INDEX | VETIGASTROPODA | PULMONTA (LYMNAEOIDEA-PUPILLOIDEA) | PULMONATA (ORTHALICOIDEA) |
PULMONATA (ACHATINOIDEA-SAGDOIDEA) | PULMONATA (GASTRODONTOIDEA-HELICOIDEA) | BIBLIOGRAPHY


PART 3 (ORTHALICOIDEA)

 

Superfamily ORTHALICOIDEA

 

Family ORTHALICIDAE Albers, 1860

 

Subfamily ORTHALICINAE Albers, 1960

 

  DISTRIBUTION.-   Tropical South America, the West Indies, south Florida, Central America and Mexico.

 

  TAXONOMY.-  The nomenclature employed in this study follows Breure & Schouten (1985).   Six genera are recognized in the subfamily Orthalicinae.  One occurs in Central America and Mexico.

 

 

Genus Orthalicus Beck, 1837

 

Orthalicus Beck, 1837.  Von Martens, 1893; Biol. Cent. Amer.: 179-182.-  Rehder, 1945; Nautilus, 59: 29-31.-  Richardson, 1993: 97-117.

Oxystyla Schluter, 1838.-  Pilsbry, 1899; Man. Conch., 12: 101-105. 

Zebra Shuttleworth, 1858.

 

  TYPE SPECIES.-  Buccinum zebra Müller, 1774 (Rehder, 1945).

 

  DISTRIBUTION.-  From Bolivia and Brazil north to Mexico, the West Indies and south Florida.  Within the Central American-Mexican region generally the genus occurs below 1500 m altitude.  In Mexico Orthalicus occurs at lower elevations along the east coast from southeastern Tamaulipas south to Isthmus of Tehuantepec, northern Chiapas, the Yucatan Peninsula and the Petén region of Guatemala.  On the west coast the genus occurs from central Sinaloa southeast to Chiapas and Guatemala, but it occurs at higher elevations than on the east coast.  Orthalicus continues south through Central America, generally at less than 1000 m altitude.

 

  TAXONOMY.-  Strebel (1909) monographed the Orthalicinae.  He described numerous species and subspecies, many of which were based on undocumented specimens.  In several instances he ignored nomeclatural priorities, he used the same subspecific name for variations within different species, and he employed unconventional taxonomic nomenclature.  In many instances he divided single specimen lots into varieties, subspecies and even species.  In other instances he placed under a given species name superficially similar appearing specimens from very widely separated localities.  These actions cause many taxonomic problems, and a major review of the genus is needed. Tentatively, twenty-seven species plus twelve of Orthalicus are recognized in this study.  Richardson (1993) provides a list of references for the species.  

 

 

Orthalicus boucardi Pfeiffer, 1860

 

Orthalicus boucardi Pfeiffer, 1860; Proc. Zool. Soc. London: 138; pl. 51, fig. 7.-  Fischer & Crosse, 1875; Miss. Sci. Mex.: 451; pl. 18, figs. 3, 3a, 3b.-  Strebel, 1882: 40-41; pl. 1, fig. 1, 2a-g.-  Von Martens, 1893; Biol. Cent. Amer.: 187; pl. 11, figs. 4, 5, 11.-  Richardson, 1993: 98-99. 

Orthalicus boucardi-ponderosus Strebel, 1882: 39; pl. 1, figs. 4a, 4b (shell).  (TL: Tehuantepec).

Orthalicus ponderosus-boucardi Strebel, 1882; 39; pl. 1, figs. 3a, 3b.  (TL: Tehuantepec).

Oxystyla longa var. boucardi (Pfeiffer).  Pilsbry, 1899; Man. Conch., 12: 127-128; pl. 20, figs. 20-26, 29 (shell). 

Oxystyla longa var. strebeli Pilsbry, 1899; Man. Conch., 12: 128; pl. 22, figs. 7-9.

Zebra boucardi (Pfeiffer).  Strebel, 1909; Rev. Unterfam. Orthalicinen: 72-76; pl. 14, figs. 210, 216, 219, 223 (shell).

Zebra boucardi heteromorph melanocheila Strebel, 1909; Rev. Unterfam. Orthalicinen: 74; pl. 14, figs. 217, 218.

Zebra boucardi heterom. xanthus Strebel, 1909; Rev. Unterfam. Orthalicinen: 74.

Zebra boucardi heterom. albinus Strebel, 1909; Rev. Unterfam. Orthalicinen: 74.  (TL: Cerro Negro, near Santa Efigenia, Tehuantepec, Oaxaca).

Zebra boucardi form adustus Strebel, 1909; Rev. Unterfam. Orthalicinen: 74-74; pl. 14, fig. 228.

Zebra boucardi form zebriola Strebel, 1909; Rev. Unterfam. Orthalicinen: 75; pl. 14, figs. 229, 230.

Zebra boucardi form zoniferus Strebel, 1909; Rev. Unterfam. Orthalicinen: 75; pl. 14, figs. 224, 225.  (TL: Tehuantepec).

 

  TYPE LOCALITIES.-  As notes above with TL.

 

  DISTRIBUTION RECORDS.-  MICHOACÁN: no specific locality (Strebel, 109).  OAXACA: Villa Alta; Tehuantepec; Cerro Negro, Tehuantepec; Sierra de Betaza (17.25º N, 96.15º W) (Von Martens, 1893).

 

 

Orthalicus decolor Strebel,1882

 

Orthalicus decolor Strebel, 1882: 37; pl. 7, figs. 2-4.-  Richardson, 1993: 100.

Oxystyla decolor (Strebel).  Pilsbry, 1899; Man. Conch., 12: 131-132; pl. 54, figs. 43-47.

Zebra mars form decolor (Strebel).  Strebel, 1909; Rev. Unterfam. Orthalicinen: 62; pl. 11, figs. 167.

 

  TYPE LOCALITY.-  Not stated.

 

  DISTRIBUTION RECORDS.-  Unknown.

 

  REMARKS.-  It is doubtful if this taxon is part of the Mexico-Central American fauna.  Strebel (1909) notes its similarity to Orthalicus mars (Pfeiffer, 1861), an Ecuadorian species

 

 

Orthalicus delphinus (Strebel, 1909)

 

Zebra delphinus Strebel, 1909; Rev. Unterfam. Orthalicinen: 30-31.

Orthalicus delphinus (Strebel).  Jacobson, 1958; Amer. Mus. Nov., (1899): 11-12.-  Richardson, 1993: 100-101.

Zebra delphinus form nebulosus Strebel, 1909; Rev. Unterfam. Orthalicinen: 31; pl. 2, fig. 47 (shell).

Oxystyla delphinus form nebulosa (Strebel).  Dall, 1926; Proc. Calif. Acad. Sci., (4) 15: 473-474.

