. Philos. Soc., 77: 255.- F. C. Baker, 1945; Moll. Fam. Planorbidae:
118, 388, 392; pl. 76, fig. 10 (reproductive anatomy); pl. 78 figs., 17-19
(shell).
TYPE LOCALITY.- Martinique
(Pilsbry, 1934: 60).
DISTRIBUTION RECORDS.- The Lesser
Antilles, Venezuela and Colombia.
NICARAGUA, Dept. Chontales:
San Agustin, nr. Acoyapa (Von Martens, 1988).
GUATEMALA, Dept. Guatemala:
ditch along railroad at Laguna, Lago Amatitlan (Hinkley, 1920). QUINTANA ROO: swamp a few miles N of San
Miguel (Richards, 1937). YUCATAN: Xanaba
Cenote Grande, 8 km SW of Chichen Itza (Bequaert & Clench, 1936).
Drepanotrema
(Fossulorbis) cultratum anitense (Cooper, 1893)
Planorbis
(Anisus) anitensis Cooper, 1893; Proc. Calif. Acad. Sci., ser. 2, 3:
341-342; pl. 14, fig. 8 (shell).
Drepanotrema
cultratum anitense
(Cooper). F. C. Baker, 1945; Moll. Fam.
Planorbidae: 118, 488; pl. 126, figs. 17-19 (shell).
TYPE LOCALITY.- Santa Anita, Baja
California Sur.
DISTRIBUTION RECORDS.- BAJA
CALIFORNIA SUR: lagoon, Santa Rita (F. C. Bater, 1945).
Drepanotrema
(Fossulorbis) cultratum dueñasianum (Tristram, 1861)
Planorbis
dueñasensis Tristram,
1861; Proc. Zool. Soc. London: 232.
Drepanotrema
cultratum dueñasianum
(Tristram). Pilsbry, 1934; Proc. Acad.
Nat. Sci. Phila., 86: 60; pl. 11, figs. 6, 6a, 6b (shell).- F. C. Baker, 1945; Moll. Fam Planorbidae:
118, 490; pl. 17, figs. 9-12 (shell).
TYPE LOCALITY.- Lago de Dueñas,
Dept. Sacatepequez, Guatemala.
DISTRIBUTION RECORDS.- GUATEMALA,
Dept. Guatemala: pools along railroad
nr. Lago de Amatitlan (Pilsbry, 1934). Dept. Sacatepequez: Lago de Dueñas.
Drepanotrema
(Fossulorbis) cultratum labrosum Pilsbry, 1934
Drepanotrema
cultratum labrosum
Pilsbry, 1934; Proc. Acad. Nat. Sci. Phila., 86: 61; pl. 11, figs. 9-11
(shell).- F. C. Baker, 1945; Moll. Fam.
Planorbidae: 118; 492; pl. 128, figs. 6-10 (shell).
TYPE LOCALITY.- Brownsvile,
Texas. Holotype ANSP 143416.
DISTRIBUTION RECORDS.- Southern
Texas south to southern Tamaulipas.
TAMAULIPAS: drift debris of the Rio Panuco (Pilsbry, 1934).
Drepanotrema
(Fossulorbis) cultratum panuco Pilsbry, 1934
Drepanotrema
cultratum panuco Pilsbry,
1934; Proc. Acad. Nat. Sci. Phila., 86: 60-61; pl. 11, figs. 4-5a
(shell).- F.C. Baker, 1945; Moll. Fam
Planorbidae: 118, 492; pl. 128, figs. 3-5 (shell).
TYPE LOCALITY.- Pasture west of
San Dieguito, San Luis Potosi.
Holotype ANSP 107619.
DISTRIBUTION RECORDS.- SAN LUIS
POTOSI: San Dieguito. TAMAULIPAS:
Tampico (Pilsbry, 1934).
Drepanotrema
(Fossulorbis) depressissimum (Moricand, 1839)
Planorbis
depressissimus
Moricand, 1839; Mém. Soc. Phys. Hist. Nat. Genèva, 8: 143-144; pl. 3, figs. 10,
11 (shell).
Drepanotrema
depressissimum
(Moricand). F. C. Baker, 1945; Moll.
Fam. Planorbidae: 118; pl. 127, figs. 17-19 (shell).- Paraense, 2000; Malac. Rev., 33/34: 14-15;
fig. 8A (shell), fig. 8B (reproductive anatomy).
TYPE LOCALITY.- Bahia, Brazil.
DISTRIBUTION RECORDS.- Widely
distributed in South America, Central America and the West Indies (Paraense,
2000). COSTA RICA, Prov. Guanacaste: Catalina (Paraense, 2000). NICARAGUA, Dept. Chontales: Acoyapa. Dept.
Rivas: Rivas (Paraense, 2000).
Drepanotrema
(Fossulorbis) kermatoides (Orbigny, 1835)
Planorbis
kermatoides
Orbigny, 1835; Mag. de Zool., (62): 27.
Drepanotrema (Fossulorbis) kermatoides
(Orbigny). F. C. Baker, 1945; Moll.
fam. Planorbidae: 148; pl. 126, figs. 13-16 (shell).- Harry & Hubendick, 1964; Meddel. Gotenb.
Mus. Zool. Avdel., 136: 23-24; figs. 78-80, 137-129 (shell).- J. B. Burch, 1980; Walkerana, 1: 196; figs.
711 (shell).- Rehder, 1966; Proc. Biol.
Soc. Wash., 79: 280.
TYPE LOCALITY.- Providence of
Lima, Peru.
DISTRIBUTION RECORDS.- Lesser
Antilles, Texas, Mexico, Central America, Venezuela, Peru and Brazil (Harry
& Hubendick, 1964). QUINTANA ROO:
Isla Cozumel (Rehder, 1966).
TAXONOMY.- Paraense (2000)
suggests that records for Drepanotrema
kermaoides from Mexico and Central America may pertain to D. depressissimum.
Drepanotrema
(Fossulorbis) sumichrasti (Crosse & Fischer, 1879)
Planorbis
sumichrasti Crosse
& Fischer, 1879; Jour. de Conchyl., 27: 342.- Fischer & Crosse, 1880: 69; pl. 33, figs.
6, 6a-d (shell).- Von Martens, 1899:
396.
Drepanotrema
sumichrasti (Crosse
& Fischer). F. C. Baker, 1945; Moll.
Fam. Planorbidae: 118; pl. 12, figs. 11-16 (shell).
TYPE LOCALITY.- Cacoprieto,
Isthmus of Tehuantepec, Oaxaca.
DISTRIBUTION RECORDS.- PANAMA, Canal Zone: Barro Colorado Island (F. C. Baker,
1945). OAXACA: Cacoprieto.
Drepanotrema
(Fossulorbis) surinamense (Clessin, 1884)
Planorbis surinamensis Clessin,
1884; in Martini & Chemnitz; System. Conch. Cab., Ed. 2, Planorbis: 126; pl. 17, fig. 11.
Drepanotrema
surinamense
(Clessin). Paraense, 2000; Malac. Rev.,
33/34: 16; fig. 10A (shell), Fig. 10B (reproductive anatomy).
TYPE LOCALITY.- Surinam.
DISTRIBUTION RECORDS.- PANAMA, Prov. Panama: Juan Diaz, ca. 6 km NE of
Ciudad Panama; Las Cumbres; Hippodrome Presidente Remón; Tocumen (Paraense,
2000). COSTA RICA: Prov. Guanacaste: Catalina (Paraense, 2000).
Subfamily BIOMPHALARIINAE
H. Watson, 1954
Genus Biomphalaria Preston, 1910
Biomphalaria Preston, 1910; Ann. Mag. Nat. Hist.,
ser. 8, 6: 533.- Paraense &
Deslandes, 1958; Rev. Brasil. Biol., 18: 65-70.- IZCN Opinion 735, 1965.
Tropicorbis
Pilsbry & Brown, 1914; Proc. Acad. Nat.
Sci. Phila., 66: 212.
Platytaphius
Pilsbry, 1924;
Proc. Acad. Nat. Sci. Phila., 78:
Australorbis Pilsbry, 1934; Proc. Acad. Nat.
Sci. Phila., 86: 55-56.
Obstructio Haas, 1939; Field Mus. Nat. Hist.,
Zool. Ser., 24: 100.
Lateorbis F. C. Baker, 1945; Moll. Fam.
Planorbidae: 85.
TYPE SPECIES.- Biomphalaria: Biomphalaria smithi Preston, 1910.
Platytaphius: Planorbis hertopleurus Pilsbry &
Vanatta, 1896. Tropicorbis: Planorbis
liebmanni Philippi, 1850 (= Planorbis
maya Morelet, 1849; = Planorbis
havanensis Pfeiffer, 1839, fide
Paraense & Deslandes, 1958). Australorbis: Planorbis glabratus Say, 1818.
Obstructio: Planorbis janeirensis Clesson,
1884. Lateorbis: Planorbis pallidus C. B. Adams, 1846.
DISTRIBUTION.- Africa, Tropical
America.
TAXONOMY.- The systematics of
this genus is very unsettled at present.
There is a large number of species in Africa and tropical America. The genus has received a great deal of study
because of its importance as intermediate hosts for human schistosome
parasites. However, many species not
directly implicated as schistosome hosts remain poorly studied. Subdivision of Biomphalaria into subgenera is not plausible at present.
Biomphalaria albicans
(Pfeiffer, 1848)
Planorbis
albicans Pfeiffer,
1839; Weigmanns Arch fúr Naturgesch.: 354.
Taphius
albicans
(Pfeiffer). Harry & Hubendick, 1964;
Göeborgs Kungl. Vetenskaps-och Vitterhets-samhölles handlingar, ser. 6, ser. B:
50-52; figs. 105-107, 149-155 (shell).
Biomphalaria
albicans (Pfeiffer). Abdel-Malek, 1969; Malacologia, 7: 190, 198,
200; fig. 8 (reproductive anatomy), figs. 17a, 17b, 18 (shell).
TYPE LOCALITY.- Cuba.
DISTRIBUTION RECORDS.- Reported
from the Greater Antilles, some Lesser Antilles, Louisiana, Texas, Mexico and
Central America (Harry & Hubendick, 1964).
COSTA RICA, Prov. Cartago: nr.
Turrialba (Abdel-Malek, 1969). BELIZE: Belize Dist.: Central Farms
(Abdel-Malek, 1969).
TAXONOMY.- Paraense (1996a)
treats this species as a synonym of Biomphalaria
helophila (Orbigny, 1835).
Biomphalaria
belizensis (Crosse & Fischer, 1878)
Planorbis
belizensis Crosse
& Fischer, 1879; Jour. de Conchyl., 27: 342.- Fischer & Crosse, 1880: 68; pl. 32, figs.
6, 6a, 6b (shell).- Von Martens, 1899:
390.
Biomphalaria
belizensis (Crosse
& Fischer, 1878).
TYPE LOCALITY.- Belize.
DISTRIBUTION RECORDS.- GUATEMALA,
Dept. Alta Verapaz: Coban; San Miguel
Tucarú (Fischer & Crosse, 1880).
BELIZE. TABASCO (Von Martens,
1899).
Biomphalaria
boucardianus (Preston, 1907)
Planorbis
boucardianus
Preston, 1907; Ann. & Mag. Nat. Hist., ser. 2: 20; 497, text-fig. 17
(shell).
Tropicorbis
boucardianus (Preston). F. C. Baker, 1945; Moll. Fam. Planorbidae:
85, 900, 506; pl. 132, figs. 31, 32 (shell); pl. 135, figs. 17-19 (shell).
TYPE LOCALITY.- Mexico.
DISTRIBUTION RECORDS.- NARARIT:
Manzanillo (F. C. Baker, 1945).
Biomphalaria fieldi
(Tryon, 1863)
Planorbis
fieldi Tryon, 1863;
Proc. Acad. Nat. Sci. Phila., 15: 281; pl. 1, figs. 4, 5 (shell).- Tate, 1870; Amer. Jour. of Conch., 5: 159.
Planorbis (Gyraulus) fieldi
Tryon. Von Martens, 1899: 394-395; pl.
21, fig. 6 (shell).
Tropicorbis
fieldi (Tryon). F. C. Baker, 1945; Moll. Fam. Planorbidae:
84, 500; pl. 132, figs. 24-30 (shell).
Biomphalaria
fieldi (Tryon). Abdel-Malek, 1969; Malacologia, 7: 198; fig.
7 (reproductive anatomy), fig. 17 (shell).
TYPE LOCALITY.- Panama.
DISTRIBUTION RECORDS.- PANAMA, Canal Zone: Gatún (F. C. Baker, 1945;
Abdel-Malek, 1969). NICARAGUA, Dept. Grenada: Lago de Nicaragua,
Grenada (Tate, 1870).
Biomphalaria
gracilenta (Gould, 1855)
Planorbis
gracilentus Gould,
1855; Proc. Boston Soc. Nat. Hist., 5: 129.
Tropicorbis (Tropicorbis) gracilentus
(Gould). F. C. Bakeer, 1945; Moll. Fam.
Planorbidae: 85, 496, 506; pl. 130, figs. 1-12 (shell); pl. 135, figs. 32-35
(shell).
TYPE LOCALITY.- Great Colorado
Desert lowlands, United States. Holotype
USNM 26477 (F. C. Baker, 1945: 496).
DISTRIBUTION RECORDS.- BAJA
CALIFORNIA SUR: 30 mi. SW of Santiago, Mexico (F. C. Baker, 1945).
Biomphalaria
havanensis (Pfeiffer, 1839)
Planorbis
havanensis
Pfeiffer, 1839; Arch. Naturgesch., 5: 354.
Tropicorbis
havanensis (Pfeiffer). F. C. Baker, 1945; Moll. Fam. Planorbidae:
pl. 13, figs. 1-11 (reproductive anatomy); pl. 46, fig. 13 (kidney); pl. 68,
fig. 8 (radula); pl. 70, fig. 4 (animal); pl. 130, figs. 18-28 (shell).
Taphius
havanensis
(Pfeffer). Paraense & Deslandes,
1958; Rev. Brasil. Biol., 18: 87-91; fig. 1 (shell), fig. 2 (pallial organs),
fig. 3 (reproductive anatomy), fig. 4 (radula).
Biomphalaria
havanensis
(Pfeiffer). Bequaert & Miller, 1973:
205-208.- Thompson, 1984; Freshwater
snails of Florida: 72; figs. 155-157 (shell).-
Yung, Pointier & Parera, 1997; Malac. Rev., 30: 115-117, fig. 1
(map), fig. 2 (reproductive anatomy), fig. 3 (shell).
Planorbis
dicipiens C. B.
Adams, 1849; Contr. Conch., 3: 43.
Planorbis
maya Morelet, 1849;
Test Noviss., I: 16.- Pilsbry, 1891;
Proc. Acad. Nat. Sci. Phila., 43: 322.- Bequaert
& Clench, 1933; Pub. Carnegie Inst. Wash., (431): 538.
Planorbis
(Gyrorbis) maya Morelet. Fischer &
Crosse, 1880: 72-73; pl. 33, figs. 4-4c (shell).
Planorbis
(Gyraulus) maya Morelet. Von Martens, 1899: 392; pl. 21, fig. 13
(shell).
Planorbis
(Tropicorbis) maya (Morelet). Bequaert
& Clench, 1936; Pub. Canegie Inst. Wash., (457): 66.
Tropicorbis
(Tropicorbis) maya (Morelet). F. C.
Baker, 1945; Moll. Fam. Planorbidae: 85; pl. 130, figs. 13-17 (shell).
Taphius
maya (Morelet). Paraense & Deslandes, 1957a; Jour. de
Conchyl., 97: 49-57; figs. 1-3 (shell), fig. 4 (animal), fig. 5 (pallial
organs), fig. 6 (reproductive anatomy), fig. 7 (radula).
Planorbis
liebmanni Philippi,
1850; in Dunker, Martini & Chemnitz,
Syst. Conch. Cab., 17: 59; pl. 10, figs. 32-34 (shell).- Pilsbry, 1891; Proc. Acad. Nat. Sci. Phila,
43: 322.- Pilsbry, 1903; Proc. Acad.
Nat. Sci. Phila., 55: 777.- Pilsbry,
1928; Proc. Acad. Nat. Sci. Phila., 80: 117.
Tropicorbis
liebmanni (Philippi). Paraense & Deslandes, 1957b: Rev. Brasil.
Biol., 17: 427-434.- Paraense, 1958;
Rev. Brasil. Biol., 18: 68-70; fig. 6 (reproductive anatomy).
TYPE LOCALITIES.- Planorbis havanensis: Havana, Cuba
(Yung, et al, 1997). Planorbis decipiens: Jamaica. Planorbis
maya: in cisterns, Ciudad Campeche, Campeche. Planorbis
liebmanni: Veracruz.
DISTRIBUTION RECORDS.- West
Indies, Mexico and Central America; South American distribution remains to be
determined. EL SALVADOR, Dept. San Salvador: San Salvador (Paraense, 2000). GUATEMALA, Dept. Sacatepequez: Antigua (Von Martens, 1899); Lago de Dueñas
(Paraense, 2000). BELIZE, Belize Dist.: Boston Village (Paraense,
2000). CAMPECHE: Isla de Carmen;
Campeche; lake at km 49, rd. from Cd.
Campeche to Merida (Paraense & Delandes, 1957b). NUEVO LEON: Topo Chico, nr. Monterey
(Pilsbry, 1903). OAXACA: Isthmus of
Tehuantepec. TAUMAULIPAS: Mesa de Solis,
nr. La Lajilla, between Padilla and Jimenez (Pilsbry, 1928). VERACRUZ. YUCATAN: Merida (Pilsbry, 1891); Tabi (Von
Martens, 1899); cave at Actun Has; hacienda Yocat; Ticul District (Bequaert
& Clench, 1933); aguado 14 km N, 2 km E of Merida; Aguado Halal, nr.
Merida; pool 3 km S of Progreso (Bequaert & Clench, 1936).
TAXONOMY.- Paraense &
Deslandes (1957b) synonymize Planorbis
liebmanni (Philippi, 1850) with Planorbis
maya (Morelet, 1849), and in turn (1958) they synonymized Planorbis maya with Planorbis havanensis (Pfeiffer, 1839). Records of Biomphalaria havanensis from Florida, Missisippi, Louisiana and Texas
(Abdel-Malek, 1969) require further study concerning their specific status.
Biomphalaria helophila
(Orbigny, 1835)
Planorbis
helophilus Orbigny,
1835; Mag. de Zool., (62): 27.
Biomphalaria
helophila
(Orbigny).- Ruíz, 1987; Universidad y Ciencias, 4: 33-43; map 1; pl. 1 (shell);
pl. 2 (radula); pl. 3 (reproductive anatomy).-
Paraense, 2000; Malac. Rev.; 33: 4; figs. 2A (shell); 2B (reproductive
anatomy).
Australorbis
albicans (Pfeiffer,
1839). Paraense and Deslandes, 1962;
Nautilus, 75: 156-161; figs. 1 (shell, radula, reproductive anatomy).
Planorbis
dentiens Morelet,
1849; Test Noviss., I: 18.- Fischer
& Crosse, 1880; pl. 33, figs. 7, 7a, 7b, 7d (shell).
Planorbis
dentiens var. edentula Fischer & Crosse, 180:
81; pl. 34, figs. 6, 6a-c (shell).
Planorbis
declivus Tate,
1870: Amer. Jour. of Conch., 5: 159 (not Planorbis
declivus Braun, 1848).
Tropicorbis
tatei Baker, 1941;
Nautilus, 54: 96-97 (replacement name for Planorbis
declivus Tate, 1870).
Tropicorbis
shimeki F. C.
Baker, 1945; Moll. Fam. Planorbidae: 218-219; pl. 134, figs. 12-14, 28 (shell).
TYPE LOCALITIES.- Planorbis helophila: Lima, Peru. Planorbis
cannarum: Belize. Planorbis dentiens: Belize. Tropicorbis
shimeki: Ometope [Isla de Ometepe], Nicaragua; holotype USNM 534290.
DISTRIBUTION RECORDS.- EL
SALVADOR, Dept. Santa Ana: Lago de
Coatepeque (Paraense, 2000). GUATEMALA, Dept. Izabal: El Prodo (Paraense, 2000).
NICARAGUA,
Isla de Ometepe (F. C. Baker, 1945). Dept. Chontales: Acoyapa (Paraense,
2000). VERACRUZ: Coatzacoalcos (F. C.
Baker, 1945). Los Tuxtlas, Laguna Escondida and Laguna Zacatal (18º35’-18º36’
N, 94º04’-95º05’ W) (Ruíz, 1987).
Biomphalaria
hondurasensis (Clesson, 1878)
Planorbis
hondurasensis
Clesson, 1878; in Martini &
Chemnitz, Syst. Conch.-Cab., Ed. 2, Limnaeidae: 164; pl. 24, fig. 2
(shell).- Pilsbry, 1920; Proc. Acad.
Nat. Sci. Phila., 72: 6.
Planorbis
(Gyraulus) hondurasensis Clesson. Von
Martens, 1899: 393.
TYPE LOCALITY.- Santa Rosa,
Honduras.
DISTRIBUTION RECORDS.- HONDURAS:
“Santa Rosa”. COSTA RICA: Rio Torres (Von Martens, 1899). Prov.
San José: ditch along rd. from San
José to La Verbena.
Biomphalaria kuhniana
(Clessin, 1885)
Planorbis
kuhnerianus
Clessen, 1883; in Martini &
Chemnitz, Syst. Conch. Cab., ed. 2, Planorbis:
108; pl. 11, fig. 12 (shell).
Planorbis
kuhniana Clessen,
1885; in Martini & Chemnitz,
Syst. Conch. Cab., ed. 2, Planorbis:
413 (substitute name for Planorbis
kuhnerianus Clessen, 1883).
Tropicorbis
kuhnianus
(Clessen). F. C. Baker, 1945; Moll. Fam.
Planorbidae: pl. 132, figs. 6-9 (shell).
Biomphalaria
kuhniana
(Clessin). Paraense, 1988; Mem. do Inst.
Ozwaldo Cruz, 83: 1-12.- Paraense, 2000;
Malac. Rev., 33: 5-6; text-figs. 3A (shell), 3b (reproductive anatomy).
Planorbis
isthmicus Pilsbry,
1920e: 78-79; text-figs. (shell).
Tropicorbis (lateorbis) isthmicus
(Pilsbry). F. C. Baker, 1945; Moll. Fam.
Planorbidae: 85, 498; pl. 131, figs. 1-7 (shell).
TYPE LOCALITY.- Planorbis kuhniana: Paramaribo,
Surinam. Planorbis isthmicus: “Chinese” wells at Panama City, Panama.
DISTRIBUTION RECORDS.- PANAMA, Prov. Panama: Panama City; Lago de
Miraflores (F. C. Baker, 1945); Las Cumbres, nr. Panama City; National
University, Tucuman; Lago Isleta, environs of Panama City (Paraense, 2000).
Biomphalaria
nicaraguana (Morelet, 1851)
Planorbis
nicaraguana Morelet,
1851; Test. Noviss., II: 14.- Von
Martens, 1899: 391-392; pl. 21, fig. 10 (shell).
Biomphalaria
nicaraguana
(Morelet). Paraense, 2000; Malac. Rev.,
33: 7-9; text-fig. 4A, 4B (shell); 4C, 4D (reproductive anatomy).
TYPE LOCALITY.- Lago de
Nicaragua. “Type” BMNH 1993.2.4.594).
DISTRIBUTION RECORDS.- NICARAGUA,
Dept. Chontales: Acoyapa, nr. Lago de
Nicaragua (Paraense, 2000). Dept. Rio San Juan: Rio San Juan (Von
Martens, 1899).
Biomphalaria obstructa
(Morelet, 1849)
Planorbis
obsructus Morelet,
1849; Test Noviss., I: 16.
Planorbula
obstructa (Morelet). Fischer & Crosse, 1880: 78; pl. 33, figs.
3, 3a-3d (shell); 34, figs. 7, 7a-7d (shell).-
Pilsbry, 1891; Proc. Acad. Nat. Sci. Phila., 43: 322.- Hinkley, 1920; Nautilus, 34: 38.- Bequaert & Clench, 1936; Pub. Canegie
Inst. Wash., (457): 68,- Richards, 1937;
Trans. Amer. Philos. Soc., 77: 255.
Planorbis
(Planorbula) obstructus (Morelet). Von
Martens, 1899: 398-399.
Tropicorbis
(Obstructio) obstrutus (Morelet).
Goodrich & van der Schalie, 1937; Misc. Pub. Mus. Zool. Univ. Mich.,
(34): 33.- F. C. Baker, 1945; Moll. Fam.
Planorbidae: 85, 128, 262, 336, 372, 388, 492, 494, 502, 504; pl. 13, figs. 1-3
(reproductive anatomy); pl. 50, fig. 11 (jaw); pl. 68, fig. 7 (radula); pl. 76,
fig. 7 (shell); pl. 128, figs. 26-33 (shell); pl. 129, figs. 4, 8, 9 (shell);
pl. 133, figs. 28-35 (shell); pl. 134, figs. 10, 11, 22 (shell).- Harry, 1950; Occ. Pap. Mus. Zool. Univ.
Mich., (524): 7, 24.- Bequaert, 1957;
Bull. Mus. Comp. Zool., 116: 214.
Biomphalaria
obstructa
(Morelet). Abdel-Malek, 1969;
Malacologia, 7: 186-190; 194-196; figs. 2A, 2B (reproductive anatomy), figs.
9-11 (shell), fig. 21A (radula).-
Paraense, 1990; Mem. do Inst. Ozwaldo Cruz, 85: 391-399.- Paraense, 2000; Malac. Rev., 33: 9-12;
text-figs. 5A, 5B, 5C (shell), 5D (reproductive anatomy); figs. 6A, 6B (shell);
6C (reproductive anatomy).
Planorbis
cannarum Morelet,
1849; Test Noviss., I: 16.
Segmentina
donbilli Tristram,
1861; Proc. Zool. Soc. London: 232.
Tropicorbis
obstructs donbilli
(Tristram). F. C. Baker, 1945; Moll.
Fam. Planorbidae: 85; pl. 129, figs. 17-25; pl. 131, figs. 15-26 (shell).
Planorbis
berendti Tryon,
1866; Amer. Jour. Conch., 2: 10; pl. 2, figs. 14-16 (shell).- Strebel, 1873: 45; pl. 5, fig. 23 (shell).
Planorbula
obstructa anodonta
Pilsbry, 1920; Proc. Acad. Nat. Sci. Phila., 71: 219 (not illustrated).
Tropicorbis
obstructus anodontus
(Pilsbry). F. C. Baker, 1945: Moll. Fam.
Planorbidae: 85, 502; pl. 133, figs. 36-40 (shell).
TYPE LOCALITIES.- Planorbis obstructus: Isla de Carmen,
Campeche. Planorbis cannarum: Belize. Segmentina donbilli: Lago de Dueñas, Guatemala. Planorbis
berendti: Orizaba, Veracruz. Planorbula obstructa anodonta: reservoir 4 miles north of Guatemala City,
Guatemala.
DISTRIBUTION RECORDS.- Gulf
coastal regions of the United States south to Belize, Guatemala, and El
Salvador. EL SALVADOR, Dept. San Salvador: San Salvador
(Paraense, 2000). GUATEMALA, Dept. Escuintla: Escuintla (Von Martens,
1899). Dept. Guatemala: Lago Amatitlan; reservoir N of Cd. Guatemala (F.
C. Baker, 1945). Dept. Izabal: Lago de Izabal, Marisco. Dept.
Sacatepequez: Lago de Dueñas, San Miguel Dueñas (Paraense, 2000). Dept.
Peten: Lago de Petén; aguado NW of
San Benito (Goodrich & van der Schalie, 1937). BELIZE, Belize
Dist.: Boston Village (Paraense, 2000).
CAMPECHE: Isla de Carmen; Aguado Yalic, Chapoton (Beqaert & Clench,
1936). CHIAPAS: Laguna Ocotal to El
Censo (Bequaert, 1957). OAXACA: San
Carlos, east of Oaxaca; Isthmus of Tehuantepec (Fischer & Crosse,
1880). QUINTANA ROO: swamp a few miles N
of San Miguel, Cozumel Island (Richards, 1937); nr. San Miguel, Cozumel Island
(F. C. Baker, 1945). SAN LUIS POTOSI:
Catamas Lake, El Abra; falls of the Rio Valles, Cd. Valles; Tamosopo (F. C.
Baker, 1945). TAMAULIPAs: Tampico (F. C.
Baker, 1945). VERACRUZ: Quilate, nr.
Misantla; Orizaba; Veracruz; Rio Tenoya, Veracruz (Strebel, 1873). YUCATAN: Ticul (Pilsbry, 1891); aguado 14 km
N, 2 km E of Merida; Aguado Yalal, nr. Merida; Piste (Bequaert & Clench,
1936); Piste (F. C. Baker, 1945); 0.5 mi. W of Progreso; irrigation cistern, 3
mi. N of merida (Harry, 1950).
Biomphalaria orbicula
(Morelet, 1849)
Planorbis
orbiculus Morelet,
1849; Test Noviss., I: 17.- Fischer
& Crosse, 1880: 70; pl. 32, figs. 9, 9a-c (shell).- Pilsbry, 1891; Proc. Acad. Nat. Sci. Phila.,
43: 322.- Hinkley, 1920; Nautilus, 34:
38.- Richards, 1937; Proc. Amer. Philos.
Soc., 77: 255.
Planorbis
(Menetus) orbiculus Morelet. Von Martens, 1899: 390-391.
Planorbis
(Tropicorbis) orbiculus (Morelet). Bequaert
& Clench, 1936; Pub. Canegie Inst. Wash., (457): 66-67.
Tropicorbis
(Tropicorbis) orbiculus (Morelet). F. C.
Baker, 1945; Moll. Fam. Planorbidae: 85, 390, 494; pl. 77, figs. 7-9 (shell);
pl. 129, figs. 1-3, 4-7, 10-16 (shell).
Tropicorbis
orbiculus dunkeri
F. C. Baker, 1945; Moll. Fam. Planorbidae: 85; pl. 129, figs. 26-36
(shell).- Harry, 1961; Malacologia, 1:
45 (a nomen nudum, holotype not
designated).
TYPE LOCALITY.- Planorbis oriculus: La Palizada,
Yucatan, on the frontier of Tabasco. Tropicorbis
orbiculus dunkeri: not stated.
DISTRIBUTION RECORDS.- Guatemala
and eastern
Biomphalaria
panamensis Dunker, 1848
Planorbis
panamensis Dunker,
1848; Proc. Zool. Soc. London: 41.-
Sowerby, in Reeve; Conch.
Icon., 22, Planorbis: pl. 12. fig.
95.
Planorbis (Gyraulus) panamensis
Dunker, 1848. Von Martens, 1899: 395.
TYPE LOCALITY.- West Panama, in a
small stream.