Zebra delphinus form pumilio Strebel, 1909; Rev. Unterfam. Orthalicinen: 33-34.

Oxystyla delphinus form nesiotica Dall, 1926; Proc. Calif. Acad. Sci., (4) 15: 474-475; pl. 35, fig. 3 (shell).

 

  TYPE LOCALITIES.-  Zebra delphinus: not given.  Zebra delphinus form nebulosus: Mazatlan, Sinaloa.  Zebra dephinus form pumilio: Mazatlan, Sinaloa.  Oxystyla delphinus form nesiotica: Maria Madre Island, Tres Marias Islands, Nayarit.  Holotype CAS 2194.

 

  DISTRIBUTION RECORDS.-  NAYARIT: Maria Magdalena Island (Dall, 1926; Jacobson, 1958).  SINALOA: Mazatlan (Strebel, 1909).

 

 

 

Orthalicus elegans Rolle, 1895

 

Orthalicus elegans Rolle, 1895; Nachr. Deutsch. Malak. Gesell., 32: 131.-  Neubert & Jannsen, 2004; Arch. für Moll., 133: 236; pl. 23, fig. 280 (shell).

Orthalicus princeps elegans Rolle.  Von Martens, 1901; Biol., Cent. Amer.: 629; pl. 44, fig. 12.-  Richardson, 1993: 110.

Oxystyla princeps var. elegans (Rolle).  Pilsbry, 1899; Man. Conch., 12: 117-18.-  Pilsbry, 1902; Man. Conch., 14: 164; pl. 24, fig. 12 (shell).

Zebra elegans (Rolle). Strebel, 1909; Rev. Unterfam. Orthalicinen: 38-39; pl. 4, figs. 66, 67. 

Orth.[alicus] colimensis Rolle, 1895; Nachr. Deutsch. Malak. Gesell., 32: 131.

 

  TYPE LOCALITY.-  Colima.  Holotype in the ZMB. 

 

  DISTRIBUTION RECORDS.-  COLIMA: not known from a definite locality.

 

 

 

Orthalicus ferussaci ferussaci Von Martens, 1863

 

Orthalicus ferussaci Von Martens, 1963; Monatsber. Akad. Wissen. Berlin, 1863: 542.-  Fischer & Crosse, 1875; Miss. Sci. Mex.: 447.-  Von Martens, 1893; Biol. Cent. Amer.: 184; pl. 10, figs. 8-10 (shell).-  Breure & Schouten, 1985; Zool. Meded. Uitg. d. Rijksmus. Nat. Hist. Leiden, 51: 29; text-fig. 16 (reproductive anatomy).-  Richardson, 1993: 101.-  Pérez & Lopéz, 2002: 223-224.

Oxystyla ferussaci (Von Martens).  Pilsbry, 1899; Man. Conch., 12: 119-120; pl. 17, figs. 13-17 (shell).

 

  TYPE LOCALITY.-  Tehuantepec, Oaxaca.

 

  DISTRIBUTION RECORDS.-  COSTA RICA, Guanacaste Prov.: Guanacaste; Salinas Bay.  GUATEMALA, Dept. Alta Verapaz: Panzos; San Geronimo.  Dept. Zacapa: Zacapa.  NICARAGUA, Dept. Chontales: Acoyapa (Pilsbry, 1899).  MICHOACÁN: El Espinal (Breure & Schouten, 1985).  OAXACA: Tehuantepec.  YUCATAN: Shkolak (Pilsbry, 1899).

 

 

 

Orthalicus ferussaci tricinctus Von Martens, 1893

 

Orthalicus melanocheilus (Valenciennes).  Fischer & Crosse, 1875; Miss. Sci. Mex.: 458 (in part); pl. 18, figs. 5, 5a (shell).

Orthalicus ferussaci var. tricinctus Von Martens, 1893; Biol. Cent. Amer.: 185; pl. 11, fig. 8 (shell).

Oxystyla ferussaci var. tricincta (Von Martens). Pilsbry, 1899; Man. Conch., 12: 120-121; pl. 18, figs. 4-6; pl. 17, fig. 18 (shell).

Orthalicus maracaibensis tricinctus Von Martens.  Richardson, 1993: 106.

 

  TYPE LOCALITY.-  Not specified.

 

  DISTRIBUTION RECORDS.-  COSTA RICA, Guanacaste Prov.: Quebrada Vijigual, Rio Saverey Valley, 150 m alt.; El Pozo, 50 m alt.   Puntarenas Prov.: Boruco, 450 m alt.; Terraba, 250-270 m alt.; Alto de Mano Tigre, 690 m alt.  NICARAGUA: no specific locality (Pilsbry, 1899).

 

 

 

Orthalicus hackeri (Strebel, 1909)

 

Zebra hackeri Strebel, 1909; Rev. Unterfam. Orthalicinen: 50; pl. 7, figs. 104, 107, 109, 110, 111 (shell).

Orthalicus hackeri (Strebel).  Richardson, 1993: 103.

 

  TYPE LOCALITY.-  Tepic, Nayarit.

 

  DISTRIBUTION RECORDS.-  NAYARIT: known only from the type locality.

 

 

 

Orthalicus leucocheilus Crosse & Fischer, 1869

 

Orthalicus leucocheilus Crosse & Fischer, 1869; Journ. De Conchyl., 17: 423.-  Fischer & Crosse, 1875; Miss. Sci. Mex.: 459; pl. 18, figs. 7, 7a (shell).- Richardon, 1993: 103.

Oxystyla leucocheila (Fischer & Crosse).  Pilsbry, 1899; Man. Conch., 12: 129-130; pl. 21, figs. 37-39 (shell).

 

  TYPE LOCALITY.-  Orizaba, Veracruz.

 

  DISTRIBUTION RECORDS.-  VERACRUZ: known only from the type locality.

 

 

 

 

Orthalicus livens Shuttleworth 1856

 

Orthalicus livens Shuttleworth 1856; Notitiae Malacologicae, 1: 64; pl. 3, fig. 8.-  Fischer & Crosse, 1872: 453, 455; pl. 18, figs. 6, 6a (shell).-  Von Martens, 1893; Biol. Cent. Amer.: 189.-  Richardson, 1993: 103.-  Neubert & Gosteli, 2005; Contr. Nat. Hist., 5: 23; pl. 20, fig. 2.

Oxystyla livens (Shuttleworth).  Pilsbry, 1899; Man. Conch., 12: 118-119; pl. 23, fig. 15 (shell).

Zebra livens (Shuttleworth).  Strebel, 1909; Rev. Unterfam. Orthalicinen: 42-43; pl. 5, figs. 73-73 (shell).

 

  TYPE LOCALITY.-  Mexico, probably near Veracruz.  Syntypes: MHNN/2.

 

  DISTRIBUTION RECORDS.-  GUATEMALA, Dept. Alta Verapaz: Coban (Strebel, 1909).  VERACRUZ: not known from a definite locality.

 

  REMARKS.-  Beck, 1837, has been credited as the author of the name livens.  He only listed the name.  Shuttleworth first described the species (Neubert & Gosteli, 2005). 

 

 

 

Orthalicus lividus Von Martens, 1863

 

Orthalicus lividus Von Martens, 1863; Monatsber. Akad. Wiss. Berlin, 1863: 542.-  Fischer & Crosse, 1875; Miss. Sci. Mex.: 448.-  Strebel, 1882: 29-31; pl. 1, fig. 8; pl. 4, fig. 6; pl. 11, fig. 18 (shell).-  Von Martens, 1893; Biol. Cent. Amer.: 186; pl. 10, figs. 11, 11a (shell).-  Solem, 1959; Occ. Pap. Mus. Zool. Univ. Mich., (611): 8.

Oxstyla livida (Von Martens).  Pilsbry, 1899; Man. Conch., 12: 124-125; pl. 19, fig. 18, 19 (shell).

Zebra lividus (Von Martens) Strebel; 1909: 48-49; pl. 7, figs. 101-106 (shell).

 

  TYPE LOCALITY.-  Volcán Jorullo, Michoacán.

 

  DISTRIBUTION RECORDS.-  MICHOACÁN: N slope of Volcán Jorullo, 2700-4200 ft. alt. (Solem, 1959).  SINALOA: Mazatlan (Pilsbry, 1899).

 

 

 

Orthalicus longus Pfeiffer, 1856

 

Orthalicus longus Pfeiffer, 1856; Malak. Blätter, 12: 39.-  Fischer & Crosse, 1875; Miss. Sci. Mex.: 450-451; pl. 18, fig. 4 (shell).-  Strebel, 1882: 43; pl. 6, figs. 1b, 1c (shell).-  Von Martens, 1893; Biol. Cent. Amer.: 189.-  Solem, 1959; Occ. Pap. Mus. Zool. Univ. Mich., (611): 6.

Oxystyla longa (Pfeiffer).  Pilsbry, 1899; Manual of Conchology, (2) 12: 126-129; pl. 20, figs. 20-26, 29; pl. 21, figs. 33-36; pl. 22; pl. 23, figs. 16-18 (shell).

Zebra longa (Pfeiffer).  Strebel, 1909; Rev. Unterfam. Orthalicinen: 68-69; pl. 13, figs. 190, 192 (shell).

 

  TYPE LOCALITY.-  Unknown.

 

  DISRIBUTION RECORDS.-  MICHOACÁN: Ario.  MORELOS: Cuernavaca.  OAXACA: Santa [Salina] Cruz Bay; San Geronimo; Tehuantepec; 10 mi. W of Tehuantepec; 20 mi. W of Tehuantepec.  Solem (1959) lists San Geronimo, of which there are two such places in Oaxaca.

 

 

 

Orthalicus maclurae Von Martens, 1893

 

Orthalicus maclurae Von Martens, 1893; Biol. Cent. Amer.: 188; pl. 11, figs. 1-3 (shell).

Oxystyla maclurae (Von Martens).  Pilsbry, 1899; Man. Conch., 12: 125; pl. 21, figs. 40-42 (shell).

Zebra maclurae (Von Martens).  Strebel, 1909; Rev. Unterfam. Orthalicinen: 69-71; pl. 13, figs. 193, 203.

Zebra maclurae form turrita Strebel, 1909; Rev. Unterfam. Orthalicinen: 71; pl. 13, figs. 204, 205.

 

  TYPE LOCALITIES.-  Ortalicus maclurae: Cacao, Golfo de Fonseca, Nicaragua [Cacao, Dept. Choluteca, Honduras ?].  Zebra maclurae form turrita: unknown.

 

  DISTRIBUTION RECORDS.-  HONDURAS: known only from the type locality.  COLIMA (Strebel, 1909).  Nayarit: Tres Marias Islands (Strebel, 1909).  OAXACA: Tehuantepec (Strebel, 1909).  TRINIDAD (Strebel, 1909).

 

  REMARKS.-  See REMARKS under Orthalicus quagus.

 

 

 

Orthalicus maculiferus (Strebel, 1909)

 

Zebra maculiferus Strebel, 1909; Rev. Unterfam. Orthalicinen: 30; pl. 3, fig. 38 (shell).

Orthalicus maculiferus (Strebel).   Richardson, 193: 105.

 

  TYPE LOCALITY.-  Not stated.

 

  DISTRIBUTION.- Unknown.

 

 

 

Orthalicus melanocheilus melanocheilus (Valenciennes, 1833)

 

Bulimus melanocheilus Valenciennes, 1833: 246; pl. 55, figs. 3a, 3b.

Orthalicus melanocheilus (Valenciennes).  Von Martens, 1865; Malak. Blätt., 12: 45, 70.-  Von Martens, 1893; Biol. Cent. Amer.: 181, 190; pl. 11, figs. 6, 7, 7a.-  Rolle, 1895: 130.-  Breure & Schouten, 1985; Zool. Meded. Uitg. d. Rijksmus. Nat. Hist. Leiden, 51: 32; text-fig. 18 (reproductive anatomy).

Oxystyla melanocheilus (Valenciennes).  Pilsbry, 1899; Man. Conch., 12: 122-123; pl. 18, figs. 1-3.-  McGinty, 1939; Nautilus, 53: 6.

Orthalicus zebra (Müller).  Binney & Bland, 1869; Land & Freshwater shells N. Amer.: 217; text-fig. 371, text-fig. 367 (jaw).

 

  TYPE LOCALITY.-  Arbitrarily restricted to northwestern Mexico by Von Martens (1865).

 

  DISTRIBUTION RECORDS.-  COLIMA: no specific locality (Rolle, 1895).  GUERRERO: 17 mi. SW of Acapulco (McGinty, 1939); 19.3 mi. W of San Marcos, 300 ft. alt. (Breure & Schouten, 1985).  NAYARIT: Tres Maries Islands (Pilsbry, 1899).  SINALOA: Mazatlan (Pilsbry, 1899).

 

 

 

Orthalicus melanocheilus mariae (McGinty, 1939)

 

Oxystla melanocheilus mariae McGinty, 1939; Nautilus, 53: 6; pl. 2, fig. 6 (shell).

 

  TYPE LOCALITY.-  Puerto Marquez, near Acapulco, Gurrero.  Holotype ANSP 174048.

 

  DISTRIBUTION RECORDS.-  GUERRERO: known only from the type locality.