DISTRIBUTION
Biomphalaria
petenensis (Morelet, 1851)
Planorbis
petenensis Morelet,
1851; Test. Noviss., II: 15.
Planorbis
(Gyrorbis) petenensis Morelet. Fischer
& Crosse, 1880; 74; pl. 33, figs. 5, 5a-c (shell).
Planorbis
(Gyraulus) petenensis Morelet. Von
Martens, 1899: 393-394.
Tropicorbis (Lateorbis) petenensis
(Morelet). F. C. Baker, 1945; Moll. Fam.
Planorbidae: 85, 502, 518; pl. 133, figs. 1-3 (shell); pl. 141, figs. 2-4
(shell).
TYPE LOCALITY.- Lago de Itza
Petén, Dept. Petén, Guatemala.
DISTRIBUTION RECORDS.- COSTA
RICA, Prov. San José: Rio Torres, nr.
San José. Prov. Puntarens: Laguna between Lagarto and Boruca (Von Martens,
1899). GUATEMALA, Dept. Petén: Lago Petén Itza; Lago de Peten, Remate (F. C. Baker,
1945). OAXACA: Cacoprieto, isthmus of
Tehuantepec (Von Martens, 1899); Tehuantepec (F. C. Baker, 1945).
Biomphalaria retusus
(Morelet, 1849)
Planorbis
retusus Morelet,
1849; Test Noviss., I: 17.- Pilsbry,
1891; Proc. Acad. Nat. Sci. Phila., 43: 322.
Planorbis (Gyrorbis) retusus
Morelet. Fischer & Crosse, 1880: 73;
pl. 32, figs. 10-10c (shell).
Planorbis
(Gyraulus) retusus Morelet. Von
Martens, 1899: 393.
TYPE LOCALITY.- Isla de Carmen,
Campeche.
DISTRIBUTION RECORDS.- CAMPECHE:
Isla de Carmen. YUCATAN: Shkolac,
between Merida and Valladolid (Pilsbry, 1891).
Biomphalaria straminea
(Dunker, 1848)
Planorbis
stramineus Dunker,
1848; Proc. Zool. Soc. London: 42.-
Dunker, 1850; in Martini &
Chemnitz, Syst. Conch. Cab., Ed. 2, 1, Planorbis:
42; pl. 5, figs. 7-9 (shell).
Biomphalaria
straminea
(Dunker). Paraense, 2000; Malac. Rev.,
33: 12.
TYPE LOCALITY.- South America.
DISTRIBUTION RECORDS.- COSTA
RICA, Prov. Cartago: Rio Coris
(Paraense, 2000).
Biomphalaria subprona
(Von Martens, 1899)
Planorbis
(Taphius) subpronus Von Martens, 1899: 396-397; pl. 21, fig. 15 (shell).
Taphius
sbpronus (Von
Martens). F. C. Baker, 1945; Moll. Fam.
Planorbidae: 79, 498; pl. 131, figs. 36-40 (shell).- Bequaert, 1957; Bull. Mus. Comp. Zool., 116:
214-215.
Biomphalaria
subprona (Von
Martens). Paraense, 1996; Mem. do Inst.
Ozwaldo Cruz, 91: 187-190.- Paraense,
2000; Malac. Rev., 33: 12.
TYPE LOCALITY.- Amatitlan,
Tabasco [Guatemala].
DISTRIBUTION RECORDS.- COSTA
RICA: Turrialba (F. C. Baker, 1945). EL
SALVADOR: Lago de Coatepeque (Bequaert, 1957).
GUATEMALA, Dept. Guatemala: Lago
de Amatitlan (Paraense, 2000). CHIAPAS:
Laguna Ocotal, 950 m alt. (Bequaert, 1957).
Biomphalaria
“temascalensis” Ruiz, 1987
Biomphalaria
temascalensis Ruiz,
1987; Universidad y Ciencias, 4: 25-34;
pl. 1, figs. a-c (shell); pl. 2, figs. 1-4 (radula); pls. 3-4 (reproductive
system).
TYPE LOCALITY.- Estacion de
Acuacultura Tropical de Temascal, Oaxaca, Mexico (17-19º N, 95-97º40’ W). Holotype not designated.
DISTRIBUTION RECORDS.- Known only
from the type locality.
TAXONOMY.- The name Biomphalaria temascalensis Ruíz, 1987 is
invalid, because the description does not include the designation of a holotype
as required by ICZN Articles 16.4 and 62.4.
Biomphalaria
tepicensis (Von Martens, 1899)
Planorbis (Gyraulus) tepicensis Von
Martens, 1899: 393; pl. 21, figs. 14 (shell).
Tropicobis
tepicensis (Von
Martens). F. C. Baker, 1945; Moll. fam.
Planorbidae: 85, 502; pl. 133, figs. 4-9 (shell).
TYPE LOCALITY.- Tepic, “Jalisco”
[Nayarit].
DISTRIBUTION RECORDS.- NAYARIT:
Tepic. OAXACA: Tehuantepec (F. C. Baker,
1945).
Subfamily HELISOMATINAE
F. C. Baker, 1928
Genus Helisoma Swainson, 1840
Helisoma Swainson, 1840; Treatise on
Malacology, VIII: 337.- F. C. Baker,
1945; Moll. Fam. Planorbidae: 123-129.
TYPE SEPCIES.- Planorbis bicarinatus Say, 1817 (= Planorbis anceps Menke, 1830).
DISTRIBUTION.- Throughout North America south to northern Mexico.
TAXONOMY.- Two subgenera are
recognized (Burch, 1980: 198).
Subgenus Helisoma Swainson, 1840
DISTRIBUTION.- As for the genus.
TAXONOMY.- A single species is
recognized. Numerous subspecies and
varieties have been proposed (Burch, 1880: 198, 274). Two subspecies are recognized currently. One occurs in Mexico.
Helisoma anceps anceps
(Menke, 1830)
Planorbis
bicarinatus Say,
1817; Nichole’s Encycl., 1st ed.: species No. 2, pl. 1, fig. 4 (not Planorbis bicarinatus Lamarck, 1804).
Planorbis
anceps Menke, 1830;
Synopsis Methodica Molluscorum. 2nd Ed. 242
Planorbis
antrosa Conrad,
1834; Amer. Jour. Sci., 25: 343.
Helisoma
anceps anceps
(Menke, 1830). F. C. Baker, 1945; Moll.
Fam. Planorbidae: 402-403.- Taylor,
1966; Veliger, 9: 209.- Burch, 1980;
Walkerana, 1: 198, 274-275; fig. 714.
TYPE LOCALITY.- Delaware River,
Virginia.
DISTRIBUTION RECORDS.- Rio de Los
Natadores, at El Cariño, 20 mi. E. of Cuatro Cienegas (Taylor, 1966). SONORA: mouth of the Rio Yaqui (F. C. Baker,
1945),
Genus Planorbella Haldeman, 1842
Planorbella Haldeman, 1842; Monogr. NA
freshwater Moll., Physidae: 14.- Burch,
1980; Walkerana, 1: 202.
TYPE SPECIES.- Planorbis campanulatus Say, 1821.
DISTRIBUTION.- North America and
the west Indies south to Peru
TAXONOMY.- Three subgenera are
recognized. Two occur in the study
area.
Subgenus Pierosoma Dall, 1905
Pierosoma
Dall, 1905; Smithsonian
Inst. Harriman Alaska Expedition, 1899, Alaska Ser., 13: 81, 85.
TYPE SPECIES.- Planorbis trivolvis Say, 1818.
DISTRIBUTION.- North America
south to Panama and the West Indies.
TAXONOMY.- Numerous species.
Seven occur in the study area.
Planorbella
(Pierosoma) caloderma (Pilsbry, 1923)
Planorbis
caloderma
Pilsbry. Hinkley, 1920; Nautilus, 34:
46, 48 (nomen nudum).
Planorbis
caloderma Pilsbry,
1923; Nautilus, 36: 143-144.-
Helisoma
calodermum
(Pilsbry). F. C. Baker, 1945; Moll. Fam.
Planorbidae: pl. 115, figs. 2-4 (shell).
TYPE LOCALITY.- Esmeralda, Dept.
Jutiapa, Guatemala. Holotype and
paratypes ANSP 45662.
DISTRIBUTION RECORDS. GUATEMALA, Dept. Izabal: Jocolo (Hinkley,
1920). Dept. Jutiapa: Esmeralda.
Planorbella
(Pierosoma) foveale foveale (Menke, 1830)
Planorbis
fovealis Menke,
1930; Symp. Method. Mollusc.: 37.
Helisoma
foveale
(Menke). F. C. Baker, 1945; Moll. Fam.
Planorbidae: 149, pl. 101, figs. 1-5
(shell); pl. 141, figs. 12-18 (shell).-
Harry & Hubendick, 1964; Göeborgs Kungl. Vetenskaps-och
Vitterhets-samhölles handlingar, ser. 6, ser. B: 35-40; fig. 33-37 (anatomy);
figs. 73-77, 131-133 (shell).
Planorbis
caribaeus Orbigny,
1841, in Sagra, Hist. Fis. Polit. y nat. de Cuba, Moluscos: 193; pl. 13, figs.
17-19 (shell).- Pilsbry, 1891; Proc.
Acad. Nat. Sci. Phila., 43: 322.-
Pilsbry, 1920; Proc. Acad. Nat. Sci. Phila., 72: 6.- Hinkley, 1920; Nautilus, 34: 38, 48.
Planorbis
(Helisoma) caribaeus (Orbigny). Bequaert
& Clench, 1936; Pub. Canegie Inst. Wash., (457): 66.
Helisoma
caribaeum
(Orbigny). van der Schalie, 1940; Occ.
Pap. Mus. Zool. Univ. Mich., (413): 6-7.
Helisoma
(Pieronoma) caribaeum caribaeum (Orbigny).
F. C. Baker, 1945; Moll. Fam. Planorbidae: 149, 466, 434, 436, 438; pl.
99, fig. 31; pl. 100, figs. 1-17; pl. 101, fig. 15; pl. 115, fig. 1
(shell).- Bequaert, 1957; Bull. Mus.
Comp. Zool., 116: 214.
Planorbis
tumidus Dunker,
1850; in Martini & Chemnitz,
Syst. Conch. Cab., Ed. 2, Planorbis:
39; pl. 7, figs. 10-12 (shell).- Dall,
1896; Proc. U. S. Nat. Mus., 19: 369.-
Pilsbry, 1891; Proc. Acad. Nat. Sci. Phila., 43: 322.
Planorbis
ancylostomus Crosse
& Fisher, 1879; Jour. de Conchyl., 27: 341.- Fischer & Crosse, 1880: 63; pl. 32, figs.
5, 5a, 5b (shell).
Planorbis
ancylostoma var. chiapnesis Fischer & Crosse, 1886:
63; pl. 34, figs. 5, 5a, 5b (shell).
Planorbis
(Helisoma) caribaeus chiapensis Fischer & Crosse. Von
Martens, 1893; Biol. Cent. Amer.: 389.
TYPE LOCALITIES.- Planobis fovealis: Jamaica (Pilsbry,
1934). Planorbis caribeaus:
Cuba. Planorbis dunkeri: not specified.
Planorbis ancylostomus:
Veracruz, Veracruz. Planorbis ancylostoma
var. chiapensis: Chiapas.
DISTRIBUTION RECORDS.- COSTA
RICA, Prov. Cartago; E of Cartago; 4
mi. SW of Cartago; San Isidro del Tejar (Pilsbry, 1920). GUATEMALA, Naranjo (F. C. Baker, 1945). Dept.
Alta Verapaz: a small aguada ca. 8 km N of Samanzana; nr. Samac (van der
Schalie, 1940). Dept. Izabal: Jocolo (Hinkley, 1920). Dept.
Guatemala: Lago Amatitlan. CAMPECHE:
Aguado Yalic, nr. Chompoton (Bequaert & Clench, 1936). CHIAPAS: Laguna Ocotal, 950 m alt.; Rio
Amarillo at the Sumidero, nr. Las Casas (Bequaert, 1957). SONORA: Rio San Bernardino; Rio Sonoyta (Dall,
1896). VERACRUZ: Orizaba (Pilsbry,
1891); Veracruz (F. C. Baker, 1945).
YUCATAN: Shkolak (Pilsbry, 1891).
TAXONOMY.- The following
subspecies has been proposed from the study area.
Planorbella (Pierosoma) foviale guatemalense (Clessin, 1889)
Planorbis
guatemalenis Clessin,
1889; in Martini & Chemnitz,
Syst. Conch. Cab., Ed. 2, Planorbis:
209-210; pl. 32, fig. 7 (shell).
Helisoma (Pierosoma) caribaeum
guatemalense (Clessin). F. C. Baker,
1945; Moll. Fam. Planorbidae: 149, 434; pl. 99, figs. 32-36 (shell).
TYPE LOCALITY.- Guatemala.
DISTRIBUTION RECORDS.- NICARAGUA
(F. C. Baker, 1945). GUATEMALA.
Planorbella
(Pierosoma) costaricense (Preston, 1907)
Planorbis
costaricensis
Preston, 1907; Ann. Mag. Nat. Hist., Ser. 7, 20: 496-497; text-fig. 16 (shell).
Helisoma (Pierosoma) costaricense
(Preston). F. C. Baker, 1945; Moll. Fam.
Planorbidae: 149, 438; pl. 101, figs. 8-10 (shell).
TYPE LOCALITY.- Catalina,
Guanacaste Prov., Costa Rica.
DISTRIBUTION RECORDS.- COSTA
RICA, Prov. Guanacaste: Catalina (F.
C. Baker, 1945).
Planorbella contrerasi
(Pilsbry, 1920)
Planorbis
contrerasi Pilsbry,
1920; Proc. Acad. Nat. Sci. Phila., 72: 193; text-fig. 2 (shell).
Helisoma (Pierosoma) contrerasi
(Pilsbry). F. C. Baker, 1945; Moll.
Fam. Planorbidae: 140; pl. 102, figs. 2-4 (shell).
TYPE LOCALITY.- Lago de Chapala,
Jalisco. Holotype ANSP 46193.
DISTRIBUTION RECORDS.- JALISCO:
known only from the type locality.
Planorbella
(Pierosoma) tenue (Dunker, 1850)
Planorbis
tenuis Dunker,
1850; Syst. Conch. Cab., Ed. 2, Limnaecea: 45; pl. 9, figs. 14-19 (shell); pl.
16, figs. 22-25 (shell).- Strebel, 1872:
42; pl. 5, figs. 21-26, a-c (shell).-
Fischer & Crosse, 1880: 60; pl. 34, fig. 1 (shell).- Von Martens, 1899: 381-386; pl. 21, figs. 1,
7, 7a (shell).- Pilsbry, 1904; Proc.
Acad. Nat. Sci. Phila., 55: 777.-
Pilsbry, 1920; Proc. Acad. Nat. Sci. Phila., 72: 6.
Helisoma
tenuis (Dunker). Goodrich & van der Schalie, 1937; Misc.
Pub. Mus. Zool. Univ. Mich., (34): 33.
Helisoma (Pierosoma) tenue tenue
(Dunker). F. C. Baker, 1945; Moll. Fam.
Planorbidae: 149, 430; pl. 97, figs. 1-11 (shell).- Bequaert & Miller, 1987: 208.
Planorbella
(Pierosoma) tenue (Dunker). J. B. Burch,
1980; Walkerana: 204; fig. 734 (shell).
Planorbis
mexicanus Dunker,
1850; in Martini & Chemnitz,
Syst. Conch.-Cab., Ed. 2, Planorbis:
45.
Planorbis
tumens Carpenter,
1857; Cat. Mazatlan shells: 81.-
Fischer & Crosse, 1886: 62;
pl. 33, figs. 3, 3a (shell).- Clessin,
1886; in Martini & Chemnitz,
Syst. Conch. Cab., ed. 2; Limnaeiden: 88; pl. 12, figs. 7-9 (shell).- Hanna, 1923; Proc. Calif. Acad. Sci., ser. 4,
12: 516-517.
Planorbis
(Helisoma) tumens Carpenter. Von
Martens, 1899: 387.
Helisoma
tumens
(Carpenter). F. C. Baker, 1945; Moll.
Fam. Planorbidae: 436; pl. 100, figs. 31-37 (shell).
Planorbis
tenuis var. juvenilis Von Martens, 1899: 384; pl.
21, fig. 4 (shell).
Planorbis
tenuis var. uhdei Von Martens, 1899: 385; pl. 21,
fig. 2 (shell).
TYPE LOCALITIES.- Planorbis tenuis: ditches in the
neighborhood of Ciudad Mexico. Planorbis mexicanus: Mexico. Planorbis
tumens: Mazatlan, Sinaloa. Planorbis tenuis var. juvenilis:
City of Mexico. Planorbis tenuis var. uhdei:
central mexico.
DISTRIBUTION.- Guatemala, Dept. Peten: Arroyo Subín, nr. Sanat
Teresa (Goodrich & van der Schalie, 1937).
BAJA CALIFORNIA SUR: Mulege (Hanna, 1923). DURANGO.
DISTRITO FEDERAL: Ciudad Mexico; Lago de Chalco (Pilsbry, 1920).
GUANAJUATO: Jaral (Von Martens, 1899); Acambaro (Pilsbry, 1903). JALISCO; Lago de Chapala (Von Martens,
1899). MEXICO: Tlapam (Pilsbry, 1903);
Toluca; sources of the Rio Lerma, nr. Toluca (F. C. Baker, 1945). SINALOA: Mazatlan (Von Martens, 1899). VERACRUZ: Cordoba; Orizaba; Jalapa (Von
Martens, 1899).
TAXONOMY.- F. C. Baker (1945)
recognizes the following subspecies from the study area.
Planorbella (Pierosoma) tenue boucardi (Fischer & Crosse, 1880)
Planorbis
tenuis var. boucardi Fischer & Crosse, 1880: 61;
pl. 32, figs. 3a, b (shell).- Von
Martens, 1899: 384.
Helisoma
tenue boucardi (Fischer
& Crosse). F. C. Baker, 1945; Moll.
Fam. Planorbidae: 149, 430; pl. 97, figs. 12-17 (shells).
TYPE LOCALITY.- Environs of
Ciudad Mexico.
DISTRIBUTION RECORDS.- DISTRITO
FEDERAL: known only from the environs of Ciudad Mexico.
Planorbella (Pierosoma) tenue chapalensis (Pilsbry, 1920)
Planorbis
chapalensis Pilsbry, 1920; Proc. Acad. Nat. Sci. Phila.,
72: 192-193; fig. 1 (shell).
Helisoma
tenue chapalensis (Pilsbry). F. C. Baker, 1945; Moll. Fam. Planorbidae:
149; 440; figs. 1, 5-10 (shell).
TYPE LOCALITY.- Lago de
Chapala, Jalisco. Holotype ANSP 46194.
DISTRIBUTION RECORDS.- JALISCO:
known only from the type locality.
Planorbella (Pierosoma) tenue exaggeratum (Von Martens, 1899)
Planorbis
tenuis Philippi. Pilsbry, 1891; Proc. Acad. Nat. Sci. Phila.,
43: 321-322; pl. 15, fig. 4 (shell).
Planorbis
tenuis var. exaggeratus Von Martens, 1899:
385.- Pilsbry, 1903; Proc. Acad. Nat.
Sci. Phila., 55: 777.
Helisoma
tenue exaggeratum (Von
Martens). F. C. Baker, 1945; Moll. Fam.
Planorbidae: 149, 430; pl. 97; figs. 18-25 (shell).
TYPE LOCALITY.- Lago de
Patzcuaro, Michoacán.
DISTRIBUTION RECORDS.- MICHOACÁN: known only from the type locality.
Planorbella (Pierosoma) tenue pertenue (F. C. Baker, 1940)
Planorbis
tenuis var. applanatus Von Martens, 1899: 384-385;
pl. 21, fig. 3 (shell) (not Planorbis
applanatus Thomae, 1845).
Helisoma
tenuis pertenuis F.
C. Baker, 1940; Nautilus, 54: 97 (replacement name for Planorbis tenuis applanatus Von Martens).
Helisoma
tenue applanatum (Von
Martens). F. C. Baker, 1945; Moll. Fam.
Planorbidae: 149, 430, 432; 434; pl. 97, figs. 26-28; pl. 98, figs. 1-8; pl.
99, figs. 1-3; pl. 101, fig. 16 (shell).
TYPE LOCALITY.- Plateau of
Mexico.
DISTRIBUTION RECORDS.- CHIHUAHUA:
Lanos River (F. C. Baker, 1945). SONORA:
Rio Yaqui, nr. Ciudad Obregon (F. C. Baker, 1945). VERACRUZ: Orizaba (F. C. Baker, 1945). BAJA CALIFORNIA NORTE: Vallecitos (F. C.
Baker, 1945).
Planorbella (Pierosoma) tenue strebelianum (Fischer & Crosse, 1880)
Planorbis
tenuis var. strebialunus Fischer
& Crosse, 1880: 63.
Helisoma
tenue strebelianum (Fischer
& Crosse). F. C. Baker, 1945; Moll.
Fam. Planorbidae: 149, 434; pl. 99, figs. 16-20, 25-30 (shell).
TYPE LOCALITY.- Laguna de Los
Cocos, Rio Tenoya and ditches in environs of Veracruz.
DISTRIBUTION RECORD.- SAN LUIS
POTOSI: Penasco, 5 mi. N of San Luis Potosi; Rio Coy (F. C. Baker, 1945). Tamaulipas: Alta Mira (F. C. Baker,
1945). VERACRUZ: environs of Veracruz.
Planorbella
(Pierosoma) trivolvis (Say, 1817)
Planorbis
trivolvis Say,
1817; Nicholson’s Encyclopedia, American Ed. 3: pl. 2, fig. 2 (shell).
Planorbella
(Pierosoma) trivolvis trivolvis (Say). J. B. Burch, 1980; Walkerana, 1: 205;
figs. 735 (shell).
Helisoma
trivolvis
(Say). Paraense, 2000; Malac. Rev., 33:
2; text-figs. 1A (shell), 1B (kidney), 1C (reproductive anatomy).
TYPE LOCALITY.- Lake Erie,
North America.
DISTRIBUTION RECORDS.- Widely
distributed in North America from the Atlantic coast westward to Missouri and
northward to the Arctic Circle and Alaska.
Records from Mexico and Central America seem questionable. COSTA RICA, Prov. Cartago: Rio Coris (Paraense, 2000). NICARAGUA, Dept. Grenada: Lake Nicaragua,
Grenada (Paraense, 2000). GUATEMALA, Dept. Alta Verapaz: Lago Petencito, nr.
Cobán (Paraense, 200o). BELIZE, Belize Dist.: Boston Village (Paraense,
2000).
Planorbella
(Pierosoma) wyldi (Tristram, 1861)
Planorbis
tumens
Carpenter. Fischer & Crosse, 1880:
62; pl. 32, figs. 4, 4a-c (shell)
Planorbis
wyldi Tristram,
1861; Proc. Zool. Soc. London: 232.
Planorbis
tenuis var. wyldi Tristram. Fischer & Crosse, 1880; 62; pl. 34, fig.
2 (shell).
Planorbis
(Helisoma) wyldi Tristram. Von Martens,
1899: 386-387; pl. 21, fig. 5 (shell).
Helisoma (Pierosoma) wyldi
(Tristram). F. C. Baker, 1945; Moll.
Fam. Planorbidae: 149, 436; pl. 100, figs. 20-30.
Planorbis
salvini Clessin,
1884; in Martini & Chemnitz,
Syst. Conch. Cab., Ed. 2, Planorbis:
207-208; pl. 31, fig. 8 (shell).
Helisoma
wyldi salvini
(Clessin). F. C. Baker, 1945; Moll.
Fam. Planorbidae: 149, 436; pl. 100, figs. 18-20 (shell).
TYPE LOCALITY.- Planorbis wyldi: Lago de Dueñas, Dept.
Sacatepequez, Guatemala. Planorbis salvini: Guatemala.
DISTRIBUTION RECORDS.- Known in
Central America from Costa Rica north to Guatemala. COSTA RICA, Prov. Limón: Rio Reventazón at Ujarras;
Alta Coca valley nr. Talamanca. Prov. Puntarenas: along Rio Diquis,
below Terraba (Von Martens, 1899).
NICARAGUA, Dept. Grenada: nr.
Volcan Monbacho (Von Martens, 1899); San Carlos (F. C. Baker, 1945). EL SALVADOR, Dept. San Miguel: Lago de Olomega (F. C. Baker, 1945). Dept.
Santa Ana: Lago de Guija; Lajo de Metapan (F. C. Baker, 1945). GUATEMALA, Dept. Alta Verapaz:
Coban. Dept. Guatemala: Aceituno; Naranjo.
Dept. Sacatepequez: Lago de
Dueñas; Santiago Zamora (Von Martens, 1899).
Subgenus Seminolina Pilsbry, 1934
Seminolina Pilsbry, 1934; Proc. Acad. Nat.
Sci. Phila., 86: 31.- F. C. Baker, 1945;
Moll. Fam. Planorbidae: 129-134.
TYPE SPECIES.- Planorbis scalare Jay, 1839.
DISTRIBUTION.- The natural
distribution is confined to peninsular Florida.
It has been introduced widely in other parts of the world.
TAXONOMY.- Two species are
recognized.
Planorbella duryi
(Wetherby, 1879)
Planorbis
duryi Wetherby,
1879; Jour. Cincinnati Soc. Nat. Hist., 2: 99; fig. 4 (shell).
Helisoma
(Seminolina) duryi duryi (Wetherby).
Pilsbry, 1934; Proc. Acad. Nat. Sci. Phila., 86: 38-40; pl. 10, figs.
1-3 (shell).- F. C. Baker, 134, 442,
444; pl. 103, figs. 20-30 (shell); pl. 104, figs. 1-7 (shell).
Planorbella
duryi
(Wetherby). Burch, 1980; Walkerana, 1:
204; figs. 718, 719 (shell).- Thompson,
1984; Freshwater snail of Fla. Man. for Identify.: 76-77; figs. 189-193.- Paraernse, 2000; Malac. Rev., 33: 2, 4.
TYPE LOCALITY.- The Florida
Everglades, Florida.
DISTRIBUTION RECORDS.- Confined
to south Florida in its natural distribution.
Introduced widely elsewhere.
COSTA RICA, Prov. Cartago: Rio
Coris (Paraense, 2000).
TAXONOMY.- Pilsbry (1934) and F.
C. Baker (1945) recognize six subspecies.
Their status has not been investigated since then.
Genus Micromenetus F. C. Baker, 1945
Micromenetus F. C. Baker, 1945; Moll. Fam.
Planorbidae: 197-190.
TYPE SPECIES.- Planorbis dilatatus Gould, 1841.
DISTRIBUTION.- Eastern North
America; Bermuda.
TAXONOMY.- Eight species are
recognized. One occurs in the study
area.
Micromenetus dilatatus
avus (Pilsbry, 1905)
Planorbis
alabamensis var. avus Pilsbry, 1905; Nautilus, 19: 34.
Promenetus (Micromenetus) alabamensis
avus Pilsbry. F. C. Baker, 1945;
Moll. Fam. Planorbidae: 190; pl. 121, figs. 40, 41; pl. 123, fig. 1; pl. 140,
fig. 25 (shell).
Micromenetus
dilatatus avus (Pilsbry). Thompson, 1983; Nautilus, 97: 68-69.- Thompson, 1984; Freshwater snail of Fla, Man.
for identification: 68; figs. 143-145 (shell).
Promenetus
minutus Taylor,
1954; Rev. Soc. Malac. Carlos de la Torre, 6: 37-38.
TYPE
LOCALITIES.- Pliocene Caloosahatchee
Formation, Florida.
DISTRIBUTION RECORDS.- Known from
South Florida, Jamaica, Haiti, and Panama.
PANAMA, Canal Zone: Allee
Stream, Barro Colorado Island (Taylor, 1954).
Family
PHYSIDAE Fischer and Crosse, 1886
DISTRIBUTION.— Europe, temperate
Asia, North America, Meso-America, South America.
TAXONOMY.- The family contains
two subfamilies and 23 genera. Taylor
(2003) states that in many cases species cannot be identified accurately by
shell characters. The locality records
listed below are taken from Taylor.
Previous published records are omitted because of the chaotic taxonomic
history of the family in Mexico and Central America.
Subfamily
APLEXINAE Starobogatov, 1967
TAXONOMY.- Taylor (2003) lists
twelve genera.
DISTRIBUTION.— Holarctic in
distribution and extending into the tropical realm in South America.
Genus Austrinauta Taylor, 2003.
Austrinauta Taylor, 2003; Rev. Biol. Trop., 51
(suppl.): 43.
TYPE SPECIES.- Physa elata Gould,
1853.
DISTRIBUTION.— Known only from
Nayarit State, México.
TAXONOMY.- One species.
Austrinauta elatus (Gould,
1853)
Physa
elata Gould, 1853;
Jour. Boston Soc. Nat. Hist., 6: 379; pl. 14, fig. 4.
Aplecta
elata (Gould). Fischer and Crosse, 1870-1902; 2: 92; pl. 32,
fig. 2.
Austrinauta
elatus (Gould). Taylor, 2003; Rev. Biol. Trop., 51 (suppl.):
45-47; figs. 15 (map), 16-19 (anatomy).
TYPE LOCALITY.— “Lower
California”. Lectotype MCZ 169130
(Johnson, 1964: 71).
DISTRIBUTION RECORDS.— Known only
from a single locality. NAYARIT:
freshwater marsh beside Matachén-Santa Cruz road, at south end of Playa
Matachén (21˚29.6’ N, 105˚11.7’ W).
Genus Amecanauta Taylor, 2003
Amecanauta Taylor, 2003; Rev. Bio. Trop., 51
(suppl.): 72.
TYPE SPECIES.- Amecanauta jaliscoensis Taylor, 2003.
DISTRIBUTION.- Nayarit and
Jalisco, México.
TAXONOMY.- A single species.
Amecanauta
jaliscoensis Taylor, 2003
Amecanauta
jaliscoensis
Taylor, 2003; Rev. Biol. Trop., 51 (suppl.): 73-74; text-fig. 15 (map),
text-figs. 49-52 (anatomy); pl. 1, fig. 3 (shell).
TYPE LOCALITY.— Jalisco: roadside ditch on W side of hwy. 200, opposite entrance to “Modulo de
Abasto” de Puerto Vallarta, 2.2 km NE of entrance to airport (20˚ 41.48’
N, 105˚13.95’ W). Holotype CAS
114813.
DISTRIBUTION RECORDS.- JALISCO:
known only from the type locality.
Genus Mexinauta Taylor, 2003
Mexinauta Taylor, 2003; Rev. Biol. Trop., 51
(suppl.): 74-76.
TYPE SPECIES.- Physa
nitens Philippi, 1841.
DISTRIBUTION.— Texas, México, Guatemala, Nicaragua, Ecuador,
Peru.