 

 

 

Orthalicus muelleri (Strebel, 1909)

 

Orthalicus zebra (Müller).  Strebel, 1882: 24; pl. 11, fig. 20 (shell).

Zebra muelleri Strebel, 1909; Rev. Unterfam. Orthalicinen: 46.

Orthalicus muelleri (Strebel).  Richardson, 1993: 107.

 

  TYPE LOCALITY.-  Mazatlan, Sinaloa [Mazatlan, Guerrero ?].

 

  DISTRIBUTION RECORDS.-  SINALOA [GUERRERO ?]: known only from the type locality.

 

 

 

Orthalicus nobilis Rolle, 1895

 

Orth.[alicus] nobilis Rolle, 1895; Nachr. Deut. malak. Gesell., 27: 131.-  Richardson, 1993: 107.

Orthalicus zoniferus var. nobilis Rolle.  Von Martens, 1901; Biol. Cent. Amer.: pl. 46, fig. 16 (shell).

Oxystyla nobilis (Rolle).  Pilsbry, 1899; Man. Conch., 12: 124.

Oxystyla zoniferus var. nobilis (Rolle).  Pilsbry, 1902; Man. Conch., 14: pl. 24, fig. 13 (shell).

Zebra nobilis (Role).  Strebel, 1909; Rev. Unterfam. Orthalicinen: 39-40; pl. 5, figs. 68, 69 (shell).

 

  TYPE LOCALITY.-  Colima.

 

  DISTRIBUTION RECORDS.-  COLIMA: not known from a specific locality.  

 

 

 

Orthalicus pallidus (Srebel, 1909)

 

Zebra pallida Strebel, 1909; Rev. Unterfam. Orthalicinen: 40; pl. 5, figs. 70-71.-  Neubert & Jannsen, 2004; Arch. für Moll., 133: 237; pl. 23, fig. 281 (holotype).

Orthalicus nobilis pallidus (Strebel).  Richardson, 1993: 107.

 

  TYPE LOCALITY.-  Colima.  Holotype SMF 26849.

 

  DISTRIBUTION RECORDS.—  COLIMA: known only from the type locality.

 

 

 

Orthalicus ponderosus ponderosus Strebel, 1882

 

Orthalicus ponderosus Strebel, 1882: 35-36; pl. 7, figs. 1, 5-8.-  Von Martens, 1893; Biol. Cent. Amer.: 190; pl. 11, figs. 10, 10a.-  Rolle, 1895; Nachr. Deut. Malak. Gesell., 32:: 130.-  Solem, 1959; Occ. Pap. Mus. Zool. Univ. Mich., (611): 7.-  Richardson, 1993:108-109.

Oxystyla ponderosa (Strebel).  Pilsbry, 1899; Man. Conch., 12: 130-131; pl. 19, figs. 14-17.-  McGinty, 1939: Nautilus, 53: 4-5; pl. 2, fig. 1 (shell).

Orthalicus boucardi-ponderosus Strebel, 1882; 39; pl. 1, figs. 4a, 4b.

Orthalicus ponderosus-boucardi Strebel, 1882: 132.

Orthalicus lividus-princeps Strebel, 1882: 31; pl. 1, fig. 6.

Zebra mars (Pfeiffer). Strebel, 1909; Rev. Unterfam. Orthalicinen: 59-61; pl. 11, figs. 170-172.

Zebra miles Strebel, 1909; Rev. Unterfam. Orthalicinen: 64-65; pl. 12, figs. 183-184. (TL: 25 miles NE of Acapulco, Guerrero).

 

  TYPE LOCALITY.-  Not stated.

 

  DISRIBUTION RECORDS.-  COLIMA: Culata, nr. Manzanillo; Colima. GUERRERO: Dos Arroyos, 25 mi. NE of Acapulco, 1000 ft. alt. (Pilsbry, 1899); vicinity of Acapulco (McGinty, 1939).  MICHOACÁN: Sierra de Coalcoman, trail from Pomare to Maruata, 700 ft. alt.; Sierra de Coalcoman, trail from Los Pozos to Ranch Quemado, 4500 ft. alt. (Solem, 1959).  NAYARIT: San Blas; Tepic.  OAXACA: Tehuantepec (Pilsbry, 1899).  

 

  REMARKS.-  Apparently Strebel took the species name Zebra miles from the type locality of “25 miles NE of Acapulco, Guerrero”.

 

 

 

Orthalicus ponderosus albatus (McGinty, 1939)

 

Oxystyla ponderosa albata McGinty, 1939; Nautilus, 53: 5-6; pl. 2, fig. 4 (shell).-  Richardson, 1993: 97.

 

  TYPE LOCALITY.-  Vicinity of Acapulco, Guerrero.  Holotype ANSP 174049.

 

  DISTRIBUTION RECORDS.-  GUERRERO: known only from the type locality.

 

 

 

Ortalicus ponderosus balesi (McGuinty, 1939)

 

Oxystyla ponderosa balesi McGinty, 1939; Nautilus, 53: 5; pl. 2, fig. 2 (shell).-  Richardson, 1993: 97.

 

  TYPE LOCALITY.-  Puerto Marquez, near Acapulco.  Holotype ANSP 174046.

 

  DISTRIBUTION RECORDS.-  GUERRERO: known only from the type locality.

 

 

 

Orthalicus princeps princeps (Broderip, 1833)

 

Bulimus princeps Broderip, 1833.

Oxystyla princeps (Broderip).  Pilsbry, 1899; Man. Conch., 12: 113-115; pl. 16, figs. 1-9.-  Pilsbry, 1926; Proc. Aacd. Nat. Sci. Phila., 78: 87-88; fig. 18 (shell).-  Pilsbry, 1930; Proc. Aacd. Nat. Sci. Phila., 82: 341-342; pl. 28, figs. 1-9.-  Richards, 1937 253.-  Richards, 1939; Proc. Amer. Philos. Soc., 81: 33.-  Basch, 1959; Occ. Pap. Mus. Zool. Univ. Mich., (612): 10.

Orthalicus princeps (Broderip).  Von Martens, 1893; Biol. Cent. Amer.: 182-183; pl. 10, figs. 3, 3a, 3b, 4-6 (shell).-  van der Schalie, 1940: 5.-  Solem, 1959; Occ. Pap. Mus. Zool. Univ. Mich., (611): 7.-  Rehder, 1966; Proc. Biol. Soc. Wash., 79: 287-288.-  Thompson, 1967; Bull. Fla. St. Mus., 11: 244.-  Breure & Schouten, 1985; Zool. Meded. Uitg. d. Rijksmus. Nat. Hist. Leiden, 51: 32; text-fig. 18 (reproductive anatomy).-  Richardson, 1993: 109-111.-  Pérez & Lopéz, 2002: 225-227.- 

O.[rthalicus] princeps var. crassiuscula Strebel, 1909; Rev. Unterfam. Orthalicinen: 23.

 

  TYPE LOCALITY.-  Bulimus princeps: Conchagua, Dept. La Union, El Salvador. 

 

  DISTRIBUTION RECORDS.-  Widely distributed from Sinaloa along the west coast and from southeastern Tamaulipas along the east coast of Mexico south to Panama.  COSTA RICA, Guanacaste Prov.: Guanacaste (Pilsbry, 1899).  GUATEMALA, Dept. Alta Verapaz: Cahabon; Coban; Senahu; Panzos (Pilsbry, 1899); Chamá (van der Schalie, 1940).  Dept. Peten: Tikal National Park (Basch, 1959).  Dept. Progreso: San Agustin (Pilsbry, 1899).  Dept. Quetzaltenango: Cerro Zunil (Pilsbry, 1899).  Dept. Retalhuleu: Retalhuleu; San Francisco Miramar; Capetillo; El Reposo (Pilsbry, 1899).  Dept. Suchitepequez:  Mazatenango (Pilsbry, 1899).  NICARAGUA: Corn Island (Richards, 1939); commonly distributed along the Pacific Versant (Pérez & Lopéz, 2002).  PANAMA, Canal Zone: Barro Colorado Island; between Tabernillo and San Pablo.  Chiriqui Prov.: Bugaba; Quipo (Pilsbry, 1926).  EL SALVADOR, Dept. Union: Conchagua (Pilsbry, 1899).  CAMPECHE: several localities (Thompson, 1967).  CHIAPAS: Monte Libano, 600 m alt.; Monte Libano to El Censo, 600-700 m alt.; San Lorenzo, midway between Ocosingo and El Real (Bequaert, 1957);  Escuintla (Solem, 1959).  COLIMA: Colima.  NAYARIT: Tepic (Pilsbry, 1899); Tres Marias Islands (Dall, 1926) (this record may be based on misidentifications of Orthalicus delphinus (Jacobson, 1958: 12). OAXACA: Panistlahuaca; Ixstapa; Cerro de Acatepec, near Tuletepec (Pilsbry, 1899); 3.4 mi. WNW Zanatepec, ca. 275 ft. alt. (Breure & Schouten, 1985).  QUINTANA ROO: Cozumel Island (Richards, 1937; Rehder, 1966).  SAN LUIS POTOSI: Agua Buena (21°57’33” N, 99°22’37” W, 440 m alt.); Las Cascadas, Tamasopa (21°56’05” N, 99°25’00” W); Jalpilla (21°23’39” N, 98°52’37” W)  (Correa, et al 1998).  SINALOA: Mazatlan; Presidio de Mazatlan (Pilsbry, 1899). TAMAULIPAS: localities in the southeastern part of the state. (Correa & Catstro, 2002).  VERACRUZ: Papantla; Misantl; Playa Vicente; Coatepec, above Jalapa; Almolonga, below Jalapa; Mirador; Cordoba; Atoyac; Veracruz (Pilsbry, 1899), Rancho El Sol, Naranjos (21°20’00” N, 97°43’16” W); San Juan Cuajinampa (21°11’53” N, 97°30’00” W); El Bajío, Carr. Naranjos-Tuxpan (20°57’17”N, 97°25’57” W); Carr. Tuxpan-Poza Rica, km 234 (20°49’11” N, 97°30’00” W); La Ordeña, Papantla (20°29’43”N, 97°18’27” W) (Correa, 1999).  YUCATAN: Merida; Tabi; Shkolak; Tekanto; Ticul; Silam (Pilsbry, 1899); Chichen Itza (Bequaert & Clench, 1936).

 

 

Orthalicus princeps crossei Von Martens, 1893

 

Orthalicus zoniferus crossei Von Martens 1893; Biol. Cent. Amer.: 186.-  Haas & Solem, 1960; Nautilus, 73: 131.

Oxystyla princeps crossei (Von Martens).  Pilsbry, 1899; Man. Conch., 12: 116; pl. 17, fig. 12. -  Richardson, 1993: 110.

 

  TYPE LOCALITY.-  Belize.

 

  DISTRIBUTION RECORDS.-  BELIZE: Rio Frio Cave, Double Fall (Haas & Solem, 1960).

 

 

 

Orthalicus princeps deceptor (Pilsbry, 1899)

 

Orthalicus obductus Shuttleworth.  Fischer & Crosse; 1875: 456 (exclusive of South American references and localities).-  Von Martens, 1893; Biol. Cent. Amer.: 187 (exclusive of South American references and localities).

Zebra obductus (Shuttleworth).  Strebel, 1909; Rev. Unterfam. Orthalicinen: 34; pl. 3, figs. 4a, 4b, 7b, 7c (shell) (exclusive of South American references and localities).

Oxystyla princeps var. deceptor Pilsbry, 1899; Man. Conch., 12: 116-117; pl. 24, figs. 19-24 (shell).-  Pilsbry, 1926; Proc. Aacd. Nat. Sci. Phila., 78: 341.

Orthalicus princeps deceptor (Pilsbry).  Richardson, 1993: 110.

 

  TYPE LOCALITY. Polvón, Dept. León, Nicaragua.  Lctotypa ANSP 5099a (H. B. Baker, 1963: 227).

 

  DISTRIBUTION RECORDS.-  BELIZE: Belize.  GUATEMALA, Dept. Alta Verapaz: Coban; San Geronimo.  NICARAGUA, Dept. Leon: Polvón. PANAMA: islands in Panama Bay. 

 

 

 

Orthalicus princeps fischeri Von Martens, 1893

 

Orthalicus princeps var. fischeri Von Martens, 1893; Biol. Cent. Amer.: 183; pl. 10, fig. 7 (shell). 

Oxystyla princeps fischeri (Von Martens).  Pilsbry, 1899; Man. Conch., 12: 116; pl. 16, fig. 9 (shell).-  Pilsbry, 1930; Proc. Aacd. Nat. Sci. Phila., 82: 342.

Orthalicus princeps fischeri (Von Martens).  Richardson, 1993: 110.

 

  TYPE LOCALITY.-  El Reposo and San Isidro, Dept. Suchitepequez, Guatemala.

 

  DISTRIBUTION RECORDS.-  GUATEMALA: known only from the type locality.

 

 

 

Orthalicus princeps perlongus (Pilsbry, 1899)

 

Oxystyla princeps var. perlonga Pilsbry, 1899; Man. Conch., 12: 117; pl. 24, fig. 25.

Orthalicus princeps perlongus (Pilsbry).  Richardson, 1993: 110.