TAXONOMY.—
Eight species, seven of which occur in México and Central America.
Mexinauta aurantia (Carpenter,
1857)
Aplexa
aurantia Carpenter,
1857; Cat. Mazatlan shells: 179.
Bulinus
aurantius
(Carpenter). Binney, 1865; Smiths. Misc.
Coll., 7: 97; figs. 166, 167.
Stenophysa
aurantia
(Carpenter). Taylor, 1966; Veliger, 9:
111.
Mexinauta
aurantia (Carpenter). Taylor, 2003; Rev. Biol. Trop., 51 (suppl.):
77-80; text-fig. 53 (map); text-figs. 54-55 (anatomy); pl. 2, figs. 4-9
(shell).
TYPE LOCALITY.- Mazatlán,
Sinaloa. Syntype BMNH 815819.
DISTRIBUTION RECORDS.— Pacific
coastal region of southern México and northwestern Costa Rica. JALISCO: marsh at Bahia Tenacatita, 1.4 km S
of Mex. Hwy. 200 (19°18.3’ N , 104°48.4’ W).
GUERRERO: nr. Acapulco; Llano Largo, 3 km NE Puerto Marques. COSTA RICA: Guanacaste Prov.; Parque Nacional Palo Verde, edge of marsh 100 m E
of W end of airstrip (10°20.68’ N, 85°20.60’ W); other localities in immediate
vicinity.
Mexinauta gracilentus (Fischer
and Crosse, 1886)
Aplecta
aurantia var. gracilenta Fischer and Crosse, 1886; 2:
87; pl. 39, fig. 4 (shell).
Mexinauta
gracilentus (Fischer
and Crosse). Taylor, 2003; Rev. Biol.
Trop., 51 (suppl.): 76.
TYPE LOCALITY.- Cobán, Dept. Alta
Verapaz, Guatemala. Holotype in MHNP.
DISTRIBUTION RECORDS.- GUATEMALA:
known only from the type locality.
Mexinauta impluviatus (Morelet,
1849)
Physa
impluviata Morelet,
1849; Test Noviss., I: 18.
Aplecta
impluviata (Morelet). Fischer and Crosse, 1886: 91; pl. 30, fig. 3
(shell); pl. 39, fig. 5 (shell).
Mexinauta
impluviatus (Morelet). Taylor, 2003; Rev. Biol. Trop., 51 (suppl.):
80-82; fig. 53 (map); figs. 56-57 (anatomy); pl. 3, figs. 9, 11-12 (shell).
TYPE LOCALITY.— Ditches in
Guatemala City, Guatemala. Syntypes in
the BMNH (3 uncatalogued specimens).
DISTRIBUTION RECORDS.- CHIAPAS:
pond at Pueblo Nuevo, Solistahuacán; San Cristobal de las Casas, 2200 m alt.;
km 65, S of Tuxtla Gutierrez along rd. to Nuena Concordia, 2700 ft. alt.; 15 km
S of Ocozocuautla, along rd. to Villa Flores, 2800 ft. alt. GUATEMALA, Dept. Chimaltenango:
Yepocapa, in fountain; Parque Nacional Los Aposentos, 2.5 km S of
Chimaltenango; ditch nr. Parque Chimaltenango.
Dept. Guatemala: Laguna El
Naranjo, Finca El Naranjo, ca. 5 km NW of Ciudad Guatemala, 1560 m alt.;
environs of Guatemala City; Guatemala City.
Dept. Sacatepéquez: Antigua;
Laguneta Quilisimate, between Santiago Zamora and Santa Catarina Barahona, 1460
m alt.; small stream entering Lago de Dueñas, above Santiago Zamora. Dept.
Santa Rosa: Rio Taxisco, Taxisco. Dept. Zacapa; Zacapa.
Mexinauta laetus (Von
Martens, 1898)
Physa
impluviata var. laeta Von Martens, 1898: 359; pl. 20,
figs. 2-3 (shell).
Mexinauta
laetus (Von Martens). Taylor, 2003; Rev. Biol. Trop., 51 (suppl.):
76.
TYPE LOCALITY.- Vera Paz,
Guatemala. Lectotype ZMB 51234a (Kilias,
1961).
DISTRIBUTION RECORDS.- Known only
from the type locality.
Mexinauta nicaraguana (Morelet,
1951)
Physa
nicaraguana Morelet,
1851; 2: 16.- Von Martens, 1898; 366;
pl. 20, figs. 4, 4a.
Mexinautan
nicaraguana (Morelet). Taylor, 2003; Rev. Biol. Trop., 51 (suppl.):
76.
TYPE LOCALITY.- Lake Nicaragua,
Nicaragua. Syntype BMNH 1893.2.4.610.
DISTRIBUTION RECORDS.- Known only
from the type locality.
Mexinauta nitens (Philippi,
1841)
Physa
nitens Philippi,
1841, in Martini & Chemnitz, 32:
5, pl. 1, figs. 1-2.
Aplecta
nitens
(Philippi). Fischer and Crosse, 1876; 2:
88; pl. 39, figs. 1-1b.
Mexinauta
nitens (Philippi). Taylor, 203; Rev. Biol. Trop., 51 (suppl.):
82-85; fig. 53 (map); figs. 58-65 (anatomy); pl. 1, figs. 5-7 (shell).
Bulinus
berlandierianus
Binney, 1865; Amer. Jour. Conch., 1: 51; pl. 7, fig. 8 (shell).
Aplecta
nitens var. acutalis Fischer and Crosse, 1886; 2:
88; pl. 39, figs. 2-2a.
Aplecta
nitens var. gigantea Fischer and Crosse, 1886; 2:
89; pl. 39, figs. 3-3a.
TYPE LOCALITY.- Physa nitens: “Mexiko”. Bulinus
berlandierianus: Texas, in the vicinity of Matamoros, Tamaulipas.
DISTRIBUTION RECORDS.- Coastal
lowlands along the Gulf of México from the vicinity of Brownsville, Texas south
to Campeche. TAMAULIPAS: Hacienda Acuña, 808 m alt. VERACRUZ: Charco de Buenos Aires; Misantla;
Veracruz; Jalapa; Córdoba; Cerro de las
Mesas, between Ignacio de la Llave and Piedras Negras; Cosamaloapan. TABASCO.
Pasture pond at km 21.1, NW of Taxco, on rd. Nacajuco to Villahermosa;
pond beside Mex. Hwy. 180, 1 km N of rd. to Ignacio Allende; 2.3 km NE of Playa
Miramar; marsh about 1 km N of junction Tacotalpa-Tapijulpa rd. with rd. to
Teapa; pond beside Jalapa-Tacolapa rd. 2 km S of rd. to Guanal, N of Rancho
Nuevo. CAMPECHE. Ditch beside Mex. Hwy. 180, 0.5 km E of Nuevo
Progreso.
Mexinauta princeps (Phillipi,
1846)
Physa
princeps Phillipi,
1846; in Küster, Martini &
Chemnits, Syst. Conch.-Cab., ed. 2 (Limnaeaceen): 66; pl. 1, fig. 11.
Aplexa
princeps (Phillipi). Bequaert & Clench, 1933; Pub. Carnegie
Inst. Wash., (431): 539.
Stenophysa
princeps (Phillipi). Rehder, 1966; Proc. Biol. Soc. Wash., 79:
280.
Mexinauta
princeps (Phillipi). Taylor, 2003; Rev. Biol. Trop., 51 (suppl.):
86-88.; text-fig. 53 (map); text-figs. 66-67 (anatomy); pl. 2, figs. 1-3
(shell); pl. 3, fig. 13 (shell).
TYPE LOCALITY.— Yucatán. Holotype ANSP 21184a (H. B. Baker, 1964:
155).
DISTRIBUTION RECORDS.- Yucatán Peninsula, from Yucatán and Quintana
Roo, México through northern Guatemala and Belize (Taylor, 2003). QUINTANA ROO: Marsh 0.5 km W of Puerto Morelos; Laguna de
Cobá, Cobá, Rio Honda. YUCATÁN: 4 km S
of Purto Progresso; Dzadz Cenote, ca. 1 km SW of Chichén Itzá. GUATEMALA, Dept. Petén: Aguada 0.5 km from La Libertad rd. to San Francisco;
small dry aguada just SW of San Benito (opposite Flores on mainland). BELIZE, Corazal
Dist.: Louisville Forest. Orange Walk Dist.: Orange Walk.
Genus Mayabina Taylor, 2003
Mayabina Taylor, 2003; Rev. Biol. Trop., 51
(suppl.): 88-89.
TYPE SPECIES.- Physa spiculata Morelet, 1849.
DISTRIBUTION.— From Oaxaca and
Veracruz, México, south to Costa Rica; Ecuador to northernmost Chile (Taylor,
2003).
TAXONOMY.- Taylor (2003) lists
eleven species, ten of which occur in México and Central America.
Mayabina bullula (Crosse
& Fischer, 1882)
Aplecta
bullula Crosse
& Fisher, 1882; Jour. de Conchyl., 30: 334.- Fischer & Crosse, 1886, 2: 91; figs.
6-6b.
Mayabina
bullula (Crosse
& Fischer). Taylor, 2003; Rev. Biol.
Trop., 51 (suppl.): 92-94; fig. 68 (map); figs. 70-75 (anatomy); pl. 3, figs.
1-3 (shell).
TYPE LOCALITY.— Tuxpan,
Veracruz. Holotype lost (Taylor, 2003).
DISTRIBUTION RECORDS.- Known only
from the vicinity of the type locality.
VERACRUZ. Tuxpan; pool on W side
of Mex. Hwy. 180, 8.8 km N of Rio Octopan (19°33.1’ N, 96°23.6’ W).
Mayabina petenensis Taylor,
2003
Mayabina
petenensis Taylor,
2003; Rev. Biol. Trop., 51 (suppl.): 96-97; fig. 69 (map); fig. 77 (anatomy),
pl. 3, fig. 10 (shell).
TYPE LOCALITY.- Guatemala, Dept.
Petén, aguada at NE side of La Libertad
(16°47.3’ N, 90°06.49’ W); 200 m alt.
DISTRIBUTION RECORDS.- Known only
from the Dept. Petén, Guatemala. Taylor
(2003) lists numerous localities from there.
Mayabina pliculosa (Von
Martens, 1898)
Physa
fuliginea var. pliculosa Von Martens; 1898: 361; pl.
20, figs. 11-12.
Stenophysa
pliculosa (Von
Martens). Tonn, et al, 1964; Rev. Biol. Trop., 12: 60.
Mayabina
pliculosa (Von
Martens). Taylor, 2003; Rev. Biol.
trop., 51 (suppl.): 97-99; fig. 69 (map); figs. 78-79 (anatomy); pl. 4, fig.
10-11 (shell).
TYPE LOCALITY.- Rio Reventazón at Ujarrás, Cartago Prov.,
Costa Rica.
DISTRIBUTION RECORDS.- Western
Costa Rica from the Central Valley northwestward to the southern tip of the
Nicoya Peninsula (Taylor, 2003). COSTA
RICA, Alajuela Sur Prov.: Sachi
Sar. Cartago
Prov.: Rio Turialba, Turialba; Turialba; Alto de Ochomogo; Aguas Calientes,
nr. Cartago; Quebrada Barahona, 1380 m alt.; Juan Viññs. Guanacaste
Prov.: ditch 3-4 km toward Hacienda La Taboga, from Cañas-Bebedero rd. Puntarenas
Prov.: La Hacienda, 1 km SW of
Concepción-Bebedero rd. (9°45.08’ N, 85°00.44’ W). San José Prov.: Pavas; Uruca; San José; Rio
Torres, San José; Rio Torres; Ocloro.
Mayabina polita
Taylor, 2003
Aplecta
cisternina var. abbreviata Fischer and Crosse, 1886; 2: 95;
pl. 30, fig. 9 (shell). (Not of Beck,
1838.)
Aplecta
cisternina var. gracilis Fischer & Crosse, 1886, 2:
95; pl. 30, figs. 10-10b (shell). (Not
of Noulet, 1854.)
Aplexa spiculata var. gracilis
(Fischer and Crosse). Bequaert &
Clench, 1936; Pub. Canegie Inst. Wash., (457): 70; pl. 2, figs. 5-8 (shell).
Aplecta
cisternina var. minor Fischer & Crosse, 1886; 2: 95;
pl. 30, fig. 8 (shell). (Not of Beck,
1838.)
Mayabina
polita Taylor,
2003; Rev. Biol. Trop., 51 (suppl.): 99-102; fig. 69 (map); figs. 40-84
(anatomy); pl. 5, figs. 1-2 (shell).
TYPE LOCALITIES.— Aplecta abbreviata: Mérida, Yucatán;
holotype lost. Aplecta gracilis: vicinity of Mérida, Yacatan; holotype lost. Mayabina
polita: pasture pool 50 m W of Rio Tulija, 1.5 km S of Mex. Hwy. 186 toward
Zopo Norte (17°39.6’ N, 92°24.7’ W).
Holotype CAS 114783.
DISTRIBUTION.- Southeastern
México, in Tabasco and northern Chiapas through the Yucatán Peninsula in
eastern Campeche, Yucatán and Quintana Roo.
Taylor (2003) provides numerous locality records.
Mayabina
sanctijohannis Taylor, 2003
Mayabina
sanctijohannis Taylor,
2003; Rev. Biol. Trop., 51 (suppl.): 102-104; figs. 15, 19 (maps); fig. 85
(anatomy); pl. 3, figs. 5-6 (shell).
TYPE LOCALITY.— Barra del
Colorado, Limón Prov., Costa Rica (10°46.37’ N, 83°35.27’ W). Holotype: CAS 114790.
DISTRIBUTION RECORDS.— COSTA
RICA, Limón Prov.: Barra del
Colorado; marsh on left bank of Rio Colorado.
Mayabina spiculata
(Morelet, 1849)
Physa
spiculata Morelet,
1849, 1: 18.- Von Martens, 1898: 366.
Aplecta
spiculata
(Morelet). Fischer & Crosse, 1886:
93; pl. 27, fig. 13 (shell).
Aplexa
spiculata (Morelet). Bequaert & Clench, 1936; Pub. Canegie
Inst. Wash., (457): 69; pl. 2, figs. 1-4.-
Harry,1950; Occ. Pap. Mus. Zool. Univ. Mich., (524): 23.
Physa
(Aplexa) spiculata Morelet. Branson
& McCoy, 1965; Univ. Colorado Stu., Ser. Biology, 13: 12.
Stenophysa
spiculata
(Morelet). Rehder, 1966; Proc. Biol.
Soc. Wash., 79: 280.
Aplexa
nitens var. spiculata (Morelet). Pilsbry, 1891; Proc. Acad. Nat. Sci. Phila.,
43: 325.
Physa
cisternina Morelet,
1851, 2: 15.
Aplecta
cisternina (Morelet). Fischer & Crosse; 1886, 2: 94; pl. 30,
fig. 7 (shell).- Pilsbry, 1891; Proc.
Acad. Nat. Sci. Phila., 43: 325.
Mayabina
spiculata
(Morelet). Taylor, 2003; Rev. Biol.
Trop., 51 (suppl.): 104-107; fig. 68 (map); figs. 86-87 (anatomy); pl. 3, figs.
4, 7 (shell).
TYPE LOCALITY.- Cd. Campeche,
Campeche Types lost.
DISTRIBUTION RECORDS.- Yucatan
Peninsula in eastern Campeche, Yucatan and Quintana Roo (Taylor, 2003). CAMPECHE: Cd. Campeche; Pantel Aguada. Nr.
Champoón; aguada 3.5 mi. S of Champotón; 26 km SE Chapotó; Colonia Lopez
Mateos, 93 km E of Silvituc. QUINTANA
ROO: San Gerbacio, Isla Cozumel; Ruinas Kuhunlich; potreros at Tomás
Garrido. YUCATAN: cienaga 2 km SW of
Progreso, Cerro Isla; south side of cienaga nr. Progreso; irrigation cistern
ca. 5 km N of Mérida; charcos at Pemex plant, 4 km S of Puerto Progreo; Izamal;
Cenote Shkolak, and Tekanto; Hatal Aguada, nr. Mérida; Yunku Aguada, nr. Yunku;
Chichen-Itzá; aguada S of Libre Unión.
Mayabina tapanensis (Crosse
& Fischer, 1882)
Aplecta
tapanensis Crosse
& Fischer, 1882; Jour. de Conchyl., 29: 334.
Aplecta
spiculata var. tapanensis Fischer & Crosse, 1886,
2: 93: pl. 30, figs. 2-2b (shell).
Physa
tapanensis (Crosse
& Fischer). Von Martens, 1898: 367.
Aplexa
tapanensis guatemalensis (Crosse
& Fischer). Hinkley, 1920; Nautilus, 34: 38.
Mayabina
tapanensis (Crosse
& Fischer). Taylor, 2003; Rev. Biol.
Trop., 51 (suppl.): 107-109.
TYPE LOCALITY.- Near Tapana [San Pedro Tapanatepec}, Oaxaca,
Mexico (16°21’ N, 94°12’ W). Holotype
lost (Taylor, 2003).
DISTRIBUTION RECORDS.-
Southeastern Oaxaca to southern Guatemala (Taylor, 2003). OAXACA: San Pedro Tapanatepec, in northern
part of town, in a marsh 100 m E of km post 90 on Mex. Hwy. 190. CHIAPAS: 65 km S of Tuxtla Gutierrez, along
rd. to Nueva Concordia. GUATEMALA, Dept. Guatemala: pools along railroad across Laguna Amatitlán from Laguna
Station. Dept.Sololá: San
Buenaventura, 1 km NW of Panajachel, 1565 m alt.; Lago de Atitlán at mouth of
Rio Catarina, 1.5 km NW of Panahachel.
Mayabina tempisquensis
Taylor, 2003
Mayabina
tempisquensis Taylor,
2003; Rev. Biol. Trop., 51 (suppl.): 109-110; text-fig. 15, 68 (map); text-fig.
90 (anatomy); pl. 4, fig. 4 (shell).
TYPE LOCALITY.— Parque Nacional
Palo Verde, edge of marsh 100 m E of W end of airstrip, Guanacaste Prov., Costa
Rica (10°20.68’ N, 85°20.6’ W). Holotype
CAS 146092.
DISTRIBUTION.- COSTA RICA, Guanacaste Prov.: edge of marsh 100 m E
of W end of airstrip, Parque Nacional Palo Verde; ornamental pool inside Hotel
Las Espuelas, 2 km SE of La Libertad; Rio Cañas, 9 km S of Cartagena.
Genus Tropinauta Taylor, 2003
Tropinauta
Taylor, 2003; Rev.
Biol. Trop., 51 (suppl.): 110-111.
TYPE SPECIES.- Tropinauta sinusdulcensis Taylor, 2003.
DISTRIBUTION.- Costa Rica.
TAXONOMY.- A single species is
currently recognized.
Tropinauta
sinusdulcensis Taylor, 2003
Tropinauta
sinusdulcensis Taylor,
2003; Rev. Biol. Trop., 51 (suppl.): 111; figs. 91-94 (anatomy).
TYPE LOCALITY.- A small stream in
pasture 3 km SE of Gulfito, Puntarenas Prov., Costa Rica (8°36.68’ N, 83°08.48’
W). Holotype CAS 146095.
DISTRIBUTION RECORDS.- COSTA
RICA: known only from the type locality.
TAXONOMY.— Taylor (2003) lists
eleven genera.
DISTRIBUTION.— Holarctic in
distribution, extending into the neotropical realm in Middle America.
Genus Chiapaphysa Taylor, 2003
Chiapaphysa
Taylor, 2003; Rev.
Biol. Trop., 51 (suppl.): 167.
TYPE SPECIES.— Chiapaphysa grijalvae Taylor, 2003.
DISTRIBUTION.— Chiapas, Mexico,
and Costa Rica.
TAXONOMY.- Two species are
included in the genus.
Chiapaphysa grijalvae Taylor,
2003
Chiapaphysa
grijalvae Taylor,
2003; Rev. Biol. Trop., 51 (suppl.): 168-170; fig. 165 (map); figs. 166-167
(anatomy); pl. 8, fig. 7 (shell).
TYPE LOCALITY.— Rio Suchiapa, 2
km SE of Suchiapa, Chiapas (16°36.4’ N, 93°05.0’ W). Holotype CAS 114818.
DISTRIBUTION RECORDS.- Chiapas, in streams of both the Pacific Ocean
and the Gulf of Mexico. CHIAPAS: Rio La
Venta, Las Flores, Mex. Hwy. 190, 18 km E of Cintalapa (16º41.6’ N, 93º33.5’
W); Rio Suchiapa, 2 km SE of Suchiapa (16º36.4’ N, 93º05.0’ W); Rio Santo
Domingo, Puente Santo Domingo, 9 km S of Cupía (16º37.4’ N, 92º59.8’ W); Rio
Santo Domingo, 2 km NE of Julián Grajales (16º30.0’ N, 92º57.4’ W); Rio
Quemado, ca. 1 km SW of Vicente Guerrero (16º25.7’ N, 92º43.5’ W); Baños del
Carmen on Rio Quemado, ca. 1 km from rd. to La Angostura (16º25.4’ N, 92º43.0’
W); Rio San Francisco Ocotal at Mex. Hwy. 200 (16º07.6’ N, 93º47.7’ W); Rio Los
Amates, Villa Flores (16º14.3’ N,
93º01.0’ W); Rio Pando, 1.5 km S of Villa Flores (16º12.8’ N, 93º16.1’
W); Rio Ningunito 1 km SW of Revolución Mexicana (16º09.6’ N, 93º04.8’ W); Rio
Custepeques 2 km SE of Independencia (16º04.6’ N, 92º49.8’ W); irrigation ditch
1.5 km WNW of Independencia (16º05.8’ N, 92º51.7’ W).
Chiapaphysa pacifica Taylor,
2003
Chiapaphysa
pacifica Taylor,
2003; Rev. Biol. Trop., 51 (suppl.): 170-172; fig. 165 (map); fig. 168
(anatomy); pl. 8, fig. 8 (shell).
TYPE LOCALITY.— Rio Tenorito,
Hacienda La Pacifica, Dept. Guanacaste, Costa Rica (10°29.02’ N, 85°09.58’
W). Holotype CA 114784.
DISTRIBUTION RECORDS.- Known only from Costa Rica. COSTA RICA, Prov. Guanacaste: Rio Salina, 1 km NW of Soley (11º01.05’ N,
85º40.46’ W); Parque Nacional de Guanacaste, Quebrada Aserradero (10º53.82’ N,
85º33.74’ W); Parque Santa Rosa, Sendero Las Mesas (10º50.95’ N, 85º36.65’ W)
Parque Nacional Santa Rosa, Sendero Los Patos (10º49.57’ N, 85º37.84’ W); Rio
Tempesquito, Vado Esperanza (10º47.38’ N, 85º33.11’ W); Rio Cabuya; Rio
Tenerito, Hacienda La Pácifica, at rd. to “Ecomuseo” (10º29.02’ N, 95º09.58’
W); Hacienda La Pácifica, ditch 0.9 km W of Lechería (10º28.19’ N; 85º08.97’
W); rivulet 1.65 km S of Parque Nacional Barra Honda headquarters (10º09.92’ N,
85º22.84’ W).
Genus Haitia Clench & Aguayo, 1932
Haitia
Clench &
Aguayo, 1932; Proc. New England Zoological Club, 13: 37.- Taylor, 2003; Rev. Biol. Trop., 51 (suppl.):
128.
TYPE SPECIES.- Physa elegans Clench & Aguayo, 1932.
DISTRIBUTION.- Tropical and
temperate North America, Central America, the West Indies, South America in
Colombia, and along the Pacific slope from Peru to central Chile (Taylor,
2003).
TAXONOMY.- Taylor recognizes
fifteen species, five of which occur in
Mexico and Central America.
Haitia lacustris (Clessin,
1886)
Physa
lacustris Clessin,
1886; Syst. Conch. Cab., ed. 2, 338: 334; pl. 48, fig. 9.
Haitia
lacustris (Clessin). Taylor, 2003; Rev. Biol. Trop. 51 (suppl.):
133.
TYPE LOCALITY.- Lago de
Coatepeque, Dept. Sanata Ana, El Salvador.
Holotype BMNH 93.2.4.1370.
DISTRIBUTION.- EL SALVADOR: known
only from the type locality.
TAXONOMY.— Possibly only an
ecotype of Haitia mexicana (Taylor,
2003).
Haitia mexicana (Philippi,
1841)
Physa
mexicana Philippi,
1841; in Küster, Martini & Chemnitz, Syst. Conch. Cab., ed. 2,
(Limnaeaceen): 5; pl. 1, figs. 3, 4 (shell).
Haitia
mexicana (Philippi). Taylor, 2003; Rev. Biol. Trop., 51 (suppl.):
140-146; text-figs. 136-139 (anatomy); pl. 7, figs. 1-15 (shell).
Taylor lists the following forty-two
names as synonyms.
Physa
solida Philippi,
1841; in Küster, Martini & Chemnitz, Syst. Conch. Cab., ed. 2, (Limnaeaceen):
6.
Physa
squalida Morelet,
1851; Test. Noviss., II: 16.
Physa
humerosa Gould,
1855; Proc. Boston Soc. Nat. Hist., 5: 128.
Physa
virgata Gould,
1855; Proc. Boston Soc. Nat. Hist., 5: 128.
Physa
fosheyi Lea, 1864;
Proc. Acad. Nat. Sci. Phila., 16: 114.
Physa
grosvenori Lea,
1864; Proc. Acad. Nat. Sci. Phila., 16: 114.
Physa
halei Lea, 1864;
Proc. Acad. Nat. Sci. Phila., 16: 114.
Physa
tenuissima Lea,
1864; Proc. Acad. Nat. Sci. Phila., 16: 114.
Physa
parva Lea, 1864;
Proc. Acad. Nat. Sci. Phila., 16: 115.
Physa
striata Lea, 1864;
Proc. Acad. nat. Sci. Phila., 16: 115.
Physa
traski Lea, 1884;
Proc. Acad. Nat. Sci. Phila., 16: 115.
Physa
mexicana var. minor Von Martens, 1865; Malak. Blätt.,
12: 57.
Physa
mexicana var. ovslis Von Martens, 1865; Malak. Blätt.,
12: 58.
Physa
mexicana var. parva Von Martens, 1865; Malak. Blätt.,
12: 58.
Physa
mexicana var. minima Von Martens, 1865; Malak. Blätt.,
12: 58.
Physa
sparsestriata
Tryon, 1865; Amer. Jour. Conch., 1: 224.
Physa
distinguenda Tryon,
1865; Amer. Jour. Conch., 1: 225.
Physa
politissima Tryon,
1865; Amer. Jour. Conch., 1: 226.
Physa
dorbigniana Lea,
1867; Jour. Acad. Nat. Sci. Phila., 6: 166.
Lymnaea
ambigua Pease,
1870; Amer. Jour. Conch., 6: 6.
Lymnaea
compacta Pease,
1870; Amer. Jour. Conch., 6: 6.
Physa
mexicana minor
Strebel, 1874: 51.
Physa
berendti “Danker”
Strebel, 1874: 55.
Physa
boucardi Crosse
& Fischer, 1882; Jour. de Conchyl., 29: 334.
Physa
strebeli Crosse
& Fischer, 1882; Jour. de. Conchyl., 29: 335.
Physa
tehuantepecensis
Crosse & Fischer, 1882; Jour. de Conchyl., 29: 335.
Physa
mexicana var. acutissima Fischer & Crosse, 1886:
100.
Physa
mexicana var. conoidea Fischer & Crosse, 1886:
100.
Physa
mexicana var. plicata Fischer & Crosse, 1886:
100.
Physa
tolucensis Fischer
& Crosse, 1886: 100.
Physa
intermedia Fischer
& Crosse, 1886: 104.
Physa
polakowskii Clessin,
1886; in Martini & Chemnitz,
Syst. Conch. Cab., Physa, 338: 352.
Physa
cupreonitens Cockerelll,
1889; Jour. Conchol., 6: 63.
Physa
penicillata
Hemphill, 1890; Cat. N. Amer. Shells: 19.
Physa
osculans rhyssa
Pilsbry, 1899; Proc. Acad. Nat. Sci. Phila., 51: 401.
Physa
rhomboidea Crandall,
1901; Nautilus, 15: 44.
Physa
virgata mut. alba
Cockerell, 1902; Jour. Malac., 9: 138.
Physa
cradalli F. C.
Baker, 1906; Trans. Acad. Sci. St. Louis, 16: 8.
Physa
balteata Preston,
1907; Ann. & Mag. Nat. Hist., 20: 497.
Physa
bottimeri Clench,
1924; Nautilus, 38: 12.
Physa
marci F. C. Baker,
1924; Nautilus, 38: 15.
Physa
humerosa interioris Pilsbry,
1932; Nautilus, 45: 139.
TYPE LOCALITY.- Physa mexicana: “Mexiko”. Holotype:
destroyed in SMF during 1939-1945 war.
DISTRIBUTION RECORDS.- Continuous
from the Central Valley of California southward over the Mexican Plateau to
Oaxaca and Tabasco, and south through Central America at least to Colombia; eastward
through southern Nevada, central and southern Utah, Arizona and New Mexico
across the southern Great Plains from Colorado to Illinois, Missouri and
Louisiana. Patchy occurrences in
Washington, Oregon and Idaho (Taylor, 2003).
In
Mexico recorded by Taylor from the states of Campeche, Colima, Distrito
Federal, Guerrero, Michoacán, Oaxaca and Tabasco. In Costa Rica recorded form the Providencias
Cartago, Heredia, Puntarenas and San José.
Haitia moreleti Taylor,
2003
Haitia
moreleti Taylor, 2003;
Rev. Biol. Trop., 51 (suppl.): 146-147; fig. 15 (map); fig. 140 (anatomy), pl.
6, fig. 2 (shell).
TYPE LOCALITY.— Marshy border of
Lago de Petén-Itzá, Santa Elena, Dept. Petén, Guatemala. Holotype CAS 114821.
DISTRIBUTION RECORDS.— GUATEMALA:
known only from the immediate vicinity of the type locality (Taylor, 2003).
Haitia patzcuarensis (Pilsbry,
1891)
Physa
osculans var. patzcuarensis Pilsbry, 1891a:
9.—Pilsbry, 1891b: 323; pl. 15, fig. 5.
Haitia
patzcuarensis (Pilsbry). Taylor, 2003; Rev. Biol. Trop., 51 (suppl.):
149-150; text-fig. 143 (penial complex), text-fig. 144 (egg mass); pl. 6, fig.
1 (shell).
TYPE LOCALITY.— Lago de
Patzcuaro, Michoacán, Mexico. Holotype
ANSP 61629.
DISTRIBUTION RECORDS.— MICHOACAN:
known only from Lago de Patzcuaro.