 

  TYPE LOCALITY.  Panama. 

 

  DISTRIBUTION RECORDS.-  PANAMA: not known from a definite locality.

 

 

 

Orthalicus princeps trifractus (Pilsbry, 1899)

 

Oxystyla princeps var. trifracta Pilsbry, 1899; Man. Conch., 12: 115; pl. 17, figs. 10, 11 (shell).

 

  TYPE LOCALITY.-  El Salvador.  Lectotype ANSP 28080 (H. Baker, 1963: 231).

 

  DISTRIBUTION RECORDS.-  EL SALVADOR: known only from the type locality.

 

 

Orthalicus pseudolongus (Strebel, 1909)

 

Zebra pseudo-longus Strebel, 1909; Rev. Unterfam. Orthalicinen: 71-72; pl. 14, figs. 208, 209 (shell).

 

  TYPE LOCALITY.  Unknown.

 

  DISTRIBUTION.-  Unknown

 

 

 

Orthalicus quagus (Strebel, 1909)

 

Zebra quaga Strebel, 1909; Rev. Unterfam. Orthalicinen: 41; pl. 6, figs. 92-94, 96 (shell).

Zebra quaga form turrita Strebel, 1909; Rev. Unterfam. Orthalicinen: 42; pl. 5, fig. 78 (shell).

Zebra livens form aberrans Strebel, 1909; Rev. Unterfam. Orthalicinen: 43-44; pl. 5, figs. 74-77 (shell), figs. 70-80 (shell).-  Neubert & Jannsen, 2004; Arch. für Moll., 133: 236; pl. 23, fig. 282 (syntype).

Orthalicus quaga (Strebel).  Solem, 1959; Occ. Pap. Mus. Zool. Univ. Mich., (611): 8; pl. 2, fig. 19 (shell).-  Richardson, 1993: 112.

 

  TYPE LOCALITIES.-  Zebra quaga: not given.  Zebra quaga form turreta: Colima.  Zebra livens form aberrans: Colima; syntype SMF 25454 (Neubert & Jannsen, 2004).

 

  DISTRIBUTION RECORDS.-  COLIMA: (Strebel).  MICHOACÁN: Cerro de Guzman, nr. Coalcoman (Solem, 1959).

 

  REMARKS.-  The name Zebra quaga form turrits (Strebel, 1909; Rev. Unterfam. Orthalicinen: 42) is a different taxon than Zebra maclurae form turrita (Strebel, 1909; Rev. Unterfam. Orthalicinen: 71) or Zebra sphinx form turrita (Strebel, 109: 68).

 

 

 

Orthalicus richardsoni (Strebel, 1909)

 

Zebra richardsoni Strebel, 1909; Rev. Unterfam. Orthalicinen: 36-37; pl. 4, figs. 60, 61, 62 (shell).

Orthalicus richardsoni (Strebel).  Richardson, 1993: 113.

 

  TYPE LOCALITY.-  Tepic, Nayarit.

 

  DISTRIBUTION RECORDS.-  NAYARIT: known only from the type locality.

 

 

 

Orthalicus selectus (Strebel, 1909)

 

Zebra selectus Strebel, 1909; Rev. Unterfam. Orthalicinen: 37-38; pl. 4, figs. 54, 58, 63 (shell).

Orthalicus selectus (Strebel).  Richardson, 1993: 113.

 

  TYPE LOCALITY.-  Trinidad.

 

  DISTRIBUTION RECORDS.-  GUATEMALA, Dept. Alta Verapaz: Coban (Strebel, 1909).

 

  REMARKS.-  The inclusion of this taxon in the Central American fauna is questionable.

 

 

Orthalicus sphinx sphinx (Strebel, 1909)

 

Zebra sphinx Strebel, 1909; Rev. Unterfam. Orthalicinen: 66-67; pl. 12, figs. 181, 185.

Orthalicus sphinx (Strebel, 1909).-  Richardson, 1993: 113.

 

  TYPE LOCALITY.-  Tepic, Nayarit.

 

  DISTRIBUTION RECORDS.-  NAYARIT: known only from the type locality.

 

 

 

Orthalicus sphinx latistriatus (Strebel, 1909)

 

Zebra sphinx form latistriata Strebel, 1909; Rev. Unterfam. Orthalicinen: 67; pl. 12, figs. 177, 178.

Orthalicus sphinx latistriatus (Strebel).  Richardson, 1993: 113.

 

  TYPE LOCALITY.-  Not stated.

 

  DISTRIBUTION.-  Unknown.

 

 

 

Orthalicus sphinx turrita (Strebel, 1909)

 

Zebra sphinx form turrita Strebel, 1909; Rev. Unterfam. Orthalicinen: 67.

Orthalicus sphinx turrita (Strebel).  Richardson, 1993: 113.

 

  TYPE LOCALITY.-  An island above San Blas, Nayarit.

 

  DISTRIBUTION RECORDS.-  NAYARIT: known only from the type locality.

 

  REMARKS.-  See REMARKS  under Orthalicus quagus.

 

 

 

Orthalicus tepicensis tepicensis (Strebel, 1909)

 

Zebra tepicensis Strebel, 1909; Rev. Unterfam. Orthalicinen: 45; pl. 6, figs. 97, 98 (shell).

Orthalicus tepicensis (Strebel).  Richardson, 1993: 113.

 

  TYPE LOCALITY.-  Tepic, Nayarit.

 

  DISTRIBUTION RECORDS.-  NAYARIT: known only from the type locality.        

 

 

 

Orthalicus tepicensis rollei (Strebel, 1909)

 

Zebra tepicensis form rollei Strebel, 1909; Rev. Unterfam. Orthalicinen: 45; pl. 6, figs. 95, 99, 100 (shell).

Orthalicus tepicensis rollei (Strebel).  Richardson, 1993: 113.

 

  TYPE LOCALITY.-  Colima.

 

  DISTRIBUTION RECORDS.-  COLIMA: known only from the type locality.

 

 

 

Orthalicus torrei (McGinty, 1939)

 

Oxystyla torrei McGinty, 1939; Nautilus, 52: 93-94.  McGinty, 1939; Nautilus, 53: pl. 2, fig. 7 (shell).

 

  TYPE LOCALITY.-  Vicinity of Acapulco, Guerrero.  Holotype ANSP 173342.

 

  DISTRIBUTION RECORDS.-  GUERRERO: known only from the type locality.

 

 

 

Orthalicus uhdeanus Von Martens, 1893

 

Orthalicus livens var. uhdeanus Von Martens, 1893; Biol. Cent. Amer.: 189.

Oxystyla livens var. uhdeanus (Von Martens).  Pilsbry, 1899; Man. Conch., 12: 129; pl. 23, figs. 16-16; pl. 22, figs. 3-4.