Haitia solidissima (Pilsbry,
1920)
Physa
solidissima Pilsbry,
1920; Proc. Acad. Nat. Sci. Phila., 71: 219; pl. 11, fig. 8.
Haitia
solidissima (Pilsbry). Taylor, 2003; Rev. Biol. Trop., 51 (suppl.):
135.
TYPE LOCALITY.— Laguna de Chapala,
Jalisco, Mexico. Holotype ANSP 107482.
DISTRIBUTION.— JALISCO: known
only from Laguna de Chapala.
Genus Ultraphysella Taylor, 2003
Ultraphysella
Taylor, 2003; Rev.
Biol. Trop., 51 (suppl.): 191.
TYPE SPECIES.- Ultraphysella sinaloae Taylor, 2003.
DISTRIBUTION.— Known from Sinaloa
and Nayarit, northwestern Mexico.
TAXONOMY.- One species is known
for the genus.
Ultraphysella sinaloae
Taylor, 2003
Ultraphysella
sinaloae Taylor,
2003; Rev. Biol. Trop., 51 (suppl.): 192-195; fig. 15 (map); figs. 187-191
(anatomy); pl. 9, fig. 1 (shell).
TYPE LOCALITY.- A pool at rd. 2.5
mi from Villa Unión toward Siqueiros, Sinaloa, Mexico (23°13.4’ N, 106°12.5’
W). Holotype CAS 146096.
DISTRIBUTION.— NAYARIT: Rio de
Palillo, El Palillo (21º38.1’ N, 105º08.4’ W); pond beside Mex. Hwy. 68D, 0.7
km E of exit to San Pedro Lagunillas, 0.1 km E of km post 16 (21º12.3’ N,
104º45.1’ W). SINALOA: pool in culvert on rd. from Villa Unión to Siqueiro, 4.2 km from Mex. Hwy. 15
in Villa Unión (23º13.4’ N, 106º12.5’ W); stream at Mex. Hwy. 15, 4.2 km NW of
Ricón Verde (22º55.7’ N, 105º49.7’ W); marshy pool beside Mex. Hwy. 15, 7 km S
of Ojo Agua de Palmillas, and 41 km S of Escuinapa (22º33.6’ N, 105º35.0’ W).
Superfamily ELLOBIOIDEA
Family CARYCHIIDAE
Jeffreys, 1829
Genus Carychium Müller, 1774
Carychium Müller, 1774; Verm. terr. et fluv.
Hist., 2: 125.
TYPE SPECIES.- Carychium minimum Müller, 1774.
DISTRIBUTION.- Holarctic realm in
general. Extending south to Java and the
Philippine Islands in the Old World, and to
TAXONOMY.- Eleven species occur
in
Carychium
costaricensis Von Martens, 1898
Carychium
costaricensis Von
Martens, 1898: 253; pl. 19, figs. 17, 18 (shell).- Hinkley, 1920; Nautilus, 34: 52.
TYPE LOCALITY.-
DISTRIBUTION
Carychium
mexicanum Pilsbry, 1891
Carychium
exiguum mexicanum
Pilsbry, 1891; Proc. Acad. Nat. Sci. Phila., 43: 318-319; pl. 14, figs. 7, 8, 9
(shell).- Von Martens, 1898: 352; pl.
19, figs. 15, 16 (shell).- Baker, 1930a:
2.- Goodrich & van der Schalie,
1937; Misc. Pub. Mus. Zool. Univ. Mich., (34): 30.- Pilsbry, 1948; Land Moll. N. Amer. , 2: 1060;
fig. 566c (shell).- Basch, 1959; Occ.
Pap. Mus. Zool. Univ. Mich., (612): 12.-
Correa-Sandoval, 1993; Rev. Biol. Trop., 41: 675.- Correa-Sandoval, Gutierrez & Reza, 1998:
13.- Correa-Sandoval & Castro, 2002;
Acta Zool. Mex., (86): 235.-
Correa-Sandoval & Rodriguez, 2005: 59.
Carychium
mexicanum
Pilsbry. Hubricht, 1985: 6; map 14.
TYPE LOCALITY.- Hills around
Orizaba, Veracruz, 500 ft. above the town.
Holotype ANSP 61628a (Baker, 1964: 193).
DISTRIBUTION RECORDS.- Widely
distributed across the southern United States from Georgia and Florida west to
Texas, and south to Guatemala. GUATEMALA
(Goodrich & van der Scahlie, 1937). Dept.
Escuintla: Costa Cuca (Von Martens, 1898).
Dept. Huehuetenango: Hacienda
Helvetia, in upper Cholhuitz. Dept. Izabal: Chama. Dept.
Petén: Tikal (Basch, 1959). NUEVO
LEON: Diente, nr. Monterrey (Pilsbry, 1948); Santiago (Correa-Sandoval, 1993);
near Laguna de Sanchez, Parque Vitro (25°23’24” N, 100°12’57” W); km 66, road
from Linares-san Roberto (24°45’22” N 100°02’18” N) (Correa-Sandoval &
Rodriguez, 2005). PUEBLA: near Necaxa,
2625-5500 ft. alt. (Baker, 1930a). SAN
LUÍS POTOSÍ: carr. Jalpan-Xilitla, 6 km SW of Xilitla, 830 m alt. (21°23’10” N,
99°03’56” W) (Correa-Sandoval, et al,
1998). TAMAULIPAS: Tampico; Ejida Santa
Juana, W of Nuevo Padilla (24°02’17” N, 98°59’18” W) (Correa-Sandoval &
Castro, 2002). VERACRUZ: Cordoba, 2700
ft. alt. (Baker, 1930a); Texolo (Pilsbry, 1948).
Genus Coelostela Benson, 1864
Coelostela
Benson, 1864; Ann.
Mag. Nat. Hist. (3), 13: 136.
TYPE SPECIES.- Coelostela scalaris Benson, 1864.
DISTRIBUTION.- India, southern
Arabia, Egypt, Syria and southern Spain; northeastern Mexico.
TAXONOMY.- Many species. The single Mexican species may be introduced.
Coelostele
tampicoensis (Pilsbry, 1907)
Coelostele
tampicoensis
(Pilsbry). Pilsbry, 1908; Man. Conch.,
19: 346.- Correa-Sandoval, Gutierrez
& Raza, 1998: 15.
TYPE LOCALITY.- Tampico,
Tamaulipas; in river drift. Holotype ANSP 93786a (Baker, 1964: 193).
DISTRIBUTION RECORDS.- SAN LUÍS
POTOSÍ: numerous localities (Correa-Sandoval, et al, 1998). TAMAULIPAS:
known only from the type locality.
SYSTELOMMATOPHORA
Superfamily VERONICELLOIDEA
Family
VERONICELLIDAE Gray, 1840
DISTRIBUTION.- Pantropical.
TAXONOMY.- Approximately 300
species have been proposed, but many of these probably are synonyms (Thomé,
Santos & Perdott, 1997).
Some veronicellid slugs are readily
distributed by human agency. They have
been introduced into many places in tropical and subtropical regions, often
with dire agricultural consequences because they feed upon newly emergent
seedlings. Four species are native to
the study area. Five others have been
introduced.
Genus Belocaulus Hoffmann, 1925
Belocaulus
Hoffmann, 1925;
Jenaische Zeit, für Naturw., 61: 198, 245.-
Thomé, 1975; Iheringia, Zool., 48; : 11-12.
TYPE SPECIES.- Vaginula angustipes Heynemann, 1885.
DISTRIBUTION.- Brazil, Paraguay
and Argentina. Introduced elsewhere.
TAXONOMY.- The genus includes two
species.
Belocaulus angustipes
(Heynemann, 1885)
Vaginulus
angustipes
Heynemann, 1885; Jahrb. Deut, Malakozool. Gesell., 12: 275-277.- Simroth, 1914; Mem. Soc. Neuchât. Sci. nat.,
35: 324-325; pl. 14, figs. 8-11.- Thomé,
1969; Arch. für Moll., 99: 334-335; pl. 7, figs. 7-9 (animal); text-figs. 5-7,
39 (reproductive anatomy).
Belocaulus
angustipes
(Heynemann).- Hoffmann, 1925; Jenaische
Zeit, für Naturw., 61: 198.-
Lopez-Pitone and Thomé, 1981; Rev. Bras. Biol., 41: 586-591.- Thomé, 1993; Biociências, 1: 70.- Thomé, Santos & Pedott, 1997; Proc. Biol.
Soc. Wash., 110: 521.
TYPE LOCALITY.- Taquera, Rio
Grande do Sul, Brazil. Lectotype SMF
194290 (Thomé, 1969: 334).
DISTRIBUTION RECORDS.- Argentina,
Brazil, Paraguay, Colombia. Introduced
into Honduras, Florida, Alabama, Louisiana and Texas. HONDURAS: no specific locality (Thomé, et al, 1997).
Genus Diploselenodes Thomé, 1975
Diploselenodes Thomé, 1975; Ihringia, Zoologie,
(48): 13-14.
TYPE SPECIES.- Vaginula bielanbergi Semper, 1885.
DISTRIBUTION.- Chile, Brazil,
Ecuador, Venezuela, Guayana, Lesser Antilles, Puerto Rico; Hispaniola, Central
America. Introduced elsewhere.
TAXONOMY.- Six species are
recognized (Thomé, 1975).
Diplosolenodes
occidentalis (Guilding, 1825)
Onchidium
occidentalis
Guilding, 1825, Trans. Linn Soc., 14: 323: pl. 9, figs. 9-12.
Vaginulus
occidentalis
(Guilding). Fischer, 1871; Nouv. Arch.
Mus. Hist. Nat., 7: 164.
Vaginulus (Latipes) occidentalis
(Guilding). H. B. Baker, 1925; Proc.
Acad. Nat. Sci. Phila., 77: 174-177; pl. 5, figs. 18, 19, 20 (reproductive
anatomy).
Cylindrocaulus
occidentalis
Hoffmann, 1925; Jenaische Zeit, für Naturw., 61: 144, 234.
Diplosolenodes
occidentalis
(Guilding).- Thomé, 1975; Ihringia,
Zoologie, (48): 14.- Thomé, 1985; Rev.
Bras. Zool., 2 (6): 411-417.- Thomé,
1993: 71.- Thomé, Santos, & Pedott,
1997; Proc. Biol. Soc. Wash., 110: 522. -
Pérez & López, 2002: 244-246; map.
TYPE LOCALITY.- Puerto Rico.
DISTRIBUTION RECORDS.- Guayana,
Venezuela, Colombia, Panama, Costa Rica, El Salvador, Nicaragua, Honduras and
Dominica. Introduced into Hawaii and
Bahamas (Thomé, Santos & Pedott, 1997: 522). COSTA RICA, Puntarenas Prov.: Rio Surubres, Bonnefil Finca (Thomé, et al, 1997). HONDURAS: Bantua [?] (Thomé et al, 1997). NICARAGUA, Pacific versant of the country
(Pérez & López, 2002). Dept. Rio San Juan (Pérez & López, 2002).
Dept. Masaya: Ticuantepe (Thomé, et
al, 1997). PANAMA: Panama Prov.: Punta de Piña; Panama
City. Canal Zone: Flamenco Island (Thomé, et al, 1997). EL SALVADOR:
(Thomé, et al, 1997).
Diplosolenodes
olivacea (Stearns, 1871)
Veronicalla
olivacea Stearns,
1871; Conch. Mem., 8: 1.
Vaginulus (latipes) olivaceus
(Stearns). H. B. Baker, 1925; Proc.
Acad. Nat. Sci. Phila., 77: 177-178; pl. 5, figs. 15, 16, 17 (reproductive
anatomy).
?Cylindrocaulus
olivaceus
(Stearns). Hoffmann, 1925; Jen. Zeit,
für Naturw., 61: 232; pl. 5, fig. 45d.
Diplosolenodes
olivacea
(Stearns).- Thomé, 1975; Ihringia,
Zoologie, (48): 14.- Thomé &
Lopez-Pitoni, 1976; Rev. Bras. Biol., 36: 710-712.
TYPE LOCALITY.- Nicaragua, Dept.
León, Polvón (12º29’ N, 87º00’ W). (H.
B. Baker, 1925). Lectotype USNM 39160.
DISTRIBUTION.- NICARAGUA: known
only from the type locality. COSTA RICA,
Limon Prov.: Pacuarito (Hoffmann,
1925).
TAXONOMY. This may be a synonym
of Diplosoleodes occidentalis (Baker,
1925c).
Genus Leidyula H. B. Baker, 1925
Leidyula
H. B. Baker, 1925;
Proc. Acad. Nat. Sci. Phila., 77: 158.–
Hoffmann, 1927: 211.- Pilsbry,
1948; Land Moll. N. Amer., 2: 1063.-
Forcart, 1967; Ann. Natal Mus., 18: 251-252.- Thomé, 1975; Ihringia, Zoologie, (48): 17-19.
TYPE SPECIES.- Vaginula moreleti Fischer, 1971.
DISTRIBUTION.- Guatemala, Mexico,
Jamaica, Puerto Rico, Cuba, southeastern United States.
TAXONOMY.- About twelve species
are recognized.
Leidyula floridana
Leidy, 1851
Vaginulus
floridanus Leidy,
1851; in Binney, Terrestrial
air-breathing mollusks of the United States and the adjacent territories of
North America: 198, 251; pl. 4.
Veronicella
floridana
(Leidy). Binney, 1878; Terr. Moll. N.
Amer., 5: 241; fig. 140, pl. 5, fig. p (jaw & radula).- H. B. Baker, 1925; Proc. Acad. Nat. Sci.
Phila., 77: 167; pl. 4, figs. 12-14 (reproductive anatomy).- Pilsbry, 1948; Land Moll. N. Amer., 2:
1063-1064; figs. 568a, 568b (animal); fig. 568c (reproductive anatomy).
Leidyula
floridana
(Leidy). Naranjo-Garcia, Thomé &
Castillejo, 2007; Rev. Mex. Biod., 78: 42.
TYPE LOCALITY.- Punta Rasa,
Florida. Restricted to Meta-Lee-Chee
Key, Charlotte Harbour, Florida (Thomé, 1989).
Holotype USNM 180860.
DISTRIBUTION RECORDS.- CHIAPAS:
at Rio Coatancito, Tapachula. NUEVO
LEÓN: Hualahuises; Presa Los Magueyes, km 118, Cd. Victoria-Linares Hwy.; Cerro
de la Silla, Guadalupe; Presa la Boca, Santiago; Las Adjuntas, Santiago. VERACRUZ: Poza Rica; Tlapacoyan
(Naranjo-Garcia, Thomé & Castillejo, 2007).
Leidyula moreleti
(Fischer, 1871)
Vaginula
moreleti Fischer,
1871; Arch. Mus. Hist. Nat., 7: 168-169, 175; pl.11, figs. 5, 6
(holotype).- Crosse & Fischer, 1872;
Jour. de Conchyl., 20: 59.- Fischer
& Crosse, 1878; Miss. Sci. Mex., I: 682; pl. 24, fig. 14; pl. 29, figs.
6-14 (anatomy).- Bequaert & Clench,
1936; Publ. Carnegie Inst. Wash., (457): 66.
Veronicella (Leidyula) moreleti
(Crosse & Fischer). Pilsbry, 1919;
Proc. Acad. Nat. Sci. Phila., 71: 219.-
H. B. Baker, 1925; Proc. Acad. Nat. Sci. Phila., 77: 165-167; pl. 4,
figs. 7-11 (anatomy).
Leidyula
moreleti
(Fischer). Thomé, 1971; Iheringia, Zool.
(40): 32-34.- Thomé, Santos &
Pedott, 1997; Proc. Biol. Soc. Wash., 110: 529.- Thomé, 1993: 71.- Correa-Sandoval, Gutierrez & Reza, 1998;
Acta Zool. Mex., (73): 13.-
Correa-Sandoval, 2000; Acta Zool. Mex., (97): 8.- Correa-Sandoval & Castro, 2002; Acta
Zool. Mex., (86): 235.- Naranjo-Garcia,
Thomé & Catillejo, 2007; Rev. Mex. Biod., 78: 42-44.
Vaginulus
mexicanus Strebel
& Pfeffer, 1882: 130-131, 140-141, 145; pl. 19, figs. 1-19, 21, 23, 26-27
(anatomy).- Thomé, 1969; Arch. Moll.,
99: 353-354; pl. 9, figs. 19-21 (animal); text-figs. 23-25, 46 (reproductive
anatomy).
Veronicella
mexicana betheli
Cockerell, 1913; Nautilus, 27: 1.
TYPE LOCALITIES.- Vaginula moreleti: Palenque,
Chiapas. Holotype designated as pl. 11,
figs. 5 and 6, in Fischer, 1871
(Thomé, 1971). Vaginulus mexicanus: Mexico; lectotype SMF 194283 (Thomé, 1969)
DISTRIBUTION.- Colombia,
Nicaragua, Honduras, Guatemala and Mexico.
Introduced into Florida and the West Indies (Thomé, Santos & Pedott,
1997: 529). GAUTEMALA, Dept. Izabal: Jacola; Livingston
(Pilsbry, 1919; Baker, 1925); Puerto Barios (Cockerell, 1913). HONDURAS (Cockerell, 1913).- NICARAGUA, Dept. León: Polvón. MEXICO,
CAMPECHE: Isla de Carmen; Cd. Campeche (Thompson, 1967). CHIAPAS: Palenque (type locality). NAYARIT: Tepic (Baker, 1925);
Cacaprieto. OAXACA: Mecos; Tehuantepec,
Cd. Valles (Baker, 1925). SAN LUIS
POTOSÍ: Cd. Valles (Hinkley, 1907); Huichihuayán; Mecos; Tamazunchale (Thomé, et al, 1997); km 5 on road
Tamazunchale-El Taiman (21º14’23” N, 98º49’44” W); (Correa-Sandoval, et al, 1998). TABASCO (Baker, 1925). TAMAULIPAS: various localities in SE part of
the state (Correa-Sandoval & Castro, 2000).
VERACRUZ: Cuatotolapam (Baker, 1925c); Las Tuxtlas Biological Station
(Naranjo-Garcia, et al, 2007); km.
post 234, road from Tuxpan-Poza Rica (20º49’11” N, 97º30’00”
W)(Correa-Sandoval, 2000). YUCATAN:
Aguada 14 km N and 2 km E of Mérida; 1.6 km N of Mérida (Bequaert & Clench,
1936); Mérida; X’makuil (Naranjo-Garcia, et
al, 2007).
Genus Phyllocaulis Colosi, 1922
Phyllocaulis Colosi, 1922; Ann. Mus. Nac. Hist.
Nat. Buenos Aires, 31: 486.- Thomé,
1975; Ihringia, Zool., (48): 21-24.-
Thomé, 1976; Iheringia, Zool., 49: 67-90.
TYPE SPECIES.- By subsequent
designation: Phylocaulus borellini
Colosi, 1921 (H. B. Baker, 1925; Nautilus, 39: 15) (= Vaginulus soleiformis Orbigny, 1835).
DISTRIBUTION.- Brazil, Uruguay,
Argentina, Chile and Bolivia. Introduced
elsewhere.
TAXONOMY.- About twenty species
are recognized (Thomé, 1975, 1976). One
had been introduced into western Mexico, but it has not been found there since
it was reported in 1925.
Phyllocaulis gayi
(Fischer, 1871)
Vaginulus
gayi Fischer, 1871;
Arch. Mus. Hist. Nat., 7: 172.
Vaginulus (Phyllocolis) gayi
(Fischer). H. B. Baker, 1925; Nautilus,
39: 15.
Phyllocolis
gayi
(Fischer). Thomé, 1971; Iheringia,
Zool., (4): 27-29; pl. 1, figs. 4-6 (animal); text-figs. 1-3, 18 (reproductive
anatomy).- Naranjo-Garcia, Thomé &
Castillejo, 2007; Rev. Mex. Biod., 78: 44-45.
TYPE LOCALITY.- Valdivia,
Chile. Lectotype in the Museum
d’Historie Naturelle, Paris (Forcart, 1952).
DISTRIBUTION RECORDS.- Southern
Chile (Naranjo-Garcia, Thomé & Castillejo, 2007). SINALOA: Mazatlán (Hoffmann, 1925).
Genus Sarasinula Grimpe & Hoffmann, 1924
Sarasinula
Grimpe &
Hoffmann, 1924; Zool. Anz., 68:
177.- Thomé, 1975; Ihringia, Zool.,
(48): 25-27.
TYPE SPECIES.- By original
designation: Vaginulus plebeius
Fischer, 1868.
DISTRIBUTION.- Indo-Pacific
region. Neotropical region: Brazil,
Peru, Ecuador, Colombia, Venezuela, Lesser Antilles. Introduced elsewhere.
TAXONOMY.- Twelve species are
recorded from the neotropical region (Thomé, 1975). Additional species occur in the Indo-Pacific
region. Two species have been introduced
into Mexico and Central America.
Sarasinula dubia (Semper,
1885)
Vaginua
dubia Semper, 1885;
Landmollusken, 7: 296.- Thomé, 1972:
252-253.
Sarasinula
dubia (Semper).- Thomé, 1993; Biociências, 1: 71.- Thomé, Santos & Pedott, 1997; Proc. Biol.
Soc. Wash., 110: 530.- Naranjo-Garcia,
Thomé & Castillejo, 2007; Rev. Mex. Biod., 78: 46-47.
TYPE LOCALITY.- St. Thomas, Virgin
Islands. Lectotype ZMB 39057a
(Naranjo-Garcia, Thomé & Casillejo, 2007).
DISTRIBUTION.- Brazil, Venezuela,
Colombia, Honduras and México.
Introduced into Texas, Florida, the West Indies and some Pacific Islands
(Thomé, et al, 1997). Its natural distribution in Central America
and Mexico is questionable. COSTA RICA, Puntarenas Prov.: Golfito. HONDURAS: La Lima (Thomé, et al, 1997). EL SALVADOR, Dept. La Libertad: Santa
Telca. CHIAPAS: ca. 1 km E of Finca
Custepec, E side of Rio Custec; 5 km E of Cañada Teopisca. COLIMA: Callejones (18º47’53” N, 103º38’29”
W). JALISCO: El Grullo; Pueblo Careyes,
km 54 on Fed. Hwy. 200, Barra de Navidad–Puerto Vallara; Puerto Vallara. MORELOS: Tetecalita, 16.5 km S, 65. km E of
Cuernavaca; 25 km W of Huamantla.
OAXACA: 4 km NW of Tepantepec (16º23’54” N, 94º12’54” W). QUERETARO: Jalpan; Xilitla. SAN LUÍS POTOSÍ: Cascada de Tamasopo, 4 km E
of Tamasopo. SINALOA: Mazatlan. VERACRUZ: Cuitláhuac (all Mexican records
from Naranjo-Garcia, et al, 2007).
Sarasinula plebeia (Fischer,
1868)
Vaginulus
prebeius Fischer,
1868; Jour. de Conchyl., 16: 145.
Sarasinula
plebeia (Fischer).- Andrews & Dundee, 1987; Ceiba, 28:
163-172.- Thomé, 1993; Biociências, 1:
71.- Thomé, Santos & Pedott, 1997;
Proc. Biol. Soc. Wash., 110: 530-531.-
Naranjo-Garcia, Thomé & Catillejo, 2007; Rev. Mex. Biod., 78: 45.
TYPE LOCALITY.- Nouméa, New
Caledonia (Thomé, 1971).
DISTRIBUTION.- Native to Chile,
Brazil, Venezuela, Colombia, Costa Rica, El Salvador, Honduras, Guatemala and Mexico. Introduced into Florida, Texas, the West
Indies, Australia, Asia and Africa (Thomé, Santos & Pedott, 1997:
530-531). COSTA RICA, Puntarenas Prov.: Golfito. EL SALVADOR, Dept. La Libertad: Santa Telca.
HONDURAS: no specific locality.
NICARAGUA: no specific locality.
MEXICO, VERACRUZ: Los Tuxtlas Biological Station (Naranjo-Garcia, et al, 2007). YUCATAN: Merida (Andrews & Dundee, 1987).
Genus Veronicella Blainville, 1817
Veronicella Blainville, 1817; Journal de
Physique, de Chimie, et d'Histoire Naturelle..., 85 (12): 437-444.- H. B. Baker, 1925: 158.- Thomé, 1975; Ihringia, Zoologie, (48): 32-35.
TYPE SPECIES.- Veronicella laevis Blainville, 1817, by
monotypy.
DISTRIBUTION.- Jamaica,
Hispaniola, Cuba, Ecuador and Chile.
Introduced elsewhere.
TAXONOMY.- Seven species are
recognizes (Thomé, 1975, 1988).
Veronicella sloanei (Cuvier,
1816)
Onchdium
sloanei Cuvier,
1816; Reg. An., II: 411.
Veronicella
sloanei
(Cuvier).- H. B. Baker, 1925; Proc.
Acad. Nat. Sci. Phila., 77: 163-165; pl. 3, figs. 5, 6.- Thomé, 1988; Iheringia, Zool. (67): 13-35.-
Thomé, 1993; Biociências, 1: 70.- Thomé,
Santos & Pedott, 1997; Proc. Biol. Soc. Wash., 110: 532.
Leidyula
sloanei (Cuvier). Thomé, 1975; Ihringia, Zool., (48): 18.
TYPE LOCALITY.- Jamaica.
DISTRIBUTION.- Jamaica, Cayman Islands.
Introduced into Barbados, Bahama Islands, Bermuda, Colombia (Old
Providence Island), Nicaragua and Honduras (Thomé, Santos & Pedott, 1997:
532. HONDURAS: no specific locality
given. NICARAGUA: Machuca.
STYLOMMATOPHORA
Superfamily SUCCINEOIDEA
Genus Catinella Pease, 1870
TYPE SPECIES.- Catinella rubida Peace, 1870.
DISTRIBUTION.- Northern Europe,
North America,
TAXONOMY.- Three subgenera are
recognized. One occurs in the study
area.
Subgenus Mediappendix Pilsbry, 1948
Mediappendix Pilsbry, 1948; Land Moll. N. Amer.,
2: 843.
TYPE SPECIES.- Succcinea campestris vagans Pilsbry,
1900.
DISTRIBUTION.- North America,
Europe and temperate
TAXONOMY.- Nine species are
recognized. Two occur in
Catinella
(Mediappendix) avara (Say, 1824)
Succinea
avara Say, 1824;
Major Long’s Second Expedition (
Catinella
avara (Say). Burch, 1962: 67.- Hubricht, 1985: 16; map 134.
Succinea
vermeta Say, 1829;
New Harmony Disseminator, 2: 230.
Quickella
vermeta (Say).
Hubricht, 1958; Nautilus, 72: 60.
Catinella
vermeta (Say). Grimm, 1960; Nautilus, 74: 12.- Hubricht, 1985: 16.
Catinella
texana Hubricht,
1961; Nautilus, 75: 61.
TYPE LOCALITY.- Northwest
territory. Holotype ANSP 59542.
DISTRIBUTION RECORDS.- Widespread
across North America. CAMPECHE: Pantel
Aguada,nr. Champoton (Bequaert & Clench, 1936). MEXICO: San Juan Teotihuacan (H. B. Baker,
1930). YUCATAN: San Ignacio (H. B. Baker,
1930).
Catinella
(Mediappendix) rehderi Pilsbry, 1948
Quickella
rehderi Pilsbry,
1948; Land Mol. N. Amer., 2: 845-847; fig. 457: b, fig. 457A: a-l (shell); fig.
456: D (reproductive anatomy).
Catinella
rehderi (Pilsbry). Smith, Miller, Christensen & Roth, 1990;
Proc. Calif. Acad. Sci., 47: 128.
TYPE LOCALITY.- 5 miles west of
Davenport, Lincoln Co., Washington.
Holotype ANSP 147757a.
DISTRIBUTION.- Washington and Montana south to Baja California Sur. BAJA CALIFORNIA NORTE: Isla Coronados; Tecate Valley, nr. U. S. border; 6.8 km S of
San Misión, 150 m alt.; Punta Banda; stream bank W of Ensenada rd. ca. 15 km N
of San Tomás; 0.8 km N San Vicente, 90 m alt.; San Telmo Mesa; 40 km N of El
Rosario; Miller’s Landing; 14 km E of Miller’s Landing; 13 km S of Miller’s
Landing; mesa just S of Rancho Mezquital (Smith, et al, 1990). BAJA CALIFORNIA SUR: Bajia San Bartolomé; 19
km SW of San Miguel Comondú; Bajia San Pedrito, 6.3 km SE of Todos Santos; 0.3 km
SSE ofSan Jose del Cabo, on rd. to La Playa (Smith, et al, 1990).
Genus Oxyloma Westerlund, 1885
TYPE SPECIES.- Succinea duneri
Pfeiffer, 1865.
DISRIBUTION.- Northern continents
and South Africa.
TAXONOMY.- Three subgenera are
recognized. One occurs in Mexico.
Subgenus Neoxyloma Pilsbry, 1948
Neoxyloma Pilsbry, 1948; Land Moll. N. Amer.,
2: 775.
TYPE SPECIES.- Succinea effuse Pfeifer, 1853.
DISTRIBUTION.- North America.
TAXONOMY.- About a dozen species
are recognized. Two species and one
subspecies occur in Mexico.
Oxyloma
(Neoxyloma) nuttalliana Lea, 1841
Succinea
nuttalliana Lea,
1841; Proc. Amer. Philos. Soc., 2: 32.
Oxyloma
nuttalliana
(Lea).- Pilsbry, 1948; Land Moll. N.
Amer., 2: 794-795.- Smith, Miller, Christensen & Roth, 1990; Proc. Calif.
Acad. Sci., 47: 129.
TYPE LOCALITY.- Oregon.
DISTRIBUTION.- BAJA CALIFORNIA
SUR: 19 km SW of San Miguel Comandú (Smith, et al, 1990).
Oxyloma
(Neoxyloma) tlapamensis tlapamensis Pilsbry, 1899
Succinea
tlapamensis
Pilsbry, 1899; Proc. Acad. Nat. Sci. Phila., 51: 401.- Von Martens, 1901; Biol. Cent. Amer.:
647.- Pilsbry, 1904; Proc. Acad. Nat.
Sci. Phila., 55: 776; pl. 48, figs. 4, 4a (shell).
TYPE LOCALITY.- Near Tlapam, State of Mexico. Lectotype ANSP 77207a (H. B. Baker, 1963:
215.
DISTIBUTION RECORDS.- MEXICO:
known only from the type locality.
Oxyloma
(Neoxyloma) tlapamensis cuitseana Pilsbry, 1899
Succinea
tlapamensis cuitseana Pilsbry,
1899; Proc. Acad. Nat. Sci. Phila., 51: 401.-
Von Martens, 1901; Biol. Cent. Amer.: 647.- Pilsbry, 1904; Proc. Acad. Nat. Sci. Phila.,
55: 776
TYPE LOCALITY.- Lago de Cuitseo,
near Huango, Michoacán. Lectotype ANSP
77208a (H. B. Baker, 1963: 215.