Zebra uhdeanus (Von Martens).  Strebel, 1909; Rev. Unterfam. Orthalicinen: 63; pl. 10, figs. 158, 161, 162.

 

  TYPE LOCALITY.-  Michoacán.

 

  DISTRIBUTION RECORDS.-  GUERRERO: Acapulco (Strebel, 1909).  MICHOACÁN: not known from a specific locality (Von Martens, 1893).

 

 

 

Orthalicus zoniferus Strebel, 1882

 

Orthalicus zoniferus Strebel, 1882: 28-29; pl. 1, figs. 7a, 7b (shell).-  Von Martens, 1893; Biol. Cent. Amer.: 186; pl. 10, figs. 12, 13.-   Solem, 1959; Occ. Pap. Mus. Zool. Univ. Mich., (611): 7-8.-  Breure & Schouten, 1985; Zool. Meded. Uitg. d. Rijksmus. Nat. Hist. Leiden, 51: 35; text-fig. 21 (reproductive anatomy).

Oxystyla zonifera (Strebel).  Pilsbry, 1899; Man. Conch., 12: 124-125; pl. 23, figs. 10-14 (shell).

Zebra zoniferus (Strebel).  Strebel, 1909; Rev. Unterfam. Orthalicinen: 51-56; pl. 8, figs. 112-115, 118-119 (shell).

 

  TYPE LOCALITY.-  Iguala, Guerrero.

 

  DISTRIBUTION RECORDS.-  COLIMA: no specific locality (Rolle, 1895)  GUERRERO: Iguala; Rancho del Platanillo, near Iguala; Chilpancingo; Venta de Zopilote; Monte Pelegrina, N of Acapulco (Pilsbry, 1899); Laguna Coyuca (Solem, 1959).  JALISCO: hill 1.0 mi. NW of Zapotiltic (Breure & Schouten, 1985). MICHOACÁN: Tafetan (19º25’ N, 100º51’ W); between Tzitzio and Heutamo; 1.2 mi. S of Charapendaro [Charapandiro] (19º15’ N, 101º58’ W) (Solem, 1959).  OAXACA: no specific locality (Von Martens, 1893). 

 

 

 

Subfamily BULIMULINAE Tryon, 1867

 

  DISTRIBUTION.-  Tropical and subtropical Americas, South Pacific Islands, southern Australia.

 

  TAXONOMY.-  Numerous genera and species.  The taxonomy employed follows Breure (1978, 1979).  Richardson (1995) gives a bibliography of the species.  The study area is inhabited by 132 species and 53 subspecies.

 

 

Genus Plekocheilus Guilding, 1828

 

  TYPE SPECIES.-  Voluta aurisseleni Born, 1780.

 

  DISTRIBUTION.  Widely distributed in South America.

 

  TAXONOMY.-  Five subgenera are recognized (Breure, 1979).  One enters the southeastern most region of Panama.

 

 

 

Subgenus Eudolichotis Pilsbry, 1896

 

  TYPE SPECIES.-  Bulimus distortus Bruguière, 1789.

 

  DISTRIBUTION.-  Northern areas of Colombia, Venezuela, Guayana, Surinam, French Guayana, Trinidad, Grenada and Pará, Brazil.  Southeastern Panama.

 

  TAXONOMY.-  About a dozen species are recognized. A single species occurs in the study area.

 

 

 

Plekocheilus (Eudolichotis) distorta panamensis (Pilsbry, 1910)

 

Auris distorta panamensis Pilsbry, 1910; Proc. Acad. Nat. Sci. Phila., 62: 507-508; pl. 37, figs. 8-9.

 

  TYPE LOCALITY.-  Between Tabernillo and San Pablo, Panama.  Holotype ANSP 101314.

 

  DISTRIBUTION.-  PANAMA: known only from the type locality.

 

 

 

Genus Berendtia Crosse & Fischer, 1869

 

Berendtia Crosse & Fischer, 1869; Jour. de Conchyl, 17: 191.-  Christensen & Miller, 1975: 44-46.-  Breure, 1978; Uitg. Rijkmuseum Nat. Hist. Leiden (164): 77-78.

 

  TYPE SPECIES.-  Clausilia (Balea ?) taylori Pfeiffer, 1861.

 

  DISTRIBUTION.-  Central Baja California Sur.

 

  TAXONOMY.-  The genus is monotypic.

 

 

 

Berendtia taylori (Pfeiffer, 1861)

 

Clausilia (Balea ?) taylori Pfeiffer, 1961; Proc. Zool. Soc. Lond.: 27; pl. 2, fig. 7. 

Berendtia taylori (Pfeiffer).  Crosse & Fischer, 1870; Jour. de Conchyl., 18: 15; pl. 5, figs. 11-13.-  Fischer & Crosse, 1875; Miss. Sci. Mex.: 304-306; pl. 14, figs. 1, 1a, 1b (hsell); pl. 16, fig. 1 (jaw), fig. 2, 3 (radula), fig. 4 (nervous system).-  Pilsbry, 1903; Man. Conch., 15: 57; pl. 18, figs. 30-36 (shell).-  Hanna, 1923; Proc. Calf. Acad. Sci., (4), 12: 506; pl. 8, fig. 10 (shell).-  Christensen & Miller, 1975; Nautilus, 89: 44-46; text-fig. 1 (reproductive anatomy), fig. 5 (shell).-  Breure, 1978; Uitg. Rijkmuseum Nat. Hist. Leiden (164): 159; figs. 261 (reproductive anatomy).-  Smith, Miller, Christensen & Roth, 1990; Proc. Calif. Acad. Sci., 47: 124-125; text-fig. 27 (map). 

Cylindrella (Urocoptis) newcombiana Gabb, 1868; Amer. Jour. Conch., 3: 237; pl. 16, fig. 3 (shell).

 

  TYPE LOCALITY.-   Clausilia (Balea ?) taylori: unknown.  Cylindrella newcombiana: Lower California, especially above Moleje [Molegé].

 

  DISTRIBUTION RECORDS.- BAJA CALIFORNIA SUR: restricted to the Sierra de la Giganta.  Smith, et al (1990) list numerous localities within this range.

 

 

 

Genus Bulimulus Leach, 1814 

 

  TYPE SPECIES.-  Helix exilis Gmelin, 1791

 

  DISTRIBUTION.-  South America from Argentina and Paraguay north to the Greater Antilles and southeastern Mexico.

 

  TAXONOMY.-  Breure (1979) list about 100 species and subspecies names that have been proposed in the genus.  Seven species and three subspecies are recognized in Mexico and Central America.

 

 

 

Bulimulus coriaceus (Pfeiffer, 1856)

 

Bulimus coriaceus Pfeiffer, 1856; Proc. Zool. Soc. Lond.: 318.