DISTRIBUTION RECORDS.- MICHOACAN:
known only from the type locality.
Genus Succinea Draparnaud, 1801
Succinea Draparnaud, 1801; Tableau des
Mollques terrestres et fluviatiles de la France: 55.
TYPE SPECIES.- Helix putris Linnaeus, 1756.
DISTRIBUTION.- The northern
Hemisphere, North Africa, Australia and some Pacific islands.
TAXONOMY.- Five subgenera are
recognized. Subgeneric assignment is based on anatomical data, for
which the vast majority of the Mexican and Central American species are
lacking. Two species of known anatomy
are so assigned. The rest are left
provisionally in the subgenus Succinea.
Subgenus Calcisuccinea Pilsbry, 1948.
Calcisuccinea Pilsbry, 1948; Land Moll. N. Amer.,
2: 820.
TYPE SPECIES.- Succinea campestris Say, 1817.
DISTRIBUTION.- North America.
TAXONOMY.- About a dozen species
are recognized. Two species and three
subspecies that occur in Mexico are assigned to this subgenus.
Succinea
(Calcisuccinea) campestris Say, 1817
Succinea
campestris Say,
1817; Jour. Acad. Nat. Sci. Phila., 1: 281.-
Pilsbry, 1891; Proc. Acad. Nat. Sci. Phila., 43: 318.
Succinea (Calcisuccinea) campestris
Say. Pilsbry, 1948; Land Moll. N. Amer.,
2: 826-828; fig. 448 (shell); figs. 449a (reproductive anatomy).
TYPE LOCALITY.- Sea Islands of
Georgia and Cumberland Island; Amelia Island, N. E. Florida.
DISTRIBUTION RECORDS.- Coastal
areas of North Carolina, South Carolina, Georgia and Florida. MEXICO, Distrito Federal: lago de Texcoco
(Pilsbry, 1901).
REMARKS.- This species has not
been recorded from Mexico subsequent Pilsbry’s report in 1891.
Succinea
(Calcisuccinea) luteola luteola Gould, 1848
Succinea
luteola Gould,
1848; Boston Soc. Nat. Hist., 3: 37.- Fischer
& Crosse, 1878; Miss. Sci. Mex., 1:658; pl. 27, figs. 2, 2a, 2b
(shell).- Dall, 1896; Proc. U. S. Nat.
Mus., 19: 365.- Von Martens, 1898; Biol.
Centr. Amer.: 331.- Bequaert &
Clench, 1936; Pub. Canegie Inst. Wash., (457): 65.- Goodrich & van der Schalie, 1937: 31.- Pilsbry, 1948; Land Moll. N. Amer., 1:
828-830; fig 448: B (reproductive anatomy); figs. 450: a-g (shell).- Branson, McCoy & Sisk, 1964; Southwestern
Naturalist, 9: 104.- Correa-Sandoval,
Gutierrez & Reza, 1898: 14.- Correa-Sandoval,
2000; Acta Zool.Mex., (79):: 8.- Correa-Sandoval
& Castro, 2002; Acta Zool. Mex., (86): 236.- Correa-Sandoval & Rodriguez, 2005:
59.
Succinea
texasiana Pfeiffer,
1848; Mon. helic. Vivent., 2: 526.
TYPE LOCALITY.- Galveston, Texas
(Pilsbry, 1948: 830).
DISTRIBUTION RECORDS.- Widely
distributed from Louisiana west to Arizona and south through much of
Mexico. GUATEMALA (Goodrich & van er
Schalie, 1937). CHIHUAHUA: Lago de
Palomas (Dall, 1896). GUERRERO; Venta de
Zopilote (Von Martens, 1898). NUEVO
LEON: km 8 on road from Linares-San Roberto (24°48’54” N, 88°37’10” W); km 27
on road from Linares-Iturbide (24°44’43” N, 99°46’37” W); km 114, road from San
Roberto-Matehuala (24°32’42” N, 100°16’40’ W) (Correa-Sandoval & Rodriguez,
2005). SAN LUÍS POTOSÍ: numerous
localities (Correa-Sandoval, et al,
1998). SONORA: Rio Bavispa, 21 mi. S of
Agua Prieta (Branson, et al,
1964). TAMAULIPAS: numerous localities
in southern part of state (Correa-Sandoval & Castro, 2002; Acta Zool. Mex.,
(86): 236). VERACRUZ: Rancho El Sol,
Naranjos (21°20’00” N, 97°43’16” W); San Juan Cuajinampa (21°11’53” N,
97°30’00” W); El Bajío, carr. Naranjos- Tuxpan (20°57’17” N, 97°25’57” W);
carr. Tuxpan- Poza Rica, km 234 (1 km al E) (20°49’11” N, 97°30’00” W); El
Cedral, carr. Poza Rica-Tajín (20°29’11” N, 97°25’23” W); La Ordeña, Papantla
(20°29’43” N, 97°18’27” W); Ruinas El Tajín (20°26’29” N, 97°22’30” W); La
Guadalupe, carr. Papantla- Nautla, km 60 (20°22’42” N, 96°55’23” W)
(Correa-Sandoval, 1999). YUCATAN:
Progreso (Pilsbry, 1891); Cerro Isla; Cienega, nr. Progreso (Bequaert &
Clench, 1936).
Succinea
(Calcisuccinea) luteola rudiuscula Von Martens, 1898
Succinea
luteola rudiuscula Von
Martens, 1898; Biol. Cent. Amer.: 331, 334; pl. 19, fig. 3 (shell).
TYPE LOCALITY.- Tehuacán, Puebla.
DISTRIBUTION RECORDS.- PUEBLA:
known only from the type locality.
Succinea
(Calcisuccinea) luteola
sonoreneis Fischer & Crosse, 1878
Sucinea
lineata var. sonoreneis Fischer & Crosse, 1878;
Miss. Sci. Mex., 1:662; pl. 27, figs. 8, 8a, 8b (shell).
Succinea
luteola sonorensis
Fischer & Crosse. Bequaert &
Miller, 1973: 155-156.
TYPE LOCALITY.- Vicinity of the
Rio Yaqui, Sonora.
DISTRIBUTION RECORDS.- SONORA: 33
mi. W of Santa Ana; 1 mi. N of Santa Ana; 2 mi. N of Granados, on Rio Bavispe
(Bequaert & Miller, 1973).
Succinea
(Calcisuccinea) luteola subtilis Von Martens, 1898
Succinea
luteola subtilis Von
Martens, 1898; Biol. Centr. Amer.: 331; pl. 19, fig. 3 (shell).- Hinkley, 1907; Nautilus, 21: 78.
TYPE LOCALITY.- Vera Cruz.
DISTRIBUTION RECORDS.- SAN LUIS
POTOSI: Valles (Hinkley, 1907).
VERACRUZ: type locality.
Subgenus Succinea Draparnaud, 1801
TAXONOMY.- The following species
are left in this subgenus, pending further study of their anatomies. Twenty-two species and three subspecies occur
in the study area.
Sucinea
ampulacea Von Martens, 1898
Succinea
ampulacea Von
Martens, 1898; Biol. Centr. Amer.: 340; pl. 19, fig. 11 (shell).
TYPE LOCALITY.- Ameca, Jalisco.
DISTRIBUITON RECORDS.- JALISCO:
known only from the type locality.
Succinea
brevis Dunker, 1850
Succinea
brevis Dunker,
1850; in Pfeiffer, Malak. Blätt., 7:
84.- Fischer & Crosse, 1878; Miss.
Sci. Mex., 1:65; pl. 26, figs. 14, 14a, 14b (shell) Von Martens, 1898; Biol. Centr. Amer.: 339;
pl. 19. fig. 7 (shell).- Bequaert, 1957;
Bull. Mus. Comp. Zool., 116: 225.
TYPE LOCALITY.- Zimapan, Hidalgo.
DISTRIBUTION.- CHIAPAS: El Real,
600 m alt. (Bequaert, 1957). HIDALGO:
Zimapan.
Succinea
californica Crosse and Fischer, 1990
Succinea
californica Crosse
and Fischer, 1878; Jour. de Conchyl., 26: 63.-
Fischer & Crosse,1878: 663; pl. 27, figs 9, 9a, 9b (shell).- Pilsbry, 1948; Land Moll. N. Amer., 2:
841-842; fig. 457d (shell).- Smith, Miller, Christensen & Roth, 1990; Proc.
Calif. Acad. Sci., 47: 128.
TYPE LOCALITY.- San Tomás, Baja
California Norte.
DISTRIBUTION.- BAJA CALIFORNIA
NORTE: Santo Tomás; Santo Tomás Valley (Smith, et al, 1990).
Succinea
carmenensis Fischer & Crosse, 1878
Succinea
undulate var. carmenensis Fischer & Crosse, 1878;
Miss. Sci. Mex., 1:658, 659; pl. 27, figs. 6, 6a (shell).- Von Martens, 1898; Biol. Centr. Amer.: 341.
Succinea
carmenensis Fischer
& Crosse. Bequaert & Clench,
1933; Pub. Carnegie Inst. Wash., (431): 537.-
Richards, 1937; Proc. Amer. Phil. Soc., 77: 225, pl.4, fig. 5 (shell).-
Goodrich & van der Schalie, 1937: 31.-
Rehder, 1966; Proc. Biol. Soc. Wash., 79: 283.
TYPE LOCALITY.- Isla de Carmen,
Campeche.
DISTRIBUTION REORDS.- GUATEMALA, Dept. Petén (Goodrich & van der
Schalie, 1937). CAMPECHE: Isla de
Carmen. QUINTANA ROO: arid swamp a few
mi. N of San Miguel, Cozumel Island (Richards, 1937); Tulum (Rehder, 1966). YUCATAN: Santa Ana, nr. Calcehtok; Chichen
Itza (Bequaert & Clench, 1933).
Succinea
clarionensis Dall, 1926
Succinea
clarionensis Dall,
1926; Proc. Calif. Acad. Sci., (4) 15: 485-486; pl.35, fig. 2 (shell).
TYPE LOCALITY.- Clarion Island; 1040 ft. alt. Holotype CAS 2209.
DISTRIBUTION RECORDS.- NAYARIT:
known only from the type locality.
Succinea
colorata Fischer & Crosse, 1878
Succinea
colorata Fischer
& Crosse, 1878; Miss. Sci. Mex., 1:657; pl. 27, figs. 5, 5a, 5b
(shell).- Von Martens, 1898; Biol. Centr.
Amer.: 336.
TYPE LOCALITY.- Tabasco.
DISTRIBUTION RECORDS.- TABASCO:
San Juan Bautista (Von Martens, 1898).
Succinea
cordovana Fischer & Crosse, 1878
Succinea
undulate var. cordovana Fischer & Crosse, 1878;
Miss. Sci. Mex., 1:658; pl. 27, figs. 7, 7a.
Succinea
salleana var. cordovana Von Martens, 1898
TYPE LOCALITY.- Cordoba,
Veracruz.
DISTRIBUTION RECORDS.- VERACRUZ:
known only from the type locality.
Succinea
costaricensis Von Martens, 1898
Succinea
costaricensis Von
Martens, 1898; Biol. Centr. Amer.: 338; pl. 19, fig. 6 (shell).- Villalobos, C., J. Monge-Najera, Z.
Barrientos & J. Franco, 1995; Rev. Boil. Trop., 43: 181-186.
TYPE LOCALITY.- Not specified.
DISTRIBUTION RECORDS.- COSTA
RICA, Prov. San José: San José and at
La Palma, 1161-1600 m alt. (Von Martens, 1898).
Prov. Puntarenas: Savana de Guacimo, in the valley of the Rio Brus;
thermal springs of Djiri Durunia, valley of the Rio Diquis, 900 m alt. (Von
Martens, 1898).
Succinea
globispira Von Martens, 1898
Succinea
globispira Von
Martens, 1898; Sitzungs-Berichte der Gesllschaft naturforschenden Freude zu
Berlin: 158.- Von Martens, 1901; Biol.
Central Amer.: 641; pl. 44, fig. 12 (shell).-
Dall, 1900; Proc. Acad. Nat. Si. Phila., 52: 97.
TYPE LOCLAITY.- Cocos Island.
DISTRIBUTION RECORDS.- COSTA
RICA: Cocos Island (von Martens, 1901).
Succinea
guadelupensis Dall, 1900
Succinea
(rusticana Gould var.) guadelupensis Dall, 1900: 102-103; pl.
8, fig. 12 (shell).
Succinea
guadelupensis Dall.-
Pilsbry, 1927; Proc. Calif. Acad. Sci. (4), 16: 171-172.- Pilsbry, 1948; Land Moll. N. Amer., 2: 840;
fig. 455: I (shell).- Smith, Miller, Christensen
& Roth, 1990; Proc. Calif. Acad. Sci., 47: 128.
TYPE LOCALITY.- Guadalupe Island, Baja California Norte. Holotype in the USNM.
DISTRIBUTION.- BAJA CALIFORNIA
NORTE: Known only from Guadalupe Island (Smith, et al, 1990).
Succinea
guatemalensis Morelet, 1849
Succinea
guatemalensis Morelet,
1849; Test Noviss., I: 16.- Fischer
& Crosse, 1878: 667; pl. 26, figs.
11,11a (shell).- Von Martens, 1898;
Biol. Centr. Amer.: 337; pl. 19, fig. 9 (shell).- Pilsbry, 1920; Proc. Acad. Nat. Sci. Phila.,
72: 6.- Pérez & López, 2002:
111-112, fig. (map).
TYPE LOCALITY.- On banks of small streams, Cd. Guatemala, Guatemala.
DISTRIBUTION RECORDS.- COSTA
RICA, Prov. Cartago: Turrialba (Von
Martens, 1898); Juan Viñas (Pilsbry, 1920).
Prov. Puntarenas: Turubares;
Rio de las Cascadas, at the foot of the Great Fall, valley of the Rio Diquis (Von
Martens, 1898). NICARAGUA: Pacific
versant, common (Pérez & López, 2002).
GUATEMALA, Dept. Alta Verapaz:
Panzos (Von Martens, 1898). Dept.
Sacatepequez: Antigua (Von Martens, 1898).
Succinea
haustrellum Rehder, 1942
Succinea
haustrellum Rehder,
1942; Jour. Wash. Acad. Sci., 33: 350; fig. 19 (shell).
TYPE LOCALITY.- Pedernal,
Guanacaste Prov., Costa Rica; 200 m. alt.
Holotype USNM 536013.
DISTRIBTION RECORDS.- COSTA RICA.
Prov. San José.: San José (Rehder,
1942).
Succinea
hortulana Morelet, 1851
Succinea
hortulana Morelet,
1851; Test. Noviss., II: 14.- Fischer
& Crosse, 1878; Miss. Sci. Mex., 1:668; pl. 26, figs. 12, 12a
(shell).- Von Martens, 1898; Biol.
Centr. Amer.: 334-335.
TYPE LOCALITY.- Adhering to wall
in gardens, Suburbs of Cd. Guatemala, Guatemala.
DISTRIBUTION RECORDS.- GUATEMALA,
Dept. Sacatepequez: Antigua.
Succinea
lutosa Pilsbry, 1926
Succinea
lutosa Pilsbry,
1926; Proc. Acad. Nat. Sci. Phila., 78: 102-103; fig. 26 (shell).
TYPE LOCALITY.- Alhajuela, Canal
Zone, Panama. Holotype ANSP 141010a (H.
B. Baker, 1963: 215).
DISTRIBUTION RECORDS.- PANAMA,
Taboga Island; Prov. Panama: Panama
City (Pilsbry, 1926).
Succinea
mcgregori Pilsbry, 1898
Succinea
mcgregori Pilsbry,
1898; Proc. Acad. Nat. Sci. Phila., 50: 354; text-fig. (shell).- H. B. Baker, 1963; Proc. Acad. Nat. Sci.
Phila., 115: 215.
TYPE LOCALITY.- Clarion Island,
Revillagegido Islands, Nayarit.
Lectotype 72770a (H. B. Baker, 1963: 215).
DISTRIBUTION RECORDS.- NAYARIT:
known only from the type locality/
Succinea
panamensis Pilsbry, 1920
Succinea
panamensis Pilsbry,
1920; Proc. Acad. Nat. Sci. Phila., 71:
218; pl. 11, fig. 4 (shell).- Pilsbry,
1926; Proc. Acad. Nat. Sci. Phila., 78: 103; text-fig. 27 (shell).- Pilsbry, 1930; Proc. Acad. Nat. Sci. Phila.,
82: 351.
TYPE LOCALITY.- Las Cascades,
Canal Zone, Panama. Holotype ANSP 48522a
(H. B. Baker, 1963: 215)
DISTRIBUTION RECORDS.- PAMAMA, Canal Zone: Las Cascades (Pilsbry,
1926).
Succinea
panucoensis Pilsbry, 1910
Succinea
panucoensis
Pilsbry, 1910; Proc. Acad. Nat. Sci. Phila., 61: 546; text-fig. 6 (shell).
- H. B. Baker, 1963; Proc. Acad. Nat.
Sci. Phila., 115: 215.
TYPE LOCALITY.- “Alligator” Lake, San Luis Potosi. Lectotype ANSP 99504a (H. B. Baker, 1963:
215).
DISTRIBUTION RECORDS.- SAN LUIS
POTOSI: Pujal; Lago de Catamas (Pilsbry, 1910).
TAMAULIPAS: Tampico (Pilsbry, 1910).
Succinea
pueblensis Fischer & Crosse, 1878
Succinea
pueblensis Fischer
& Crosse, 1878; Miss. Sci. Mex.; 669; pl. 26, figs. 15-15c (shell).
TYPE LOCALITY.- Neighborhood of
Puebla [City], Puebla.
DISTRIBUTION RECORDS.- PUEBLA:
known only from the type locality.
Succinea
recisa Morelet, 1851
Succinea
recisa Morelet,
1851; Test. Noviss., II: 14.- Fischer
& Crosse, 654; pl. 26, figs. 13, 13a.-
Tate, 1859; Amer. Jour. Conch., 5: 158.-
Von Martens, 1898; Biol. Centr. Amer.: 339-340.- Pilsbry, 1910; Proc. Acad. Nat. Sci. Phila.,
62: 503.- Pilsbry, 1920; Proc. Acad.
Nat. Sci. Phila., 72: 6.- Richards,
1938; Proc. Amer. Philo. Soc., 79: 173.-
Pérez & López, 2002: 113-114, map.
TYPE LOCALITY.- Under bark of
trees, around Lago de Izabal, Dept. Izabal, Guatemala.
DISTRIBTION RECORDS.- PANAMA
(Tate, 1859). Canal Zone: Las Cascades (Pilsbry, 1910). COSTA RICA, Prov. Cartago: Juan Viñas (Pilsbry, 1920). NICARAGUA, Pacific versant, common (Pérez
& López, 2002). Dept. Matagalpa: Matagalpa (Von Martens). Dept.
Rio San Juan: near the Rio San Juan (Von
Martens, 1898). Region Autonoma Atlantico
Sur: Corn Island (Richards, 1939). HONDURAS, Roatan Island: between Coxen Hole and French Harbor; West End
(Richards, 1938). GUATEMALA, Dept. Sacatepequez: Antigua (Von Martens,
1898).
Succinea
rusticana Gould, 1846
Succinea
rusticana Gould,
1846; Proc. Boston Soc. Nat. Hist., 2: 187.-
Pilsbry, Land Moll. N. Amer., 2: 824-825; fig. 446a-d (shell).- Smith,
Miller, Christensen & Roth, 1990; Proc. Calif. Acad. Sci., 47: 128.
TYPE LOCALITY.- Oregon.
DISTRIBUTION.- Pacific coast
region from Alaska south to Baja California Norte (Roth & Sadeghian,
2006). BAJA CALIFORNIA NORTE: Ojos
Negros, in a slow stream through meadows (Smith, et al, 1990). BAJA CALIFORNIA SUR: Sierra Laguna (Smith, et
al, 1990).
Succinea
socorroensis Dall, 1926
Succinea
socorroensis Dall,
1926; Proc. Calif. Acad. Sci., (4) 15: 486; pl. 35, fig. 1 (shell)
TYPE LOCALITY.- On the north
slope of Mount Evermann, Socorro Island; 2800 ft. alt. Holotype CAS 2210.
DISTRIBUTION
RECORDS.- NAYARIT: known only from the
type locality.
Succinea undulate
undulata Say, 1829
Succinea
undulata Say, 1829;
New Harmony Disseminator of Useful Knowledge, 2: 230.- Fischer & Crosse, 1878; Miss. Sci. Mex.,
1:656; pl. 27, figs. 4, 4a, 4b (shell).-
Von Martens, 1898; Biol. Centr. Amer.: 335-336.
TYPE LOCALITY.- Mexico.
DISTRIBUTION RECORDS.- GUANAJUATO:
Irapuato (Von Martens, 1898). JALISCO: Sayula (Von Martens, 1898). MEXICO.
Succinea undulata
morchi Dunker, 1889
Succinea
undulata morchi
Dunker, 1889, in Paetel, Catalog
Conchyl. Sammlung., 2: 366.- Von Martens,
1898; Biol. Centr. Amer.: 336; pl. 19, fig. 8 (shell).
TYPE LOCALITY.- Zimapan, Hidalgo.
DISTRIBUTION RECORDS.- HIDALGO:
known only from the type locality.
Succinea
virgata virgata Von Martens, 1865
Succinea
virgata Von Martens,
1865; Malak. Blätter, 12: 50; pl. 1, figs. 6, 7 (shell).- Von Martens, 1898; Biol. Centr. Amer.: 334;
pl. 19, fig. 4 (shell).- Pilsbry, 1904;
Proc. Acad. Nat. Sci. Phila., 55: 776.
TYPE LOCALITY.- Veracruz.
DISTRIBUTION RECORDS.-
OAXACA. PUEBLA: Paso de Macho;
Tehuacan (Von Martens, 1898). VERACRUZ: Sierra de Atoyac (Von Martens, 1898);
Texolo (Pilsbry, 1904).
Succinea virgata
hogeana Von Martens, 1898
Succcinea
virgata hogeana Von
Martens, 1898; Biol. Centr. Amer.: 334; pl. 19, fig. 5 (shell).
TYPE LOCALITY.- Oaxaca.
DISTRIBUTION RECORDS.- OAXACA:
known only from the type locality.
Succinea virgata
microspira Von Martens, 1898
Succinea
virgata var. β
Fischer & Crosse,1878; Miss. Sci. Mex., 1: 659; pl. 27, figs. 3, 3a, 3b
(shell).
Succinea
virgata microspira
Fischer & Crosse, Von Martens, 1898; Biol. Centr. Amer.: 334.- H. B. Baker, 1930; Occ. Pap. Mus. Zool. Univ.
Mich., (220): 5.
TYPE LOCALITY.- Puebla.
DISTRIBUTION RECORDS.- PUEBLA:
Necaxa (H. B. Baker, 1930).
Superfamily ACHATINELLOIDEA
Family
TORNATELLINIDAE Cooke and Kondo, 1961
Genus Tornatellinides Pilsbry, 1910
Tornatellinides Pilsbry, 1910; Nautilus, 23: 123.
TYPE SPECIES.- Tornatellina simplex Pease.
DISTRIBUTION.- Japan,
TAXONOMY.- Two subgenera are
recognized. The typical subgenus occurs
on the
Tornatellides
mexicana Dall, 1926
Tornatellides
mexicana Dall, 1926;
Proc.
TYPE LOCALITY.- On the south
slope of Mt. Evermann, Socorro Island, Revillogigedo Islands, Nayarit;
2000-2800 ft. alt. Holotype CAS 2207.
DISTRIBUTION RECORDS.- NAYARIT: known only from the type locality on
Tornatellides
clarionensis Dall, 1926
Tornatellides
clarionensis Dall,
1926; Proc. Calif. Acad. Sci., (4) 15: 485; pl. 35, fig. 9 (shell).
TYPE LOCALITY.- Clarion Island,
Revillogigedo Islands, Nayarit; 1040 ft. alt.
Holotype CAS 2209.
DISTRIBUTION RECORDS.- NAYARIT:
known only from the type locality on Clarion Island.
Superfamily COCHLICOPOIDEA
Family
COCHLICOPIDAE Pilbry, 1900
Genus Cochlicopa Férussac, 1821
Cochlicopa Férussac, 1821; Tableaux
Systematiques des Animaux Mollusques: 24.
Cionella Jeffreys, 1829; Trans. Linn. Soc.
London, 16: 347. Pilsbry, 1948; Land
Moll. N. Amer., 2: 1045-1047.
TYPE SPECIES.- Helix lubrica
Müller, 1774.
DISTRIBUTION.- Holarctic in
occurrence.
TAXONOMY.- Four species are
recognized in
Cochlicopa
lubrica (Müller, 1774)
Helix
lubrica Müller,
1774; Verm. Hist.: 104.
Cochlicopa
lubrica
(Müller). Pilsbry & Johnson, 1898; Nautilus, 11: 127.- Pilsbry, 1908; Man. Conch., 19: 312.- Hubricht, 1985: 6; map 24.
Cionella
lubrica (Müller). Pilsbry, 1948; Land Moll. N. Amer., 2:
1047-1049; figs. 560a, 560b (shell).-
Pilsbry, 1953; Proc. Acad. Nat. Sci. Phila., 105: 165.
TYPE LOCALITY.- Denmark.
DISTRIBUTION RECORDS.- Holarctic
realm; widespread in North America (Hubricht, 1985). CHIHUAHUA: about 8-9 km W of Colonia Juarez;
talus of cliff on the Rio Piedras Verdes, about 3.6 km below Pacheco, 5900 ft.
alt.; Sierra de la Breña, 17.5 km from Pearson, on road to Pacheco, 7000 ft.
alt. (Pilsbry, 1953). NUEVO LEON: near
the Hacienda Pablillo (Pilsbry, 1953)
Superfamily PUPILLOIDEA
Family
STROBILOPSIDAE Wenz, 1915
Genus Strobilops Pilsbry, 1893
Strobila Morse, 1856; Jour. Portland Soc.
Nat. Hist., 1: 26 (not Strobila Sars,
1835; not Strobila Sodoffsky, 1837).
Strobilops Pilsbry, 1893; Proc. Acad. Nat.
Sci. Phila., 44: 403 (substitute name for Strobila
Morse).- Pilsbry, 1927; Man. Conch., 28:
12, 56-58.- Zilch, 1959: 177.
TYPE SPECIES.- Helix labyrinthica Say, 1817.
DISTRIBUTION.- North
TAXONOMY.- Five subgenera are
recognized. Two subgenera, Strobilops s. s., and Discostrobilops, occur in the study
area. A third subgenus from the Cayman
Islands, Coelostrobilops, is included
in this list because of its geographic proximity to
Subgenus Strobilops Pilsbry, 1893
DISTRIBUTION.- North America,
TAXONOMY.- Twelve species are
recognized. Seven species occur in the
study area.
Strobilops aenea
mexicana Pilsbry, 1927
Strobilops
aenea mexicana
Pilsbry, 1927; Man. Conch., 28; 32-33; pl. 5, figs. 9-13 (shell).- Pilsbry, 1953; Proc. Acad. Nat. Sci. Phila.,
105: 165.- Correa-Sandoval, Gutierrez
& Reza, 1998; Acta Zool. Mex., (73): 14.
TYPE LOCALITY.- Diente, near
Monterey, Nuevo Leon. Holotype ANSP
77194a.
DISTRIBUTION RECORDS.- NUEVO
LEON: Diente; Monterey; along the Rio Maurisco, ca. 25 km S of Monterey
(Pilsbry, 1953). PUEBLA: Necaxa
(Pilsbry, 1927). SAN LUÍS POTOSÍ: 6 km
NW of Xilitla; 830 m alt. (21°23’10” N, 99°03’56” W); Las Pozas; 580 m alt.
(21°24’23” N, 99°00’00” W); 0.3 km NW of Las Pozas; 600 m alt. (21°24’38” N,
99°00’15” W) (Correa-Sandoval, et al,
1998).
Strobilops californica
Miller & Christensen, 1980
Strobilops
californica Miller
& Christensen, 1980; Proc. Biol. Soc. Wash., 93: 593-596; figs. 1a-c
(shell).- Smith, Miller, Christensen
& Roth, 1990; Proc. Calif. Acad. Sci., 47: 111; fig. 15 (map).
TYPE LOCALITY.- La Laguna, Sierra
de la Victoria, along slope above creek immediately behind La Laguna shelter,
Baja California Sur; 6500 ft. alt.
Holotype USNM 799595.
DISTRIBUTION RECORDS.- BAJA
CALIFORNIA SUR: known only from the type locality.
Strobilops hannai
Pilsbry, 1931
Strobilops
hannai Pilsbry,
1931; Man. Conch., 28: 59-60; pl. 16, figs. 3, 3a, 5 (shell).
TYPE LOCALITY.- Socorro Island,
Revillogegido Islands, Nayarit. Holotype
ANSP 256587a.
DISTRIBUTION RECORDS.- NAYARIT:
known only from the type locality.
Strobilops piratica
Pilsbry, 1930
Strobilops
piratica Pilsbry,
1930; Proc. Acad. Nat. Sci. Phila., 82; 257; pl. 19, figs. 9-11 (shell).- Pilsbry, 1931; Man. Conch., 28: 58-59; pl.
13, figs. 8, 9, 10, 11 (shell).
TYPE LOCALITY.- On the summit of
the ridge north of High Peak, Old Providence Island, Colombia. Holotype ANSP 150860.
DISTRIBUTION RECORDS.- OLD
PROVIDENCE ISLAND, COLOMBIA: known only from the type locality.
Strobilops salvini
(Tristram, 1863)
Helix
salvini Thristram,
1863; Proc. Zool. Soc. London: 411.
Strobila
salvini
(Tristram). Von Martens, 1892; Biol.
Cent. Amer.: 174; pl. 10, figs. 1-1c (shell).
Strobilops
salvini
(Tristram). Pilsbry, 1927; Man. Conch.,
28: 37-39; pl. 4, figs. 1 (shell).
TYPE LOCALITY.- Mountain forests
of northern Guatemala, [Dept. Alta Verapaz].
DISTRIBUTION RECORDS.- GUATEMALA:
known only from the type locality.
Strobilops strebeli
strebeli (Pfeiffer, 1862)
Helix
strebeli Pfeiffer,
1862; Malak. Blätter, 8: 31; pl. 1, figs. 5-8 (shell).
Strobilops
strebeli
(Pfeiffer). Pilsbry, 1927; Man. Conch., 28: 33-34; pl. 5, figs. 1-4 (shell).
Strobila
labyrinthica (Say).
Strebel, 1880: 43; pl. 4, fig. 6 (shell).-
Von Martens, 1892; Biol. Cent. Amer.: 173.
TYPE LOCALITY.- Mirador,
Veracruz.
DISTRIBUTION RECORDS.- VERACRUZ:
Mirador; environs of Veracruz (Strebel, 1880).
Strobilops strebeli
guatemalensis Hinkley, 1920
Strobilops
strebeli guatemalensis
Hinkley, 1920; Nautilus, 34: 52.-
Pilsbry, 1927; Man. Conch., 28: 34-35, 60; pl. 5, figs. 5, 6, 7, 8
(shell).
TYPE LOCALITY.- In beach debris
of Lago de Izabal, Jocolo, Dept. Izabal, Guatemala. Holotype ANSP 9272a.
DISTRIBUTION RECORDS.- GUATEMALA,
Dept. Izabal: known only from the
type locality.
Strobilops
veracruzensis veracruzensis
Pilsbry, 1927
Strobila
labyrinthica
Strebel, 1880: 44; pl. 4, fig. 6 (shell) (not Helix labyrinthica Say, 1817).
Strobilops
veracruzensis
Pilsbry, 1927; Man. Conch., 28: 35-36; pl. 4, fig. 6 (shell); pl. 9, figs. 1,
2, 3 (shell) (new name for Helix
labyrinthica Strebel).
TYPE LOCALITY.- Neighborhood of
Veracruz.
DISTRIBUTION RECORDS.- VERACRUZ:
known only from the type locality.
Strobilops
veracruzensis crossei Pilsbry, 1927
Helix
strebeli Pfeiffer. Fischer & Crosse, 1872: 267; pl. 12,
figs. 7a. 7b (shell) (not Helix strebeli
Pfeiffer, 1862).
Strobilops
veracruzensis crossei
Pilsbry, 1927; Man. Conch., 28: 36-37; pl. 9, figs. 4, 4a (shell) (new name for
Helix strebeli Fischer & Crosse).
TYPE LOCALITY.- Mirador,
Veracruz.
DISTRIBUTION RECORDS.- VERACRUZ:
known only from the type locality.
Subgenus Coelostrobilops Pilsbry, 1931
Coelostrobilops Pilsbry, 1931; Man. Conch., 28; 60.
TYPE SPECIES.- Strobilops wenziana Pilsbry, 1930.
DISTRIBUTION.- Grand Cayman
Island.
TAXONOMY.- A single species is
recognized.
Strobilops wenziana
Pilsbry, 1930
Strobilops
wenziana Pilsbry,
1930; Proc. Acad. Nat. Sci. Phila., 82: 238; pl. 19, figs. 1-7 (shell).- Pilsbry, 1931; Man. Conch., 28: 60-62; pl.
13, figs. 1-7 (shell).
TYPE LOCALITY.- About midway
between North Sound and Red Bay, Grand Cayman Island. Holotype ANSP 150861.
DISTRIBUTION RECORDS.- GRAND
CAYMAN ISLAND: known only from the type locality.
Subgenus Discostrobilops Pilsbry, 1927
Discostrobilops Pilsbry, 1927; Man. Conch., 28: 18,
46.- Pilsbry, 1948; Land Moll. N. Amer.,
2: 865.- Morrison, 1953; Nautilus, 65:
55.
TYPE SPECIES.- Helix hubbardi A. D. Brown, 1861.
DISTRIBUTION.- Gulf coastal
plains from Mexico northeast to Georgia and Florida, Bermuda, Cuba, Jamaica and
the Bahamas Islands; Northwest Mexico, Sonora (?).
TAXONOMY.- Three extant species
are recognized. Two occur in Mexico.
Strobilops (Discostrobilops) hubbardi (A. D. Brown, 1861)
Helix
hubbardi A. D.
Brown, 1861; Proc. Acad. Nat. Sci. Phila.: 333; text-fig. (shell).
Strobilops (Discostrobilops) hubbardi
(Brown). Pilsbry, 1927; Man. Conch., 28: 47-48; pl. 7, figs. 1, 2, 3
(shell).- Pilsbry, 1948; Land Moll. N.
Amer., 2: 865-868; figs. 1-9 (shell).-
Correa-Sandoval, Gutierrez & Reza, 1998: 14.- Correa-Sandoval & Castro, 2002; Acta
Zool. Mex., (86): 236.
Helix vendryesiana Gloyne, 1872;
Jour. de Conchyl., 19: 333.
Strobilops hubbardi (Gloyne). Gulick, 1904; Proc. Acad. Nat. Sci. Phila.,
56: 413.- Hinkley, 1907; Nautilus, 21:
76.- Pilsbry, 1948; Land Mollusca of
North America, 2: 865-868; figs. 468, 1-9 (shell).
Strobilops
hubbardi vendryesiana
(Gloyne). Pilsbry, 1931; Man. Conch.,
28: 48-50; 7, figs. 4-12 (shell); pl. 8, figs. 1-9 (shell).
Strobilops
hubbardi stevensoni
Pilsbry, 1899; Proc. Acad. Nat. Sci. Phila., 51: 404.
TYPE LOCALITIES.- Helix hubbardi: Indianola, Calhoun Co.,
Texas; holotype ANSP 124. Helix vendryesiana: Jamaica. Strobilops
hubbardi stevensoni: mangrove swamp, Biscayne Bay near State Agricultural
Station, Miami, Florida.
DISTRIBUTION RECORDS.- The Gulf
coastal plains from northeastern Mexico to Florida and Georgia; Jamaica, Cuba,
Bimini Island and Bermuda. SAN LUÍS
POTOSÍ: Valles (Pilsbry, 1931); km 48 on rd. from Cd. Valles to Agua Buena; 900
m alt. (21°52’55” N, 99°22’06” W);
Cascadas de Tamasopo; 430 m alt. (21°56’05” N, 99°25’00” W); Las
Abritas, 5 km E of El Naranjo; 840 m alt. (22°33’24” N, 99°22’22” W); 6 km. SW
of Xilitla; 830 m alt. (21°23’10” N, 99°03’56” W); Las Cascadas, Tamasopo
(21°56’05” N, 99°25’00” W) (Correa-Sandoval, et al, 1998). TAMAULIPAS:
river drift, Tampico (Hinkley, 1907); nr. Gomez Farias (23°00’00” N, 99°09’18”
W); Salto El Tigre, Santa Engracia (23°58’51” N, 99°16’53” W) (Correa-Sandoval
& Castro, 2002).
Strobilops
(Discostrobilops) sinaloa Morrison, 1953
Strobilops (Discostrobilops) sinaloa,
Morrison, 1953; Nautilus, 67; 54-55; pl. 2, figs. 4, 5, 6 (shell).- Miller & Christensen, 1980; Proc. Biol.
Soc. Wash., 93: 595.
TYPE LOCALITY.- Sinaloa ?. Holotype USNM 592719.
DISTRIBUTION RECORDS.- SINALOA:
known only from the type locality. It is
questionable about whether this species actually came from Sinaloa (Morrison,
1953).
Family VALLONIIDAE
Morse, 1864
Genus Vallonia Risso, 1820
Vallonia Risso, 1820; Hist. Nat. Eur.
Merid., 4: 101.- Pilsbry, 1948; Land
Moll. N. Amer., 2: 1019-1023.
TYPE SPECIES.- Vallonia rosalia Risso, 1826 (= Helix costata Müller, 1774).
DISTRIBUTION.- North America,
Europe and North Africa, northern and central Asia, Japan.
TAXONOMY.- Twenty-five species
are recognized (Pilsbry, 1948). One
occurs naturally in Mexico.
Vallonia
perspeciva Sterki, 1892
Vallonia
perspeciva Sterki,
1892; Nautilus, 6: 77.- Sterki, 1893; in Pilsbry, Man. Conch, 8: 257; pl. 33,
figs. 39-45 (shell).- Pilsbry, 1948; Land
Moll. N. Amer., 2: 1033-1034; fig. 553 (shell).- Pilsbry, 1953; Proc. Acad. Nat. Sci. Phila.,
105: 165.
TYPE LOCALITY.- Woodville,
Jackson Co., Alabama.
DISTRIBUTION RECORDS.- Widespread
across temperate North America, barely entering northern Mexico. CHIHUAHUA: Sierra de la Breña, 11 mi. from
Pearson, on rd. to Pacheco, 7000 ft. alt.; in cliff talus along the Rio Piedras
Verdes, 8-9 km W of Colonia Juarez; on the Rio Piedras Verdes, 3¼ km E of
Pacheco (Pilsbry, 1953).
Family VERTIGINIDAE
Stimpson, 1851
Fifty (51) species and nine
subspecies occur in the study area.
Subfamily TRUNCATELLININAE
Steenberg, 1925
Genus Columella Westerlund, 1878
Columella Westerlund, 1878; Fauna Europaea …,
2: 193.- Pilsbry, 1926; Man. Conch., 27:
232—233.
Sphaeradium Westerlund, 1887; Fauna
Palearctischen Region Binnenconchylien, 3: 125 (not Sphaeradium as limited by Von Martens, 1860).
Edentulina Clessin, 1876 (not Edentulina Pfeiffer, 1855).
TYPE SPECIES.- Pupa edentula Draparnaud, 1805.
DISTRIBUTION.- The Palearctic
region; North America as far south as Nicaragua, Hawaii.
TAXONOMY.- About ten species are
recognized. Four occur in North America
and Central America. One is known from
the study area.
Columella
polvonense (Pilsbry, 1894)
Pupa
polvonense Pilsbry,
1894; Proc. Acad. Nat. Sci. Phila., 46: 31; pl. 1, fig. 11.
Sphaeradium
polvonense Pilsbry,
1904; Proc. Acad. Nat. Sci. Phila., 55: 769.
Columella
polvonensis
(Pilsbry). Pilsbry, 1926; Man. Conch.,
27: 246; pl. 31, fig. 9 (shell).
TYPE LOCALITY.- Polvón, Dept.
DISTRIBUTION
Subfamily PUPILLINAE
Turton, 1831
Genus Chaenaxis Pilsbry & Ferriss, 1906
Chaenaxis Pilsbry & Ferriss, 1906; Proc.
Acad. Nat. Sci. Phila., 58: 145.-
Pilsbry, 1916;
TYPE SPECIES.- Bifidaria tuba Pilsbry & Ferriss,
1906.
DISTRIBUTION.- Arizona,
TAXONOMY.- A single species is
recognized.
Chaenaxis
tuba Pilsbry & Ferriss, 1906
Bifidaria
tuba Pilsbry &
Ferriss, 1906; Proc. Acad. Nat. Sci. Phila., 58: 145; fig. 6 (shell).
Chaenaxis
tuba (Pilsbry &
Ferriss). Pilsbry, 1916; Man. Conch.,
24: 2; pl. 9, figs. 1, 2, 3 (shell).-
Pilsbry, 1948; Land Moll. N. Amer., 2: 917-919; figs. 496; figs. 497: 1,
2, 3 (shell).- Branson, McCoy &
Sisk, 1964; Southwestern Naturalist, 9: 104.-
Bequaert & Miller, 1973: 172-176.-
Naranjo-Garcia, 1991: 168.
Bifidaria
tuba intuscostata Clapp, 1908; Nautilus, 22:
76, 96; pl. 7, figs. 1-11 (shell).
Chaenaxis
intuscostata
(Clapp). Pilsbry, 1916; Man. Conch., 24:
3; pl. 9, figs. 6-9 (shell).
Chaenaxis
intuscostata form brevicostata Pilsbry, 1916; Man.
Conch., 24: 4; pl. 6, figs. 4, 5 (shell).
Chaenaxis
sonorensis Pilsbry,
1953; Proc. Acad, Nat. Sci. Phila., 105: 163-164; pl. 9, figs. 8, 8a (shell).
TYPE LOCALITY.- Bifidaria tuba: drift debris of the
DISTRIBUTION
Genus Pupilla Leach, 1828
Pupilla Leach, in Fleming, 1828; British Animals: 268.- Pilsbry, 1921;
TYPE SPECIES.- Pupa marginata Draparnaud, 1801 (= Pupa muscorum Linnaeus, 1758).
DISTRIBUTION.- Temperate North
America, Europe, Africa, Asia and
TAXONOMY.- Five subgenera are
recognized. Two occur in the study area.
Subgenus Pupilla Leach, 1828
DISTRIBUTION.- Temperate North
America, Europe, Africa, Asia and
TAXONOMY.- About thirty species
are recognized. One species occurs in
the study area.
Pupilla
(Pupilla) hebes (Ancey, 1881)
Pupa
hebes Ancey, 1881;
Le naturaliste, 3: 389.
Pupilla
hebes (Ancey). Pilsbry & Ferriss, 1911; Proc. Acad. Nat.
Sci. Phila., 63: 197.
Pupilla (Pupilla) hebes
(Ancey). Pilsbry, 1921; Man. Conch., 26:
164-167; pl. 18., figs. 1-4 (shell).-
Pilsbry, 1948; Land Moll. N. Amer., 2: 936-939; figs. 503: 1-4
(shell).- Pilsbry, 1953; Proc. Acad.
Nat. Sci. Phila., 105: 164.- Bequaert
& Miller, 1973: 180-181.- Smith,
Miller, Christensen & Roth, 1990; Proc. Calif. Acad. Sci., 47: 107.
Pupa
arizonensis Gabb, in Binney, 1885; Man. Amer. land shells:
173.
Pupilla
muscorum idahoensis
Henderson & Daniels, 1917; Proc. Acad. Nat. Sci. Phila., 69: 57.
Pipilla
hebes nefas Pilsbry
& Ferriss, 1910; Proc. Acad. Nat. Sci. Phila., 62: 135.
TYPE LOCALTIY.- Pupa hebes: White Pine, White Pine Co.,
DISTRIBUTION RECORDS.-
Washington, Idaho and
Subgenus Striopupilla Pilsbry, 1921
Striopupilla Pilsbry, 1921;
TYPE SPECIES.- Pupa sterkiana Pilsbry, 1890.
DISTRIBUTION.- Baja California
Norte.
TAXONOMY.- Three species are
recognized. All occur in the study area.
Pupilla
(Stiopupilla) goniodon Pilsbry, 1927
Pupilla
goniodon Pilsbry,
1927; Proc. Calif. Acad. Sci., (4) 16: 172; pl. 7, fig. 3 (shell).- Smith, Miller, Christensen & Roth, 1990;
Proc. Calif. Acad. Sci., 47: 107-108.
TYPE LOCALITY.- Northwest
Anchorage, Guadalupe Island, Baja California Norte. Holotype CAS 2574.
DISTRIBUTION RECORDS.- BAJA
CALIFORNIA NORTE: Isla Guadalupe; Isla
San Benito del Oeste (Smith, et al,
1990).
Pupilla
(Stiopupilla) guadalupensis Pilsbry, 1927
Pupilla
guadalupensis
Pilsbry, 1927; Proc. Calif. Cad. Sci, (4) 16: 173-174; pl. 7, figs. 1, 2
(shell).
Pupilla
(Striopupilla) guadalupensis Pilsbry.- Smith,
Miller, Christensen & Roth, 1990; Proc. Calif. Acad. Sci., 47: 108.
TYPE LOCALITY.- 1000 feet above
Northeast Anchoraga, Guadalupe Island, Lower California. Holotype CAS 2575.
DISTRIUTION RECORDS.- BAJA
CALIFORNIA NORTE: known only from Isla Guadalupe (Smith, et al, 1990).
Pupilla
(Striopupilla) sterkiana (Pilsbry, 1890)
Pupa
sterkiana Pilsbry,
1890; Proc. Acad. Nat. Sci. Phila., 41: 411; pl. 2, figs. 2, 3 (shell).
Pupilla (Striopupilla) sterkiana
(Pilsbry). Pilsbry, 1921; Man. Conch.,
26: 156; pl. 19, figs. 16, 17. - Baker,
1963; Proc. Acad. Nat. Sci. Phila., 115: 202.-
Smith, Miller, Christensen & Roth, 1990; Proc. Calif. Acad. Sci.,
47: 107.
TYPE LOCALITY.- San Ramon, Lower
California. Lectotype ANSP 60466a (Baker, 1963: 202).
DISTRIBUTION RECORDS.- BAJA
CALIFORNIA NORTE: San Ramon; along ocean beach N and S of Bahia San Quintin; W
coast of San Quintin Peninsula, W of Winton Hill; Ensenada (Smith, et al,
1990).
Subfamily NEASOPUPINAE
Steenberg, 1925
DISTRIBUTION.- Madeira, South
Africa, Australia, Oceania, southern North America south to Brazil.
TAXONOMY.- Seven genera are
recognized. Four genera occur in the study area.
Genus Bothriopupa Pilsbry, 1898
Bothriopupa Pilsbry, 1898; Nautilus, 11:
119.- Pilsbry, 1917; Man. Conch., 24:
235.- Pilsbry, 1948; Land Moll. N. Amer.,
2: 1010-1011.
TYPE SPECIES.- Pupa variolosa Gould, 1848.
DISTRIBUTION.- Northern South
America, Florida, the Greater Antilles and Guatemala.
TAXONOMY.- About eight species
are recognized. Five are recorded from
the study area.
Bothriopupa
breviconus Pilsbry, 1917
Bothriopupa
breviconus Pilsbry,
1917; Man. Conch., 24: 230; pl. 28, figs. 9, 10 (shell).- Hinkley, 1920; Nautilus, 34: 44.
TYPE LOCALITY.- Mountains west of
Livingston, Dept. Izabal, Guatemala.
Holotype ANSP 107534.
DISTRIBUTION RECORDS. GUATEMALA, Dept. Izabal: mountains of Rio Cavech,
back of Cavech Village (Hinkley, 1920).
Bothriopupa
conoidea (Pfeiffer, 1853)
Pupa
conoidea Pfeiffer,
1853; Monographia heliciorum Viventium, 3: 533.
Bothriopupa
conoidea
(Pfeiffer). Pilsbry, 1917; Man. Conch.,
24: 231-232; pl. 28, figs. 7, 8, 11 (shell).-
Pérez & López, 2002: 96-97, text-fig. (distribution map).
TYPE LOCALITY.- Demerara.
DISTRIBUTION RECORDS.- Northern
South America and Nicaragua. NICARAGUA:
common along the Pacific Versant (Pérez & López, 2002).
Bothriopupa
leucodon (Morelet, 1851)
Pupa leucodon Morelet, 1851;
Testacea novissima, 2: 13.- Fischer
& Crosse, 1873; Miss. Sci. Mex. et L’Amer. Cent.: 311; pl. 14, figs. 2a, 2b
(shell).
Bothriopupa
leucodon
(Morelet). Pilsbry, 1917; Man. Conch.,
24: 232; pl. 29, figs. 1, 4 (shell).-
Goodrich & van der Scahlie, 1937: 30.
TYPE LOCALITY.- Salama, Dept.
Baja Verapaz, Guatemala.
DISTRIBUTION RECORDS.- GUATEMALA:
known only from the type locality; no precise locality (Goodrich & van der
Schalie, 1937).
Bothriopupa
tenuidens (C. B. Adams, 1841)
Pupa tenuidens C. B. Adams, 1841;
Proc. Bost. Soc. Nat. Hist., 1845: 15.-
Jacobson & Boss, 1973; Occ. Pap. on Moll., 9: 500; pl. 82, fig. 10
(lectotype).
Bothriopupa
tenuidens (C. B.
Adams). Pilsbry, 1917; Man. Conch., 24:
229-230; pl. 29, figs. 5-7, 11-13 (shell).-
Thompson & López, 1996: 49.-
Pérez & López, 2002: 98-100; text-fig. (distribution map).
TYPE LOCALITY.- Jamaica. Lectotype MCZ 270000 (Jacobson & Boss,
1973: 500).
DISTRIBUTION RECORDS.- Jamaica,
Puerto Rico, Cuba, Venezuela and Nicaragua. NICARAGUA, Dept. Managua: Lago de Xiloa (12°14’ N, 86°20’ W) (Thompson &
López, 1996); common along the Pacific Versant (Pérez & López, 2002).
Bothriopupa
variolosa (Gould, 1848)
Pupa
variolosa Gould,
1848; Proc. Bos. Soc. Nat. Hist., 3: 40.
Bothriopupa
variolosa
(Gould). Pilsbry, 1917; Man. Conch., 24:
230; pl. 29, figs. 2, 3 (shell).-
Pilsbry, 1948; Land Moll. N. Amer., 2: 1011-1012; figs. 539
(shell).- Bequaert & Clench, 1936;
Pub. Canegie Inst. Wash., (457): 65.
TYPE LOCALITY.- Florida.
DISTRIBUTION RECORDS.- Recorded
from Florida and Yucatan. YUCATAN:
Chotch Cenote, 2 mi. NE of Piste (Bequaert & Clench, 1936).
Genus Pupisoma Stoliszka, 1873
Pupisoma Stoliszka, 1873; Jour. Asiatic Soc.
Bengal, 42: 32.- Pilsbry, 1920; Man.
Conch., 26: 19.- Baker, 1927; Proc.
Acad. Nat. Sci. Phila., 79: 223 (anatomy).-
Pilsbry, 1948; Land Moll. N. Amer., 2: 1006.
TYPE SPECIES.- Pupisoma lignicola Stoliszka, 1873.
DISTRIBUTION.- Burma, India,
South Africa, Japan, Sri Lanka, Penang, India, Indonesia, Australia, New
Caledonia, Abyssinia, Gulf coastal area of the United States south to southern
Brazil.
TAXONOMY.- Five subgenera are
recognized (Zilch, 1959). One occurs in
the study area.
Subgenus Ptychopatula Pilsbry, 1889
Ptychopatula Pilsbry, 1889; Proc. Acad. Nat.
Sci. Phila, 41: 191.- Pilsbry, 1889;
Nautilus, 3: 62.- Zilch, 159: 173.
TYPE
SPECIES.- Helix caeca Guppy, 1868 (= Pupisoma
dioscoricola (C. B. Adams, 1845).
DISTRIBUTION.- Tropical and
subtropical Africa, Asia, Australia and the Americas.
TAXONOMY.- Numerous species. Six
species and one subspecies occur in the study area.
Pupisoma
(Ptychopatula) bailyi Pilsbry, 1934
Pupisoma (Ptychopatula) bailyi
Pilsbry, 1934; Man. Conch., 28: 116; pl. 24, fig. 15 (shell).
TYPE LOCALITY.- Cuernovaca,
Moreles [Morelos]. Holotype ANSP 156099.
DISTRIBUTION RECORDS.- MORELOS:
known only from the type locality.
Pupisoma
(Ptychopatula) comicolense Baker, 1927
Pupisoma comicolense Baker, 1927a;
Proc. Acad. Nat. Sci. Phila., 79: 224-226; pl. 15, fig. 4, 7 (reproductive
anatomy), fig. 5 (pallial organs), fig. 6 (radula); shell not described.- Baker, 1930a; Occ. Pap. Mus. Zool. Univ.
Mich., (220): 3-4; pl. 7, figs. 1, 2 (shell).-
Pilsbry, 1928; Nautilus, 41: 144.-
Pilsbry, 1934; Man. Conch., 28: 115-116; pl. 15, fig. 6 (shell).- Pilsbry, 1948; Land Moll. N. Amer., 2: 1006; figs.
537; a (radula), c, d (reproductive anatomy), fig. e (pallial organs).
TYPE LOCALITY.- Near Necaxa,
Puebla. Holotype UMMZ 159752.
DISTRIBUTION RECORDS.- PUEBLA:
known only from the immediate vicinity of the type locality (Baker, 1930a).
Pupisoma
(Ptychopatula) dioscoricola dioscoricola (C. B. Adams,
1845)
Helix
dioscoricola C. B.
Adams, 1845; Proc. Boston Soc. Nat. Hist., 2: 16.- Jacobson & Boss, 1963; Occ Baker, 1963;
Proc. Acad. Nat. Si. Phils., 115: 201. Pap. On Moll., 9: 346, 516; pl. 90, fig.
11 (lectotype).
Helix
punctum Morelet,
1851; Test. Noviss., II: 9.
Helix
caeca Guppy, 1868;
Proc. Sci. Assos. Trinidad: 241.-
Fischer & Crosse, 1872: 228; pl. 12, figs. 1a, 1b (shell).
Pupisoma
dioscoricola
(Adams). Pilsbry, 1920; Man. Conch., 26:
36-39; pl. 4, figs. 1-5 (shell).-
Pilsbry, 1926; Proc. Acad. Nat. Sci. Phila., 78: 133.- Pilsbry, 1948; Land Moll. N. Amer., 2:
1007-1008; figs. 538: 1-5 (shell).- Thompson,
1967; Bull. Fla. St. Mus., 11: 232.-
Thompson & López, 1996; Amer. Malac. Bull., 13: 49.- Pérez & López, 2002: 100-102, text-fig.
(distribution map).
TYPE LOCALITIES.- Helix dioscoricola: Dublin Castle,
Graywick, Jamaica; lectotype MCZ 275929 (Jacobson & Boss, 1973: 516). Helix
punctum: Merida, Yucatan. Helix caeca: Trinidad.
DISTRIBTION RECORDS.- Florida and
southern Texas to Brazil, Galapagos Islands.
PANAMA, Prov. Panama: Juan Mina (Pilsbry, 1926). NICARAGUA, Dept. Managua: Lago de
Xiloa (12°14’ N, 86°20’ W) (Thompson & López, 196); common along the Pacific
versant (Pérez & López, 2002).
CAMPECHE: 11.4 mi. E of Cayal; Cd. de Carmen (Thompson, 1967). QUINTANA ROO: 7.1 mi. NNW of Xiatil
(Thompson, 1967). TABASCO: Cardenas
(Thompson, 1967). YUCATAN: Tikul; 0.8
mi. NE of Becanchen (Thompson, 1967).
Pupisoma
(Ptychopatula) dioscoricola insigne Pilsbry, 1920
Pupisoma
dioscoricola insigne Pilsbry, 1920; Man. Conch., 26:
39-40; pl. 4; fig. 6, 7, 8 (shell).-
Baker, 1923a; 3.- Baker, 1925a:
2.- Baker, 1930a; Occ. Pap. Mus. Zool.
Univ. Mich., (220): 2.- Bequaert & Clench, 1936; Pub. Canegie
Inst. Wash., (457): 65.- Pilsbry, 1948; Land
Moll. N. Amer., 2: 1008; figs. 538: 6, 7, 8 (shell).- Basch, 1959; Occ. Pap. Mus. Zool. Univ.
Mich., (612): 12.- Correa-Sandoval &
Rodriguez, 2005; Acta Zool. Mex., 21: 59.
TYPE LOCALITY.- Brownsville,
Texas. Holotype ANSP 109013.
DISRIBUTION RECORDS.- Eastern
Mexico south to Venezuela. GUATEMALA, Dept. Peten: Tikal (Basch, 1956).
NUEVO LEON: numerous localities in southern part of state
(Correa-Sandoval & Rodriguez, 2005).
PUEBLA: Necaxa (Baker, 1930a).
SAN LUIS POTOSI: Valles River falls; Ganina River; Choy River Cave
(Pilsbry, 1920). VERACRUZ: Coatotolapan
(Baker, 1923a). YUCATAN: Izamal
(Pilsbry, 1920); Santa Ana Cenote, Villadolid (Bequaert & Clench, 1936).
Pupisoma
(Ptychopatula) mediamericanum Pilsbry, 1920
Pupisoma
mediamericanum
Pilsbry, 1920; Man. Conch., 26: 42-43; pl. 4, figs. 16, 17 (shell).- Baker, 1927a; Proc. Acad. Nat. Sci. Phila.,
79: 223-224; pl. 15, fig. 1 (reproductive anatomy), fig. 2 (pallial organs),
fig. 3 (jaw).- Baker, 1930a; Occ. Pap.
Mus. Zool. Univ. Mich., (220): 4.-
Pilsbry, 1948; Land Moll. N. Amer., 2: 1009; figs. 538: 16, 17 (shell).
TYPE LOCALITY.- Chama, Dept. Alta
Verapaz, Guatemala. Holotype and
paratypes ANSP 28270; syntypes ANSP 45727.
DISTRIBUTION RECORDS.- GUATEMALA;
known only from the type locality.
PUEBLA: Necaxa (Baker 1927a). VERACRUZ: 500 ft. above Orizaba (Pilsbry,
1920); Peñuela; Sumidero, 2625-3400 ft. alt. (Baker, 1930a).
Pupisoma
(Ptychopatula) michoacanensis Pilsbry, 1920
Pupisoma
michoacanensis
Pilsbry, 1920; Man. Conch., 26: 40; pl. 4, fig. 12.- Pilsbry, 1948; Land Moll. N. Amer., 2: 1009;
fig. 538: 12 (shell).
TYPE LOCALITY.- Morelia,
Michoacán. Holotype ANSP 77119.
DISTRIBUTION RECORDS.- MICHOACÁN:
known only from the type locality.
Pupisoma
(Ptychopatula) minus Pilsbry, 1920
Pupisoma
minus Pilsbry,
1920; Man. Conch., 26: 40-41; pl. 4, figs. 9, 11 (shell).- Pilsbry, 1948; Land Moll. N. Amer., 2:
1009-1010; figs. 538: 9-11.-
Naranjo-Garcia, 1991; Amer. Malac. Bull., 8: 168.- Pérez & López, 2002: 103-104; text-fig.
(distribution map).- Correa-Sandoval
& Rodriguez, 2005; Acta Zool. Mex., 21: 59.
TYPE LOCALITY.- Snapper Creek,
south of Coconut Grove,
DISTRIBUTION
Genus Nesopupa Pilsbry, 1900
Nesopupa Pilsbry, 1900; Proc. Acad. Nat.
Sci. Phila, 52: 432.- Pilsbry &
Cooke, in Pilsbry, 1920;
TYPE SPECIES.- Pupa tantilla Gould, 1847.
TAXONOMY.- Eleven subgenera are
recognized. One subgenus occurs in the
study area.
Subgenus Cocopupa Pilsbry & Cooke, 1920
Cocopupa Pilsbry & Cooke, 1920; Man.
Conch., 25: 323.
TYPE SPECIES.- Vertigo cocosensis Dall, 1900.
DISTRIBUTION.- Cocos Island,
Costa Rica.
TAXONOMY.- A single species is
known.
Nesopupa
(Cocopupa) cocosensis (Dall, 1900)
Vertigo
cocosensis Dall,
1900; Proc. Acad. Nat. Sci. Phila., 52: 98, pl. 8, fig. 13 (shell).
Nesopupa (Cocopupa) cocosensis
(Dall). Pilsbry & Cooke, in Pilsbry, 1920; Man. Conch., 25:
323-324; pl. 30, figs. 10, 11 (shell).
TYPE LOCALITY.- Cocos Island,
Costa Rica.
DISTRIBUTION RECORDS.- COSTA
RICA, Cocos Island: known only from the type locality.
Genus Sterkia Pilsbry, 1898
Sterkia Pilsbry, 1898; Nautilus, 11:
119.- Pilsbry, 1920; Man. Conch., 26:
49.- Pilsbry, 1948; Land Moll. N. Amer.,
2: 1012-1013.
TYPE SPECIES.-
Pupa calamitosa Pilsbry, 1889.
DISTRIBUTION.- California south
to Baja California Sur; northern South America, Guatemala, the West Indies and
south Florida.
TAXONOMY.- Two subgenera are
recognized. The genus includes six
species, all of which occur in the study area.
Subgenus Sterkia Pilsbry, 1898
DISTRIBUTION.- Southern
California and Baja California Norte.
TAXONOMY.- Two species are
recognized. Both occur in Baja
California Norte.
Sterkia
(Sterkia) calamitosa calamitosa (Pilsbry, 1889)
Pupa
calamitosa Pilsbry,
1889; Nautilus, 3: 61; pl. 1, fig. 7 (shell).-
Pilsbry, 1890; Proc. Acad. Nat. Sci. Phila., 41: 411; pl. 12, fig. 16
(shell).
Sterkia
calamitosa
(Pilsbry). Pilsbry, 1920; Man. Conch.,
26: 57; pl. 7, figs. 1-4 (shell).-
Pilsbry, 1948; Land Moll. N. Amer., 2: 1013-1014; figs. 540: 1-4
(shell).- Smith, Miller, Christensen
& Roth, 1990; Proc. Calif. Acad. Sci., 47: 110, fig. 14 (distribution
map).
TYPE LOCALITY.- Near the mouth of
the Rio San Tomas. Holotype ANSP 11602.
DISTRIBUTION RECORDS. BAJA
CALIFORNIA NORTE: Ensenada de Todos Santos (Pilsbry, 1948); hills W of Bahia
San Quintin (Smith, et al, 1990).
Sterkia
(Sterkia) calamitosa martiniana Pilsbry, 1927
Sterkia (Sterkia) calamitosa
martiniana Pilsbry, 1927; Proc. Calif. Acad. Sci. (4): 16: 186; pl. 12,
fig. 2 (shell).- Pilsbry, 1934; Man.
Conch., 28: 110; pl. 15, fig. 4 (shell).-
Pilsbry, 1948; Land Moll. N. Amer., 2: 1014.- Smith, Miller, Christensen & Roth, 1990;
Proc. Calif. Acad. Sci., 47: 110, fig. 14 (distribution map).
TYPE LOCALITY.- Isla San Martin,
Baja California Norte.
DISTRIBUTION RECORDS.- BAJA
CALIFORNIA NORTE: Isla San Martin.
Sterkia
(Sterkia) hemphilli (Sterki, 1890)
Pupa
hemphilli Sterki,
1890; Nautilus, 4: 27, 39; pl. 1, fig. 6 (shell).
Sterkia (Sterkia) hemphilli
(Sterki). Pilsbry, 1920; Man. Conch.,
26: 55; pl. 7, figs. 5-8, 11, 12 (shell).-
Pilsbry, 1927; Proc. Calif. Acad. Sci. (4): 16: 187.- Pilsbry, 1948; Land Moll. N. Amer., 2: 1013;
figs. 540: 5-8, 11, 12 (shell).
TYPE LOCALITY.- Banks of the Rio San Tomas, Baja California Norte. Holotype Carnegie Museum 62.20384.
DISTRIBUTION REDORDS.- Southern
California south to Baja California Norte.
BAJA CALIFORNIA NORTE: Ensenada
de Todos Santos; W shore of Bahia san Quintin; Punta Abreojos (Smith, et al, 1990)).
Subgenus Metasterkia Pilsbry, 1920
Metasterkia
Pilsbry, 1920; Man. Conch., 26: 50.-
Pilsbry, 1948; Land Moll. N. Amer., 2: 1014.
TYPE SPECIES.- Sterkia antilliensis Pilsbry, 1920.
DISTRIBUTION.- Northern South
America, the West Indies, Florida, Guatemala and Baja California Norte.
TAXONOMY.- Four species are
recognized. All occur in the study area.
Sterkia
(Metasterkia) antillensis Pilsbry, 1920
Sterkia (Metasterkia) antillensis
Pilsbry, 1920; Man. Conch., 26: 53-54; pl. 6, figs. 8-11 (shell).- Pérez & López, 2002: 105-107; pl. 15
(shell); text-fig. (distribution map).
TYPE LOCALITY.- El Abra, Viñales
Prov., Cuba.
DISTRIBUTION RECORDS.- Cuba,
Jamaica, Venezuela and Nicaragua.
NICARAGUA, common along the Pacific versant (Pérez & López, 2002).
Sterkia
(Metasterkia) bakeri Pilsbry, 1921
Sterkia bakeri
Pilsbry, 1921; Man. Conch, 26: 236; pl. 24, figs. 1, 2, 3 (shell).- Baker, 1923a: 3.- Baker, 1963; Proc. Acad. Nat. Sci. Phila.,
115: 200.
TYPE LOCALITY.- Hacienda de Coatotolapan, between the Rio San Juan and its tributary the Arroyo Hueyapan, Canton de Acayucan, Veracruz. Lectotype ANSP 141375 (baker, 1963: 200).
DISTRIBUTION RECORDS.- VERACRUZ:
known only from the type locality.
Holotype in the UMMZ.
Sterkia
(Metasterkia) eyriesii eyreisii (Drouet, 1859)
Pupa
eyriesii Drouet,
1859; Essai sur les mollusques terrestres et fluviatiles de la Guyane
Française: 71; pl. 2, figs. 16a, 17 (shell).
Sterkia (Metasterkia) eyriesii
eyreisii (Druuet). Pilsbry, 1920;
Man. Conch., 26: 61-62; pl. 6, figs. 1, 2, 4, 5 (shell).- Goodrich & van der Schalie, 1937; Misc.
Pub. Mus. Zool. Univ. Mich., (34): 30.-
Pilsbry, 1948; Land Moll. N. Amer., 2: 1018; fig. 542: 1, 2, 4, 5
(shell).
TYPE LOCALITY.- Ilet-la-Mère,
Guyana.
DISTRIBUTION RECORDS.- French
Guyana, Surinam, Trinidad and Guatemala.
GUATEMALA, no precise locality (Goodrich & van der Schalie,
1937). Dept. Alta Verapaz: Chamá
(Pilsbry, 1920).
TAXONOMY.- Two subspecies are
recognized. Sterkia eyriesii rhoadsi
Pilsbry, 1920, occurs in Florida.
Sterkia
(Metasterkia) clementina (Sterki, 1890)
Pupa
clementina Sterki,
1890; Nautilus, 4: 44; pl. 1, fig. 1 (shell).
Bifidaria
clementina oldroydi
Vanatta, 1916; Nautilus, 30: 48.
Sterkia
clementina
(Sterki). Pilsbry, 1920; Man. Conch.,
26: 54; pl. 7, figs. 9, 10, 13 (shell).-
Pilsbry, 1927; Proc. Calif. Acad. Sci., (4) 16: 176.- Pilsbry, 1948; Land Moll. N. Amer., 2: 1014,
1016; figs. 540: 9, 10, 13 (shell).- Smith,
Miller, Christensen & Roth, 1990; Proc. Calif. Acad. Sci., 47: 110.
TYPE LOCALITY.- Northeast
Anchorage, Isla Guadalupe, Baja California Norte. Holotype Carnegie Museum 62.20392.
DISTRIBUTION RECORDS.- BAJA
CALIFORNIA NORTE: Northeast Anchorage, Isla Guadalupe.
Subfamily PUPOIDINAE
Iredale, 1940
DISRIBUTION.- Europe, Africa,
Asia, Australia, Hawaii and North America
TAXONOMY.- Three extant genera
are recognized.
Genus Pupoides Pfeiffer, 1854
Pupoides Pfeiffer, 1854; Malacologische
Blätter, 1: 102.- Pilsbry, 1921; Man.
Conch., 26: 108, 138.- Pilsbry, 1948; Land
Moll. N. Amer., 2: 921-922.
Leucochila
Von Martens, 1860;
Die Heliceen: 205.
Leucochiloides Pfeiffer, 1978; Nomenel. Helic.
Viv.: 292.
TYPE SPECIES.- Pupoides: Bulimus nitidulus Pfeiffer, 1839.
Leucochila: Cyclostoma marginata Say, 1821. Leucochiloides:
Bulimus lardeus Pfeiffer, 1852.
DISTRIBUTION.- All continents.
TAXONOMY.- Four subgenera are
recognized. Two occur in the study area.
Three species occur in the study area.
Subgenus Pupoides Pfeiffer, 1854
DISTRIBUTION.-
North America, northern Asia, Europe and North Africa.
TAXONOMY.- A large number of
species have been named. Seven occur in
North America, two of which are found in the study area.
Pupoides (Pupoides) albilabris (C. B. Adams, 1841)
Cyclostoma
marginata Say,
1821; Jour. Acad. Nat. Sci. Phila., 2: 172 (not Cyclostoma marginatus G. Fischer, 1807).
Pupoides
maerginatus Pilsbry
& Vanatta, 1900; Proc. Acad. Nat. Sci. Phila., 52: 586.- Pilsbry, 1904; Proc. Acad. Nat. Sci. Phila,
55: 766.- Pilsbry, 1921; Man. Conch.,
26: 111; pl. 12, figs. 1-7 (shell).
Pupa
fallax “Say” Gould,
1843; Boston Jour. Nat. Hist., 4: 357.
Pupa
albilabris C. B.
Adams, 1841; Amer. Jour. Sci., 40: 271. (new name for Cyclostoma marginata Say).
Pupoides
albilabris (C. B.
Adams, 1841). Pilsbry, 1948; Land Moll.
N. Amer., 2: 921-923; figs. 499: 1-7. (shell).-
Pilsbry, 1953; Proc. Acad. Nat. Sci. Phila., 105: 164.- Bequaert & Miller, 1973: 78-79,
176-177.- Smith, Miller, Christensen
& Roth, 1990; Proc. Calif. Acad. Sci., 47: 108; fig. 12 (map).- Naranjo-Garcia, 1991: 168.
TYPE LOCALITY.- “Upper
Missouri”.
DISTRIBUTION RECORDS.- Widely
distributed in North America. BAJA
CALIFORNIA NORTE: Isla Angel de la Guardia; Isla San Lorenzo (Smith, et al, 1990). BAJA CALIFORNIA SUR: numerous localities
(Smith, et al, 1990). SINALOA: Rio Fuerte, San Blas (Pilsbry,
1953). SONORA: Rio Mayo, at Navajo; Rio
Yaqui, 11 mi. N of Cajeme; Cerro Zaporxa, E of Cameje; Arroyo San Rafael, San
Bernardo (Pilsbry, 1953); N end of Sierra El Viejo, 550 m alt. (30º25.41’ N,
112º22.5’ W); SW part of Sierra El Viejo, 500 m alt. (30º23.8’ N, 112º36.6’ W);
Arroyo Cerro de Oro, ca. 0.25 mi. from Cerro de Oro, on rd. to Rayon, 600 m
alt. (29º36.8’ N, 110º37.8’ W) (Naranjo-Garcia, 1991). CHIHUAHUA: Presa Chihuahua, 7 mi. from Cd.
Chihuahua (Bequaert & Miller, 1973).
NUEVO LEÓN: Topo Chico, nr. Monterey (Pilsbry, 1904). TAMAULIPAS: Tampico (Pilsbry, 1948); drift of
Rio Purificación, 24 mi. W of Padilla (Bequaert & Miller, 1973).
Pupoides (Pupoides) catalinensis Hanna, 1923
Pupoides
catalinensis Hanna,
1923; Proc. Calif. Acad. Sci., (4), 12: 514; pl. 10, figs. 1-4 (shell).- Pilsbry, 1926; Man. Conch., 27: 249-250;
text-fig. 17 (shell).- Smith, Miller,
Christensen & Roth, 1990; Proc. Calif. Acad. Sci., 47: 108; map fig. 12.
TYPE LOCALITY.- Santa Catalina
Island, Baja California Sur.
DISTRIBUTION RECORDS.- BAJA CALIFORNIA SUR: known only from Santa
Catalina Island (Smith, et al, 1990).
Subgenus Ischnopupoides Pilsbry, 1926
Ischnopupoides Pilsbry, 1926; Man. Conch., 27:
250.- Pilsbry, 1948; Land Moll. N. Amer.,
2: 924.
TYPE SPECIES.- Bulimus cordatus Pfeiffer, 1856.
DISTRIBUTION.- SW United States,
western Mexico and Bolivia.
TAXONOMY.- Three species are
recognized. One occurs in the study
area.
Pupoides
(Ischnopupoides) cordatus (Pfeiffer, 1856)
Bulimus
cordatus Pfeiffer,
1856; malak. Blätter, 3: 46.
Pupa
cordata (Pfeiffer).
Binney & Bland, land and freshwater shells of North America, 1: 241; fig.
418 (shell).
Pupoides (Pupoides) cordatus
(Pfeiffer). Von Martens; 1893:
330.- Pilsbry, 1921; Man. Conch., 26:
119-120; pl. 11, figs. 14, 15 (shell).
TYPE LOCALITY.- Mazatlan,
Sinaloa.
DISRIBUTION RECORDS.- SINALOA:
known only from the type locality.
Subfamily GASTROCOPTINAE
Pilsbry, 1918
DDISRIBUTION.- Australia, Africa,
North and South America, Asia, Sri Lanka, the Philippines, and Oceania.
TAXONOMY.- A single extant genus,
Gastrocopta, is recognized. Two South American genera are referred
questionably to the subfamily.
Genus Gastrocopta Wollaston, 1878
Gastrocopta Wollaston, 1878; Testacea Atlantica
…: 515.- Pilsbry, 1916; Man. Conch., 24:
6-12.
Leucochilus Boettger, 1881; Conchologische
Mittheilungen, 1: 64 (not Leucochila Von
Martens, 1860).
Bifidaria Sterki, in Pilsbry, 1891; Proc. Acad. Nat. Sci. Phila., 43: 315.
Eubifidaria
Sterki, 1893; Nautilus, 6: 101.
TYPE SPECIES.- Gastrocopta: Pupa acarus Benson, 1856. Leucochilus: Pupa armifera Say, 1821. Bifidaria: Pupa servilis Gould, 1843. Eubifidaria: “Pupa hordeacea Gabb” of Sterki, = Bifidaria cristata Pilsbry & Vanatta, 1900.
DISTRIBUTION.- Worldwide.
TAXONOMY.- Nine subgenera are
recognized (Zilch, 1959: 160-162).
Subgenus Gastrocopta Wollaston
DISRTIBUTION.- Tropical and warm
portions of America, Africa, Mascarene Island, Sri Lanka and the Philippines.
TAXONOMY.- Numerous species. Four occur in the study area.
Gastrocopta
(Gastrocopta) cristata (Pilsbry & Vanatta, 1900)
Bifidaria
procera cristata
Pilsbry & Vanatta, 1900; Proc. Acad. Nat. Sci. Phila., 52: 595; pl. 22,
figs. 4, 5 (shell).
Gastrocopta (Gastrocopta) cristata
(Pilsbry & Vanatta). Pilsbry, 1916;
Man. Conch., 24: 68-69; pl. 13, figs. 6, 8-12 (shell).- Pilsbry, 1948; Land Moll. N. Amer., 2: 911,
913; figs. 493: 6, 8-12.- Branson, McCoy
& Sisk, 1964; S.W. Nat., 9: 104.-
Bequaert & Miller, 1973: 169-170.
TYPE LOCALITY.- Camp Verde,
Yavapai County, Arizona; 3200 ft. Alt. (Bequaert & Miller, 1973).
DISTRIBUTION RECORDS.- Nebraska, Kansas, Oklahoma, west Texas, New
Mexico and Arizona and adjacent NW Mexico.
BAJA CALIFORNIA NORTE. SONORA: drift of Rio Bevispe, 21 mi. S of Agua
Prieta; drift of Rio Yaqui, 4 mi. N of Cd. Obregon (Branson, et al, 1964); drift of Rio Sonoyta,
Sonoyta (Bequaert & Miller, 1973).
Gastrocopta (Gastrocopta) pellucida hordeacella (Pilsbry, 1890)
Pupa
hordeacella
Pilsbry, 1890; Proc. Acad. Nat. Sci. Phila., 44; pl. 1, figs. f-k
(shell).- Hinkley, 1907; Nautilus, 21:
78.
Bifidaria
pellucid hordeacella
(Pilsbry). Pilsbry, 1904; Proc. Acad.
Nat. Sci. Phila., 55: 766.
Gastrocopta
pellucida hordeacella
(Pilsbry).- Pilsbry, 1916; Man. Conch., 24: 78-80; pl. 17, figs. 1-4; pl. 16
(shell).- Pilsbry, 1948; Land Moll. N.
Amer., 2: 913-915; figs. 494: a-d; figs. 495 (Shell).- Basch, 1959; Occ. Pap. Mus. Zool. Univ.
Mich., (612): 12.
Bifidaria
hordeacella parvidens Sterki,
1899; Nautilus, 12: 128.
Gastrocopta
pellucida parvidens (Sterki).
Pilsbry, 1948; Land Moll. N. Amer., 2: 916; fig. 494: c).- Hanna, 1923; Proc. Calif. Acad. Sci. (4), 13:
515.
Pupa
pellucida Pfeiffer,
1841. Fischer & Crosse, 1873; Miss.
Sci. Mex., I: 311.- Strebel, 1880:
91.- Von Martens, 1898: 328 (in part).
Gastrocopta
pellucida
(Pfeiffer). Hinkley, 1920; Nautilus, 34:
52.- Rehder, 1966; Proc. Biol. Soc.
Wash., 79: 383.- Thompson, 1967; Bull.
Fla. St. Mus., 11: 231.- Bequaert &
Miller, 1973: 79-81.- Naranjo-Garcia,
1991; Amer. Malac. Bull., 8: 167-168.-
Correa-Sandoval, 1993; Rev. Biol. Trop., 41: 675.- Correa-Sandoval, Gutierrez & Reza, 1998;
Acta Zooll. Mex., (73): 134.-
Correa-Sandoval, 2000; Acta Zool. Mex., (79): 8.- Correa-Sandoval, & Castro, 2002; Acta
Zool. Mex., (86): 235.- Thompson &
López, 1996; Amer. Malac. Bull., 13: 49.-
Pérez & López, 2002: 88-91; (distribution map in Nicaragua).- Correa-Sandoval & Rodriguez, 2005; Acta
Zool. Mex., 21: 59.
TYPE LOCALITIES.- Pupa pellucida: Cuba (Pilsbry,
1916). Pupa hordeacella: New Braunfels, Texas; holotype and paratypes ANSP
60460. Bifidaria hordeacella parvidens: around Jerome, Yavapai County,
Arizona; syntypes in the Carnegie Museum.
DISTRIBUTION RECORDS.- Widely
distributed in the West Indies, Mexico and Guatemala. Bequaert & Miller (1973) state that the
species apparently has a continuous distribution over Mexico. NICARAGUA, Dept. Managua: Lago de
Xiloa (12°14’ N, 86°20’ W) (Thompson & López, 1996); abundant records from
the Pacific versant (Pérez & López, 2002). GUATEMALA, Dept. Alta Verapaz: Chama
(Hinkley, 1920). Dept. Baja Verapaz: Salama. Dept.
Guatemala: Barranca de Villalobos,
nr. Cd. Guatemala. Dept. Peten: Tikal
(Basch, 1959). Dept. Sacatepequez:
Antigua, 4500 ft. alt. (Von Martens, 1898). BAJA CALIFORNIA SUR: numerous
localities (Smith, et al, 1990). CAMPECHE: 7.1 mi. SW of Cd. Camopeche; 8.1
mi. SW of Champoton (Thompson, 1967).
NUEVO LEON: numerous records for southern part of state (Correa-Sandoval
& Rodriguez, 2005). QUINTANA ROO:
Tulum (Rehder, 1966). SAN LUIS POTOSI:
Cd. Valles (Pilsbry, 1916); numerous localities (Correa-Sandoval, et al, 1998). SONORA: numerous localities (Naranjo-Garcia,
1991). TABASCO: San Juan Bautista (Von
Martens, 1898). TAMAULIPAS: Cd. Victoria
(Pilsbry, 1916); Tampico (Hinkley, 1907).
YUCATAN: Sisal; Merida (Fischer & Crosse, 1873); 1.0 mi. SSE of
Telchac; 0.8 mi. NE of Becanchen (Thompson, 1963). VERACRUZ: Antigua (Pilsbry,
1916); Veracruz (Strebel, 1880); Rancho El Sol, Naranjos (21°20’00” N,
97°43’16” W) Carr. 1 km NE of Tuxpan, km 234 (20°49’11” N, 97°30’00” W)
(Correa-Sandoval, 1999).
TAXONOMY.- Recognition of Gastrocopta pellucida hordeacella
(Pilsbry, 1890) as a valid subspecies is controversial. Bequaert & Miller (1973) synonymised both
hordeacella and parvidens with Gastrocopta
pellucida pellucida (Pfeiffer, 1941).
Later, Smith, et al, 1990,
resurrected hordeacella as a distinct
subspecies. Bifidaria hordeacella parvidens (Clapp, 1899) is a synonym of hordeacella. I follow these authors by assigning all
mainland populations to hordeacella.
Gastrocopta
(Gastrocopta) riograndensis (Pilsbry & Vanatta, 1900)
Bifidaria
riograndensis
Pilsbry & Vanatta, 1900; Proc. Acad. Nat. Sci. Phila., 52: 596.
Gastrocopta (Gastrocopta) riograndensis
(Pilsbry & Vanatta). Pilsbry, 1916;
Man. Conch., 24: 69-70; pl. 12, figs. 9, 10 (shell).- Pilsbry, 1948; Land Moll. N. Amer., 2: 914;
figs. 492: 9, 10 (shell).- Thompson,
1967; Bull. Fla. St. Mus., 11: 231.-
Neck, 1980; Veliger, 23: 180-182.-
Smith, Miller, Christensen & Roth, 1990; Proc. Calif. Acad. Sci.,
47: 106.
Bifidaria
procera
(Gould). Hinkley, 1907; Nautilus, 21:
77.
TYPE LOCALITY.- Lower Rio Grande
valley, river drift at Hidalgo, Hidalgo Co., Texas. Holotype ANSP 60137.
DISTRIBUTION RECORDS.- Southwest
Texas, northern Mexico. BAJA CALIFORNIA SUR: Juncalito (Smith, et al, 1990). CAMPECHE: 8.1 mi. SW of Chamopoton (Thompson,
1967). SAN LUIS POTOSI: falls below Cd. Valles; canyon below Los Canoas
(Pilsbry, 1916). TAMAULIPAS: Panuco
River Valley, Tampico (Hinkley, 1907; Pilsbry, 1916). YUCATAN: 1.0 mi. SSE of Puerto Techac
(Thompson, 1967).
TAXONOMY.- The identity of
specimens from Baja California Sur is uncertain (Smith, et al, 1990).
Gastrocopta
(Gastrocopta) servilis (Gould, 1843)
Pupa
servilis Gould,
1843; Boston Jour. Nat. Hist., 4: 356; pl. 16, fig. 14 (shell).
Pupa
pellucida Pfeiffer
(in part). Strebel, 1880: 91; pl. 4, fig. 19; pl. 15,
fig. 10 (shell).- Von Martens, 1893;
Biol. Cent. Amer.: 328.
Gastrocopta (Gastrocopta) servilis
(Gould). Pilsbry, 1916; Man. Conch., 24:
70-74; pl. 14, figs. 4, 5, 6, 7 (shell).-
Pilsbry, 1926; Proc. Acad. Nat. Sci. Phila., 78: 133.- Rehder, 1966; Proc. Biol. Soc. Wash., 79:
283.- Thompson, 1967; Bull. Fla. St.
Mus., 11: 231.- Thompson & López,
1996; Amer. malac. Bull., 13: 48.- Pérez
& López, 2002: 93-95 (text-fig. distribution map).
TYPE LOCALITY.- Matanzas, Cuba.
DISTRIBUTION RECORDS.- Generally
distributed throughout the West Indies, including Bermuda; scattered records in Central America and
Mexico. PANAMA, Canal Zone: Barro Colorado Island.
Prov. Panama: Juan Mina (Pilsbry, 1926).
NICARAGUA: Bluefields; abundant along the Pacific versant (Pérez &
López, 2002). GUATEMALA, Dept. Sacatepequez: Antigua. Dept. Baja Verapaz: Salama. Dept. Guatemala: Barranca de Villalobos, nr. Cd. Guatemala. CAMPECHE: Isla de Carmen (Pilsbry,
1916). MORELOS: Yautepec. QUINTANA ROO: Tulum (Rehder, 1966). VERACRUZ: Veracruz, on the beach (Strebel,
1880). YUCATAN: Merida (Pilsbry, 1916).
Subgenus Albinula Sterki, 1892
Albinula Sterki, 1892; Nautilus, 6:
101.- Pilsbry, 1916; Man. Conch., 24:
13-14.- Pilsbry, 1948; Land Moll. N.
Amer., 2: 874.
TYPE SPECIES.- Pupa contracta Say, 1822.
DISTRIBUTION.- Eastern North
America.
TAXONOMY.- Six species are
recognized. One occurs in the study
area.
Gastrocopta (Albinula) contracta contracta (Say, 1822)
Pupa
contracta Say,
1822; Jour. Acad. Nat. Sci. Phila, 2: 374.-
Pilsbry, 1891; Proc. Acad. Nat. Sci. Phila., 43: 315.
Bifidaria
contracta (Say).
Pilsbry, 1904; Proc. Acad. Nat. Sci. Phila., 55: 766.- Hinkley, 1907; Nautilus, 21: 78.
Gastrocopta
(Albinula) contracta contracta (Say). Pilsbry, 1916; Man. Conch., 24: 22-23;
pl. 2, figs. 9-12 (shell).- Pilsbry,
1948; Land Moll. N. Amer., 2: 880-881; figs. 474: 9-12.- Pilsbry, 1953; Proc. Acad. Nat. Sci. Phila.,
105: 161.- Correa-Sandoval, Gutierrez
& Reza, 1998: 14.
TYPE LOCALITY.- Occoquan,
Virginia.
DISTRIBUTION RECORDS.- Eastern
North America from Ontario west to Manitoba and south to Florida and Veracruz;
Cuba and Jamaica. MORELOS: Yautepec
(Pilsbry, 1891, 1904). NAYARIT: drift
debris of Rio Fuerte, San Blas (Pilsbry, 1953).
NUEVO LEON: on Rio Marisco, ca. 25 km S of Monterey (Pilsbry,
1953). SAN LUÍS POTOSÍ: Cascadas de
Tamasopo, 430 m alt. (21°56’05” N, 99°25’00” W); Carr. Tamuín-San Vicente, 3 km
after x-sec. to El Chote, 80 m alt. (21°52’51” N, 98°40’31” W); Río Coy, carr.
to Cd. Valles, 170 m alt. (21°45’36” N, 98°57’22” W); 15.5 km NE of Tamazunchale, 140 m alt. (21°12’26” N,
98°53’25” W); Río Moctezuma, Tamazunchale (21°15’21” N, 98°48’56” W)
(Correa-Sandoval, et al, 1998). SONORA: Arroyo San Rafael, San Bernardo
(Pilsbry, 1953). TAMAULIPAS: Tampico
(Hinkley, 1907). VERACRUZ: Texolo; nr.
Orizaba (Pilsbry, 1904).
Subgenus Geminidens Pilsbry, 1930
Geminidens Pilsbry, 1930; Proc. Acad. Nat.
Sci. Phila., 82: 351.- Zilch, 1959: 160.
TYPE SPECIES.- Bothriopupa geminidens Pilsbry, 1917.
DISTRIBUTION.- Venezuela,
Trinidad and Panama.
TAXONOMY.- The subgenus includes
a single species.
Gastrocopta
(Geminidens) geminidens (Pilsbry, 1917)
Bothriopupa
geminidens Pilsbry,
1917; Man. Conch., 24: 228-229; pl. 28, figs. 12, 13, 14 (shell).- Baker, 1963; Proc. Acad. Nat. Sci. Phila.,
115: 201.
Gastrocopta
geminidens (Pilsbry). Pilsbry, 1926; Man. Conch., 27: 228.
Gastrocopta (Geminidens) geminidens
(Pilsbry). Pilsbry, 1930; Proc. Acad.
Nat. Sci., Phila., 80: 351.- Haas, 1960:
13.- Thompson & López, 1996: 49.- Pérez & López, 2002: 83-88; distribution
map.
TYPE LOCALITY.- Cariaquita,
[Estado Sucre], Venezuela. Holotype ANSP
105200a (Baker, 1963: 201).
DISTRIBUTION RECORDS.- Known from
Panama, Venezuela, Margarita Island and Trinidad. PANAMA, Prov.
Panama: Old Panama City (Pilsbry, 1930).
COSTA RICA: Dept. Guanacaste (Pérez & López,
2002). NICARAGUA, Dept. Managua: Lago de
Xiloa (12°14’ N, 86°20’ W) (Thompson & López, 1996); Dept. Chontales; Dept. Masaya; Dept. San Juan del
Rio (Pérez & López, 2002).
Subgenus Immersidens Pilsbry & Vanatta, 1900
Immersidens Pilsbry & Vanatta, 1900; Proc.
Acad. Nat. Sci. Phila., 52: 606.
TYPE SPECIES.- Bifidaria ashmuni 1898.
DISTRIBUTION.- Central Plateau of
North America south to Brazil and Argentina.
TAXONOMY.- The subgenus contains
numerous species. Seven species and one
subspecies occur in the study area.
Gastrocopta
(Immersidens) allyni Roth & Christensen, 1984
Gastrocopta (Immersidens) allyni Roth
& Christensen, 1984; Proc. Biol. Soc. Wash., 97: 245-250; figs. 1, 2
(shell), fig. 3 (distribution map).- Smith,
Miller, Christensen & Roth, 1990; Proc. Calif. Acad. Sci., 47: 106; fig. 10
(distribution map).
TYPE LOCALITY.- In drift, arroyo
24 km south of La Paz, Baja California Sur.
Holotype CAS 32993.
DISTRIBUTION RECORDS.- BAJA
CALIFORNIA SUR: southern tip of peninsula, numerous locality records (Smith, et al, 1990).
Gastrocopta (Immersidens) ashmuni (Sterki, 1898)
Bifidaria
ashmuni Sterki,
1898; Nautilus, 12: 90.- Pilsbry &
Ferriss, 1906; Proc. Acad. Nat. Sci. Phila., 58: 144.
Gastrocopta
(Immersidens) ashmuni (Sterki). Pilsbry,
1916; man. Conch., 24: 41-44; pl. 6, figs. 5, 6, 7; text-figs. 13a-c
(shell).- Pilsbry, 1948; Land Moll. N.
Amer., 2: 896-898; figs. 483: 5-7; figs. 484a-c (shell). Branson, McCoy & Sisk, 1964: 104.- Bequaert & Miller, 1973: 159-163.- Naranjo-Garcia, 1991: 166-167.- Correa-Sandoval & Rodriguez, 2005: 59.
TYPE LOCALITY.- Santa Rita
Mountains, on east slope of range, 20 miles west of Crittenden, Santa Cruz
County, Arizona (Bequaert & Miller, 1973: 159).
DISRIBUTION RECORDS.- CHIHUAHUA: Rio Piedras Verdes, ca. 9 km above
Colonia Juarez (ca. 30º15’ N, 108º W) (Pilsbry, 1953); Presa Chihuahua, 7 mi. S
of Cd. Chihuahua, 4800 ft. alt. (28º35’ N, 106º05’ W). NUEVO LEON: km 66, rd. from Linares to San
Roberto (24º45’22” N, 100º02’18” W); rd. from Linares to Dr. Arroyo (24º31’05”
N, 99º58’55” W) (Correa-Sandoval & Rodriguez, 2005). SONORA: Rio Nacozari,
7 mi. S of Nacozari (Branson et al, 1964); nr. Nacozari, 3550-4200 ft. alt.;
Nogales; Sierra de Santo Niño, N of Mina El Milagro, on rd. to Sahuaipa, 4000
t. alt. (ca. 29º N, 109º30’ W); nr. Magdalena, 3650-4250 ft. alt. (ca. 30º5’ N,
111º W); nr. Zocozari, 3550-4000 ft. alt. (ca. 30º N, 110º W); Rancho Pinos
Altos, Sierra Nacori, 5800 ft. alt. (29º45’N, 108º30’W) (Bequaert & Miller,
1973); Sierra Las Minitas, 1400 m alt. (31º11.1’ N, 109º04.7’ W); slope above arroyo, along Rayon-Cerro de Oro
rd., ca. 4.2 mi. from Cerro de Oro (29º38.9’ N, 110º36.6’ W); S edge of Sierra
El Pinito, ca. 10.1 km W of Aribabi (30º52.7’ N, 110º42.7’ W); Sierra Batamote,
Mina El Milagro (29º57’10” N, 109º32’ W); SW end of Sierra Los Embudos
(31º11.8’ N, 109º05.2’ W) (Naranjo-Garcia, 1991).
Gastrocopta
(Immersidens) cochisensis (Pilsbry & Ferriss, 1910)
Bifidaria
cochisensis Pilsbry
& Ferriss, 1910; Proc. Acad. Nat. Sci. Phila, 62: 139; figs. 32, 33
(shell).
Gastrocopta (Immersidens) cochisensis
(Pilsbry & Ferriss). Pilsbry, 1916;
Man. Conch., 24: 44-46; pl. 6, figs. 1-4, 8 (shell).- Pilsbry, 1948; Land Moll. N. Amer., 2:
899-900; figs. 483: 1-4, 8; fig. 486 (shell).-
Bequaert & Miller, 1973: 163-164.
TYPE LOCALITY.- Tanner Canyon
[now Garden Canyon], Huachuca Mountains, Arizona. Holotype and paratypes ANSP 97444.
DISRIBUTION RECORDS.- CHIHUAHUA:
Rio Piedras Verdes, below Pacheco, 5900 ft. alt.; Sierra de la Breña, ca. 17.5
mi. from Mata Ortiz, on rd. to Pacheco, 7000 ft. alt. (Pilsbry, 1953). SINALOA: Rio Fuerte, San Blas (Pilsbry,
1953). SONORA: arroyo, 8 km S of Guaymas (Pilsbry, 1953).
Gastrocopta
(Immersidens) dalliana dalliana
(Sterki, 1898)
Bifidaria
dalliana Sterki,
1898; Nautilus, 12: 91.
Gastrocopta
dalliana
(Sterki). Pilsbry, 1916; Man. Conch.,
24: 49; pl. 8, figs. 10, 11 (shell).-
Pilsbry, 1948; Land Moll. N. Amer., 2: 901-902; fig. 488 (shell).
Gastrocopta (Immersidens) dalliana dalliana (Sterki). Branson, McCoy & Sisk, 1964: 104.- Bequaert & Miller, 1973: 165-167.- Naranjo-Garcia, 191: 167.
Gastrocopta
dalliana media Pilsbry,
1916; Man. Conch., 24: 50; pl. 8, figs. 10-11 (shell).
TYPE LOCALITIES.- Bifidaria dalliana: Ephraim Canyon, Nogales, Santa Cruz County, Arizona; 3800
ft. alt. (Bequaert & Miller, 1973). Gastrocopta dalliaa media: Montezuma Wells, near Rimrock, Yavapai
County, Arizona; holotype ANSP 82918.
DISRIBUTION RECORDS.- CHIHUAHUA:
cliff talus on N side of Rio Piedras Verdes, 8-9 km above Colonia Juarez, 5900
ft. alt. (ca. 30º10’ N, 108º10’ W) (Pilsbry, 1953). SONORA:
spring-fed march 60 mi. S of Nogales; San Carlos bay, 10 mi. N of
Guaymas; S end of Sierra Purico, 6300 ft. alt. (ca. 30º31’ N, 109º45’ W),
mountain 5 mi. S of Magdalena, 4250 ft. alt.;
NW side of Rio Nacozari, 4200 ft. alt. (ca. 30º35’ N, 111º W) (Bequaert
& Miller, 1973); S edge of Sierra El Pinito, ca. 10.1 km W of Aribabi (30º52.7’
N, 110º42.7’ W); N end of range, Sierra El Viejo, 550 m alt. (30º24.1’ N
112º22.5’ W); Rancho Tres Marias, W of Alamo 27º06’56” N, 109º09’18” W); N end
of Sierra El Viejo, 500 m alt. (30º23.8’ N, 112º23.6’ W) (Naranjo-Garcia,
1991).
Gastrocopta
(Immersidens) dalliana bilamellata (Sterki & Clapp, 1909)
Bifidaria
bilamellata Sterki
& Clapp, 1909; Nautilus, 22 126; pl. 8, fig. 7.- Pilsbry & Ferriss, 1910; Proc. Acad. Nat.
Sci. Phila., 62: 143; fig. 36b.
Gastrocopta
bilamellata (Sterki
& Clapp). Pilsbry, 1916; man.
Conch., 24: 51-52; pl. 8, figs. 1-4; text-fig. 18 (shell).- Pilsbry, 1948; Land Moll. N. Amer., 2: 903;
fig. 489, figs. 490: 1-4 (shell).
Gastrocopta (Immersidens) dalliana
bilamellata (Sterki & Clapp).
Bequaert & Miller, 1973: 168-169.-
Naranjo-Garcia, 1991: 167.
TYPE LOCALITY.- Foothills of
Plomosa Range, about 8 miles east of Quartzite, Yuma County, Arizona. Holotype in the Clapp Collection, Carnegie
Museum.
DISTRIBUTION RECORDS.- Southwest
Arizona and northwest Sonora. SONORA: S
side of Rio Sonora, nr. Hermosillo, 900 ft. alt. (ca. 29º N, 111º W); Arroya
San Rafael, San Bernardo, 900 ft. alt. (ca. 27º25’ N, 108º54’ W); north
foothills of Cerro Zaporxa, E of Obregon (ca. 28º N, 109º W); drift in wash W
of Pinacate Peak, ca. 45 mi. NW of Sonoyta, 1200 ft. alt. (ca. 31º50’ N,
112º50’ W); drift in Rio Sonoyta, Sonoyta, 1500 ft. alt. (ca. 31º50’ N, 112º50’
W) (Bequaert & Miller, 1973); Cerro El Oro (29º36.8’ N, 110º37.8’ W); SW
end of Sierra Los Embudos (31º11,8’ N, 109º05.2’ W) (Naranjo-Garcia, 1991).
Gastrocopta
(Immersidens) gularis Thompson & López, 1996
Gastrocopta (Immersidens) gularis
Thompson & López, 1996; Amer. Malacological Bulletin, 13: 47-53; figs. 5-8,
15-19, 21, 25 (shell).- Pérez &
López, 2002: 86-88; (text-fig, distribution map).
TYPE LOCALITY.- Lago de Xiloa,
Dept. Managua, Nicaragua (12°14’ N, 86°20’ W).
Holotype FLMNH 247775.
DISTRIBUTION RECORDS. COSTA RICA,
Prov. Guanacaste: Parque Nacional Santa Rosa, banks of the Rio Nisparal
(10°48’33” N, 85°39’25” W); 2.5 km N of Estacion Argelia (10°47’50” N,
85°39’15” W); Sendero Guayacan (10°21’02” N, 85°21’08” W) (Thompson &
López, 1996). NICARAGUA, Dept. Boaco: Las Canoas (13°13’6: N, 85°52’50” W). Dept.
Leon: Lago Asososca (12°26’ N, 86°40’
W). Dept.
Managua: Platanal (12°27’15” N,
86°05’34” W); Tamarindo (12°29’30” N, 86°05’25” W). Masaya Dept.: Lago Apoyo (11°55’ N, 86°03’ W). Dept. Matagalpa: Dario
(12°43’ N, 86°12’ N) (Thompson & López, 1996). Dept.
Chontales; Dept. Masaya; Dept. San Juan del Rio (Pérez & López, 2002).
Gastrocopta
(Immersidens) prototypus (Pilsbry, 1899)
Bifidaria
prototypus Pilsbry,
1899; Proc. Acad. Nat. Sci. Phila., 51: 400.-
Pilsbry, 1904; Proc. Acad. Nat. Sci. Phila., 55: 766; pl.50, figs 7,
7a.- Pilsbry & Ferriss, 1910; Proc.
Acad. Nat. Sci. Phila., 62: 142, fig. 35.-
Baker, 1963; Proc. Acad. Nat. Si. Phila., 115: 202.
Gastrocopta (Immersidens) prototypus
(Pilsbry). Pilsbry, 1916; Man. Conch.,
24: 47-48; pl. 7, figs. 1-5 (shell).
Gastrocopta (Immersidens) prototypus
prototypus (Pilsbry). Bequaert &
Miller, 1973: 164-165.
Bifidaria
cochisensis oligobasodon Pilsbry
& Ferriss, 1910; Proc. Acad. Nat. Sci. Phila., 62: 141; figs. 34a-c
(shell).
Gastrocopta (Immersidens) prototypus
basidentata Pilsbry, 1916; Man. Conch., 24: 48; pl. 7, figs. 6, 7 (shell).
TYPE LOCALITIES.- Bifidaria prototypus: Huingo, near Lago
de Cuitzco, 40 km NW of Morelia, Michoacan (Bequaert & Miller, 1973); Lectotpe ANSP 77201a (Baker, 1963: 202). Bifidaria
cochisensis oligobasodon; Ash canyon, SE slope of Huachuca Mountains,
Cochis County, Arizona, 5000 ft. alt. (Bequaert & Miller, 1973: 165). Gastrocopta
prototypus basidentata: Guatemala
City, Guatemala.
DISTRIBUTION RECORDS.- Arizona and widely scattered localities south
to Guatemala. GUATEMALA: Dept. Guatemala: Cd. Guatemala. JALISCO: Guadalajara (Pilsbry, 1916). MICHOACAN: Huingo.
Gastrocopta
(Immersidens) rixfordi Hanna, 1923
Gastrocopta
rixfordi Hanna,
1923; Proc. Calif. Acad. Sci., (4) 13: 515-516; pl. 10, figs. 5-8 (shell).
Gastrocopta
(Immersidens) rixfordi Hanna. Pilsbry,
1926; Man. Conch., 27: 207-208; text-fig. 9 (shell).- Smith, Miller, Christensen & Roth, 1990;
Proc. Calif. Acad. Sci., 47: 106-107; fig. 11 (distribution map).
Gastrocopta
dalliana dalliana
(Sterki). Bequaert & Miller, 1973:
167 (in part; Baja California Sur records).
TYPE LOCALITY.- Isla Monserrat,
Gulf of California, Baja California Sur.
Holotype CAS 1094.
DISTRIBUTION RECORDS.- BAJA
CALIFORNIA SUR: numerous locality records (Smith, et al, 1990).
Subgenus Privatula Sterki, 1893
Privatula Sterki, 193; Nautilus, 6:
101.- Pilsbry, 1948; Land Moll. N. Amer.,
2: 893-894.
TYPE SPECIES.- Odostomia corticaria Say, 1816.
DISTRIBUTION.- Eastern North
America from Maine and Ontario west to Minnesota and south to Florida and
Louisiana; northeastern Mexico.
TAXONOMY.- A single species is
recognized.
Gastrocopta (Privatula) corticaria
(Say, 1816)
Odostomia
corticaria Say,
1816; Nicholson’s American Edition British Encyclopedia, 2: pl. 4, fig. 5
(shell).
Gastrocopta
(Privatula) corticaria (Say). Pilsbry,
1916; Man. Conch., 24: 52-53; pl. 10, figs. 1-4 (shell).- Pilsbry, 1948; Land Moll. N. Amer., 2: 894;
figs. 480: 1-4 (shell).-
Correa-Sandoval, Gutierrez & Reza, 1998: 14.
TYPE LOCALITY.- Philadelphia,
Pennsylvania. Neotype ANSP 64524a
(Pilsbry, 1948).
DISTRIBUTION RECORDS.- SAN LUÍS
POTOSÍ: km 82, on rd. from Río Verde-Cd. Valles, 1420 m alt. (21°53’54” N,
99°35’00” W); km 140 on rd. from Cd. del Maíz-Antigua Morelos, 1320 m alt.
(22°28’02” N, 99°28’25” W); Las Abritas , 5 km E of El Naranjo, 840 m alt.
(22°33’24” N, 99°22’22” W,) (Correa-Sandoval, et al, 1998).
Subgenus Vertigopsis Sterki, 1893
Vertigopsis Sterki, 1893; Nautilus, 6:
101.- Pilsbry, 1948; Land Moll. N.
Amer., 2: 886.
TYPE SPECIES.- Pupa curvidens Gould (= Vertigo pentodon Say, 1821).
DISTRIBUTION.- Eastern North
America west to Alberta, Canada and Arizona, and south to Nicaragua.
TAXONOMY.- The subgenus includes four species. Two occur in the study area.
Gastrocopta
(Vertigopsis) pentodon (Say, 1822)
Vertigo
pentodon Say, 1821;
Jour. Acad. Nat. Sci. Phila., 2: 376.
Gastrocopta (Vertigopsis) pentodon
(Say). Pilsbry, 1916; Man. Conch., 24:
28-31; pl. 3, figs 2, 3, 5-8; pl. 4; pl. 5, figs. 28-41 (shell).- Hinkley, 1920; Nautilus, 34: 39, 48, 52.- Baker, 1930a: 5.- Goodrich & van der Schalie, 1937:
30.- Pilsbry, 1948; Land Moll. N. Amer.,
2: 886-889; figs. 477: 2, 3, 5-8 (shell).-
Pilsbry, 1953; Proc. Acad. Nat. Sci. Phila., 105: 161.- Bequaert & Miller, 1973: 88-90, 157-158.- Pérez & López, 2002: 91-92.- Correa-Sandoval & Rodriguez, 2005: 59.
Bifidaria
curvidens
(Gould). Pilsbry, 1904; Proc. Acad. Nat.
Sci. Phila., 55: 766.
TYPE LOCALITY.- Pennsylvania.
DISRIBUTION RECORDS.- NICARAGUA, Dept. Jinotega; Dept. Leon: Dept. Matagalpa: (Pérez
& López, 2002). GUATEMALA, no
precise locality (Goodrich & van der Schalie, 1937). Dept.
Alta Verapaz: Chama. Dept. Izabal: Jocolo; Quirigua (Hinkley, 1920). CHIHUAHUA: talus of cliffs on Rio Piedras
Verdes, ca. 4.5 km below Pacheco (Pilsbry, 1953). NUEVO LEON: on Rio Maurisco; nr. the Hacienda
Pablillo, above Galeana, 7800 ft. alt. (Pilsbry, 1953); drift of the Rio
Purificación, 24 mi. N of Padillo (Bequaert & Miller, 1973); 0.5 km. W of
El Salto (23°56’53” N, 99°45’59” W) (Correa-Sandoval & Rodriguez,
2005). PUEBLA: Necaxa (Baker, 1930a). SAN LUIS POTOSI: blFLMNHfs N of San Dieguito
(Pilsbry, 1916). TAMAULIPAS: canyon 4
mi. W of Cd. Victoria (Pilsbry, 1904, 1916).
TAXONOMY.- Synonyms that apply to
populations outside of the study area are: Pupa
curvidens Gould, 1841, Pupa cincinnatiensis Judge, 1878, Pupa
floridana Dall, 1885, Pupa montanella
Cocherell, 1889 and Pupa curvidens
var. gracilis Sterki, 1890 (Pilsbry,
1948); and Pupa tappaniana C. B. Adams, 1842 (Bequaert & Mlller, 1973).
Gastrocopta
(Vertigopsis) pilsbryana (Sterki, 1890)
Pupa
pilsbryana Sterki,
1890; Nautilus, 3: 123.
Gastrocopta
pilsbryana
(Sterki). Pilsbry, 1916; Man. Conch.,
24: 36-38; pl. 3, figs. 10, 11, 12 (shell).-
Pilsbry, 1948; Land Moll. N. Amer., 2: 890-891; figs. 477: 10-1, fig.
478 (shell).- Pilsbry, 1953; Proc. Acad.
Nat. Sci. Phila., 105: 161.- Branson,
McCoy & Sisk, 1964; Southwestern Naturalist, 9: 104. Bequaert & Miller,
1973: 158-159.
Gastrocopta
soneri Chamberlin
& Jones, 1929; Bull. Univ. Utah, 19: 83; fig. 32 (shell).
Gastrocopta
pilsbryana amissidens Pilsbry,
1934; Man. Conch., 28: 112; pl. 24, figs. 5, 6 (shell).- Pilsbry, 1948; Land Moll. N. Amer., 2:
891-893; figs. 478 (shell).
TYPE LOCALITIES.- Pupa pilsbryana: from the Colorado
River, Arizona; holotype Carnegie Museum 62.20382. Gastrocopta
stoneri: Cedar City, Utah. Gastrocopta pilsbryana amissidens: San
Francisco Mountains, Arizona; holotype ANSP 161437.
DISRIBUTION RECORDS.- Utah,
Arizona, New Mexico and trans-Pecos Texas south to northern Mexico. CHIHUAHUA:
Sierra la Breña, ca. 17.7 km by rd. from Pearson, on rd. to Pacheco, ca. 7000
ft. alt.; along Rio Piedras Verdes, 4.5 km below Pacheco, 5900 ft. alt.
(Pilsbry, 1953). SONORA: (Branson, et al, 1964). TAMAULIPAS: (Bequaert & Miller, 1973).
Subfamily VERTIGININAE
Stimpson, 1851
TYPE GENUS.- Vertigo Müller, 1774.
TAXONOMY.- A single extant genus
is recognized (Zilch, 1959: 148-149).
Genus Vertigo Müller, 1774
TYPE SPECIES.- Vertigo pusilla Müller, 1774.
DISTRIBUTION.- Practically the
entire Holarctic Realm from sea level to 10,000 feet altitude (Pilsbry, 1948).
TAXONOMY.- Five subgenera are
recognized. Three occur in the study
area.
Subgenus Vertigo Müller, 1774
DISTRIBUTION.- As for the genus.
TAXONOMY.- About 80 species are
recognized. Three species and three
subspecies occur in the study area.
Vertigo
(Vertigo) berryi Pilsbry, 1919
Vertigo
berryi Pilsbry,
1919; Man. Conch., 25: 89; pl. 6, figs. 10, 13 (shell).
Vertigo (Vertigo) berryi Pilsbry. Pilsbry, 1948; Land Moll. N. Amer., 2: 955;
fig. 513: 10, 13 (shell).- Smith,
Miller, Christensen & Roth, 1990; Proc. Calif. Acad. Sci., 47: 108-109;
fig. 13 (map).
TYPE LOCALITY.- Mill Creek
canyon, San Bernardino Mountains, California; 4600 ft. alt. Holotype ANSP 105166.
DISTRIBUTION RECORDS.- California
and Nevada south to Baja California Norte.
BAJA CALIFORNIA NORTE: Valle Trinidad (Pilsbry, 1948).
Vertigo
(Vertigo) gouldi arizonensis Pilsbry & Vanatta, 190o
Vertigo coloradoensis arizonensis Pilsbry & Vanatta, 1900; Proc. Acad. Nat. Sci.
Phila., 52: 601; pl. 23, fig. 9 (shell).
Vertigo (Vertigo) coloradoensis
arizonensis Pilsbry & Vanatta.
Pilsbry, 1919; Man. Conch., 25: 117; pl. 12, figs. 14, 16 (shell).
Vertigo (Vertigo) gouldi arizonensis
Pilsbry & Vanatta. Pilsbry, 1948; Land
Moll. N. Amer., 2: 975-976; fig. 518: 14, 16 (shell).- Bequaert & Miller, 1973: 186-187.
TYPE LOCALITY.- Top of Mingus
Mountain, Yavapai Co., Arizona, 7600 ft. alt.
Holotype ANSP 119010.
DISTRIBUTION RECORDS.- CHIHUAHUA:
Sierra de la Breña, ca. 16 mi. SW of Colonia Juarez (Bequaert & Miller,
1973).
Vertigo
(Vertigo) gouldi coloradoensis (Cockerell, 1891)
Pupa
coloradoensis
Cockerell, 1891; British Naturalist: 100.
Virtigo
coloradoensis
(Cockerell). Sterki, 1892; Nautilus, 6:
5.- Pilsbry, 1919; Man. Conch., 25: 115.
Vertigo (Vertigo) gouldi coloradoensis
(Cockerell). Pilsbry, 1948: Land Moll.
N. Amer., 2: 974; fig. 518: 13; fig. 522 (shell).- Pilsbry, 1953; Proc. Acad. Nat. Sci. Phila.,
105: 164.- Bequaert & Miller, 1973:
184-185.
TYPE LOCALITY.- Near Swift Creek,
Custer Co., Colorado.
DISTRIBUTIN RECORDS.- Colorado,
Utah and Arizona south to northwestern Mexico.
CHIHUAHUA: along the Rio Piedras Verdes, 4.6 km from Pearson, 5900 ft. alt.
(Pilsbry, 1953).
Vertigo
(Vertigo) gouldi inserta Pilsbry, 1919
Vertigo coloradoensis inserta Pilsbry, 1919; Man. Conch., 25; 118; pl. 12, figs. 10, 11
(shell).
Vertigo (Vertigo) gouldi inserta
Pilsbry. Pilsbry, 1948; Land Moll. N.
Amer., 2: 976; fig. 10, 12 (shell).-
Pilsbry, 1953; Proc. Acad. Nat. Sci. Phila., 105: 164.- Bequaert & Miller, 1973: 187.
TYPE LOCALITY.- Bear Wallow,
Santa Catalina Mountains, Pima Co., Arizona.
Holotype ANSP 109559.
DISTRIBUTION RECORDS.- CHIHUAHUA:
Sierra de la Breña, ca. 16 mi. SW of Colonia Juarez (Bequaert & Miller,
1973).
Vertigo
(Vertigo) ovata Say, 1822
Vertigo ovata
Say, 1822 Jour. Acad. Nat. Sci. Phila., 2: 375.
Vertigo
(Vertigo) ovata (Say). Pilsbry, 1919; Man. Conch., 25: 83-88.- Pilsbry, 1948; Land Moll. N. Amer., 2: 953;
figs. 513: 1-3, 4, 7 (shell).- Bequaert
& Miller, 1973: 183.- Smith, Miller,
Christensen & Roth, 1990; Proc. Calif. Acad. Sci., 47: 109; fig. 13 (map).
Pupa
ovata (Say). Von Martens, 1898: 327.
TYPE LOCALITY.- Philadelphia,
Pennsylvania.
DISTRIBUTION RECORDS.- Widespread
in North America and the West Indies.
BAJA CALIFORNIA NORTE: On road to La Playa, 0.3 km SSE of San Jose del
Cabo; Sierra Laguna (Smith, et al,
1990). SONORA: drift of Rio Sonoyta, at
Sonoyta, 5 mi. S of the Arizona border (Bequaert & Miller, 1973). VERACRUZ: Veracruz (Von Martens, 1898;
Pilsbry, 1919, questions this record).
Subgenus Alloptyx Pilsbry, 1953
Alloptyx Pilsbry, 1953; Proc. Acad. Nat.
Sci. Phila., 105: 164.
TYPE SPECIES.- Vertigo hinkleyi Pilsbry, 1921.
DISTRIBUTION.- Arizona and
Chihuahua.
TAXONOMY.- A single species is
recognized.
Vertigo
(Alloptyx) hinkleyi Pilsbry, 1921
Vertigo
hinkleyi Pilsbry,
1921; Man. Conch., 26: 234; pl. 6, figs. 12-16 (shell).- Pilsbry, 1948; Land Moll. N. Amer., 2: 945;
fig. 542: 12-16 (shell).
Vertigo (Alloptyx) hinkleyi
Pilsbry. Pilsbry, 1953; Proc. Acad. Nat.
Sci. Phila., 105: 164-165.
TYPE LOCALITY.- Cave Canyon near
the reservoir, Huachuca Mountains, Arizona.
Holotype ANSP 46263.
DISTRIBUTION RECORDS.- CHIHUAHUA;
Sierra de la Breña on the road to Pacheco, 17.5 km from Pearson, 7000 ft. alt.
(Pilsbry, 1953)
Subgenus Angustula Sterki, 1888
Angustula Sterki, 1888; Proc. U. S. National
Museum, 11: 378.- Pilsbry, 1919; Man.
Conch., 25: 145.- Pilsbry, 1948; Land
Moll. N. Amer., 2: 944.
TYPE SPECIES.- Pupa milium Gould, 1840.
DISTRIBUTION.- Eastern North
America, west to South Dakota, Colorado and Arizona, Jamaica, Hispaniola and
Bermuda.
TAXONOMY.- Two species are recognized.
One occurs in the study area.
Vertigo
(Angustula) milium (A. A. Gould, 1840)
Pupa
milium Gould, 1840;
Boston Journal of Natural History, 3: 402.
Vertigo
milium
(Gould). Binney, 1878; Terr. Moll., 5:
215; pl. 71, fig. 1.- Hinkley, 1907;
Nautilus, 21: 77.- Pilsbry, Man. Conch.,
25: 146-149; pl. 13, figs. 1-7 (shell).-
Pilsbry, 1948; Land Moll. N. Amer., 2: 944-945; figs. 509a-e
(shell).- Pérez & López, 2002:
108-106; pl. 15 (shell); text-fig (distribution map).
TYPE LOCALITY.- Oak Island,
Chelsea, near Boston, Massachusetts.
DISTRIBUTION RECORDS.- Eastern
North America, west to South Dakota, Colorado and Arizona, Jamaica and
Hispaniola (Pilsbry, 1948). NICARAGUA:
“common in the hilly northern central part of the country” (Pérez & López,
2002). TAMAULIPAS: Tampico (Hinkley,
1907).
Genus Nearctula Sterki, 1892
Nearctula Sterki, 1892; Nautilus, 8: 5.- Smith, Miller, Christensen & Roth, 1990;
Proc. Calif. Acad. Sci., 47: 109.
TYPE SPECIES.- Pupa californica Rowell,1861 = Pupa rowelli Newcomb, 1860 (Smith, et al, 1990).
DISTRIBUTION.- San Francisco,
California south to Bahia San Quintin, Baja California Norte.
TAXONOMY.- The genus includes two
species and several subspecies. Two
species and three subspecies occur in the study area.
Nearctula rowelli
diegoensis (Sterki, 1890)
Pupa
californica var. diegoensis Sterki, 1890; Nautilus, 4:
18.
Vertigo
californica diegoensis
(Serki). Pilsbry, 1918; Man. Conch., 25:
141; pl. 9, fig. 8 (shell).- Pilsbry,
1948; Land Moll. N. Amer., 2: 996-998; figs. 533: 8, 11 (shell).
Nearctula
rowelli diegoensis
(Sterki). Smith, Miller, Christensen
& Roth 1990: 109; fig. 13 (map).
TYPE LOCALITY.- False Bay near
Asher Station, San Diego, California.
DISTRIBUTION RECORDS.- BAJA CALIFORNIA
NORTE: San Ramon; Isla San Martin (Pilsbry, 1948); Bahia Todos Santos; nr.
Bahia San Quintin; W shore of Bahia San Quintin; hills W of Bahia San Quintin
(Smith, et al, 1990).
Nearctula rowelli
catalinaria (Sterki)
Vertigo
califronica catalinaria
(Sterki). Pilsbry, 1927; Proc. Calif.
Acad. Sci., (4) 16: 174; pl. 7, fig. 5 (shell).- Pilsbry, 1934; Man. Conch., 28: 97; pl. 15,
fig. 8 (shell).
Nearctula
rowelli catalinaria
(Sterki). Smith, Miller, Christensen
& Roth 1990: 109; fig. 13 (map).
TYPE LOCALITY.- Santa Catalina
Island, Chanel Islands, California.
DISTRIBUTION RECORDS.- California
Chanel Islands south along the coast to northern Baja California Norte. BAJA CALIFORNIA NORTE: Guadalupe Island,
about 1000 ft above the landing at Northeast Anchorage; ca. 2 mi. N of S end of
island on E side; Sierra San Pedro Martir, along road to astronomical
observatory area, 2700 m alt. (Smith, et
al, 1990).
Nearctula rowelli
guadalupensis (Pilsbry, 1927)
Vertigo
californica guadalupensis
Pilsbry, 1927; Proc. Calif. Acad. Sci., (4) 16: 175; pl. 7, fig. 4
(shell).- Pilsbry, 1934; Man. Conch.,
28: 97; pl. 15, fig. 7 (shell).
Nearctula
rowelli guadalupensis
(Pilsbry). Smith, Miller, Christensen
& Roth 1990: 109.
TYPE LOCALITY.- Guadalupe Island,
Baja California Norte. Holotype CAS
2582.
DISTRIBUTION RECORDS.- BAJA
CALIFORNIA NORTE: known only from the type locality.
Nearctula degeneris
(Pilsbry, 1927)
Vertigo
degeneris Pilsbry,
1927; Proc. Calif. Acad. Sci., (4) 16: 175; pl. 7, fig. 6 (shell).- Pilsbry, 1934; Man. Conch., 28: 98; pl. 15,
fig. 9 (shell).
Nearctula
degerneris
(Pilsbry). Smith, Miller, Christensen
& Roth 1990: 110.
TYPE LOCALITY.- About 1000 feet
above the landing at
DISTRIBUTION RECORDS.- BAJA
INDEX | VETIGASTROPODA | PULMONTA (LYMNAEOIDEA-PUPILLOIDEA) | PULMONATA (ORTHALICOIDEA) |
PULMONATA (ACHATINOIDEA-SAGDOIDEA) | PULMONATA (GASTRODONTOIDEA-HELICOIDEA) | BIBLIOGRAPHY