Bulimulus coriaceus (Pfeiffer).  Fischer & Crosse, 1875; Miss. Sci. Mex.: 552; pl. 21, figs. 7, 7a (shell).-  Von Martens, 1893; Biol. Cent. Amer.: 248.-  Pilsbry, 189; Man. Conch., 11: 51-52; pl. 10, figs. 77-80 (shell).

Bulimus behrendti Pfeiffer, 1861; Malak. Blätt., 8: 168.

Bulimulus coriaceaus var. berendti (Pfeiffer).  Von Martens, 1893; Biol. Cent. Amer.: 249.

 

  TYPE LOCALITY.-  State of Veracruz.

 

  DISTRIBUTION RECORDS.-  OAXACA: Cacaprieto, near Santa Efigenia, Tehuantepec (Von Martens, 1893).  TABASCO: Teapa; San Juan Bautista (Von Martens, 1893).  VERACRUZ: Jalapa; Orizaba; Cordoba; Veracruz; Toxpan, nr. Cordoba; Atoyac (Von Martens, 1893).

 

 

 

 

Bulimulus corneus corneus (Sowerby, 1833)

 

Bulimus corneus Sowerby, 1833; Proc. Zool. Soc.London, 1833: 37.-  Neubert & Jannsen, 2004; Arch. für Moll., 123: 206; pl. 10, fig. 107 (syntype).

Bulimulus (Leptomerus) corneus (Sowerby).  Fischer & Crosse, 1875; Miss. Sci. Mex.: 548.

Bulimulus corneus (Sowerby).-  Pilsbry, 1897; Man. Conch., 11: 54-55; pl. 10, fig. 82 (shell).-  Pérez, Santamaria & López, 1996; Estud. Mus. Cienc. Naturales de Avala, 10-11: 259-265 (ecology).-  Pérez & López, 2002: 208-210.

 

  TYPE LOCALITY.-  Polvón, Dept. Chinadega, Nicaragua.

 

  DISTRIBUTION RECORDS.-  COSTA RICA, Alahuela Prov.: Las Palmares. San Jose Prov.: San Jose.  Guanacaste Prov.: Guanacaste.  Limon Prov.: Puerto Viejo, on the Rio Sarapiquí (Pilsbry, 1897). GUATEMALA, Dept. Retalhuleu: Retalhuleu.  Dept. Escuintla: Escuintla. Dept. Zacapa: Zacapa.  NICARAGUA, Pacific versant, numerous localities (Pérez & López, 2002).  Dept. Rio San Juan: San Juan,             Castillo; Toro Rapids (Pilsbry, 1897); Bluefields (Fluck, 1905).

 

 

 

 

Bulimulus corneus nubeculatus (Pfeiffer, 1853)

 

Bulimus nubeculatus Pfeiffer, 1853; Proc. Zool. Soc. Lond.: 257.

Bulimulus corneus var. nubeculatus (Pfeiffer).  Von Martens, 1893; Biol. Cent. Amer.: 247.-  Pilsbry, 1897; Man. Conch., 11: 56.

 

  TYPE LOCALITY.-  Central America.  Lectotype BMNH 1975407 (Breure, 1979; Zool. Verhandl. Uit. Rijkmuseum Nat. Hist. Leiden (168): 63).

 

  DISTRIBUTION RECORDS.-  Not known from a definite locality.

 

  TAXONOMY.-  Regarded as a questionable variety of Bulimulus corneus by Von Martens (1893) and by Pilsbry, (1897).

 

 

 

 

Bulimulus corneus minor (Von Martens, 1893)

 

Bulimulus (Leptomerus) nubeculatus (Pfeiffer).  Fischer & Crosse, 1875; Miss. Sci. Mex.: 556; pl. 25, figs. 7, 7a (shell).

Bulimulus corneus var. minor Von Martens, 1893; Biol. Cent. Amer.: 247.-  Pilsbry, 1897; Man. Conch., 11: 56; pl. 10, figs. 71, 72 (shell).

 

  TYPE LOCALITY.-  San Andres Tuxtla, Veracruz.

 

  DISTRIBUTION RECORDS.-  VERACRUZ: known only from the type locality.

 

  TAXONOMY.-  Regarded as a questionable variety of Bulimulus corneus by Von Martens (1893) and by Pilsbry (1897).

 

 

 

Bulimulus dysoni (Pfeiffer, 1846)

 

Bulimus dysoni Pfeiffer, 1846; Proc. Zool. Soc. Lond.: 39.

Bulimulus dysoni (Pfeiffer).  Fischer & Crosse, 1875; Miss. Sci. Mex.: 551.-  Von Martens, 1893; Biol. Cent. Amer.: 241.-  Pilsbry, 1891; Proc. Acad. Nat. Sci. Phila: 315.-  Pilsbry, 1897; Man. Conch., 11: 56-57; pl. 10, fig. 83 (shell).-  Breure, 1979; Zool. Verhandl. Uit. Rijkmuseum Nat. Hist. Leiden (168): 62.

 

  YPE LOCALITY.-  Honduras.  Syntypes BMNH 197453 (Breure, 1979; Zool. Verhandl. Uit. Rijkmuseum Nat. Hist. Leiden (168): 62.

 

  DISTRIBUTION RECORDS.-  HONDURAS: not known from a definite locality.  NICARAGUA, Dept. Chontales: Acoyapa (Martens, 1893).  GUATEMALA, Dept. Sacatepeques: Duenas (Pilsbry, 1897).  YUCATAN: Tabi (Pilsbry, 1891).

 

 

 

Bulimulus inermis inermis (Morelet, 1851)

 

Bulimus inermis Morelet, 1851; Test. Noviss., II: 10.

Bulimulus (Leptomerus) inermis (Morelet).  Fischer & Crosse, 1875; Miss. Sci. Mex.: 550; pl. 20, figs. 16, 17 (shell).

Bulimulus inermis (Morelet).  Pilsbry, 1897; Man. Conch., 11: 51; pl. 19, fig. 87 (shell).-  Bequaert & Clench, 1933; Pub. Carnegie Inst. Wash., (431): 534.

 

  TYPE LOCALITY.-  Palizada, state of Campeche. 

 

  DISTRIBUTION RECORDS.-  CAMPECHE: known only from the type locality.

 

 

Bulimulus inermis var. major Von Martens, 1897

 

Bulimulus inermis var. major Von Martens, 1897; Biol. Cent. Amer.: 250.-  Köhler, 2007; Mitt. Mus. für Nat. Berlin, Zool., 82: 134, fig. 37 (syntype).

 

  TYPE LOCALITY.-  Northern Guatemala, valley of the Rio Cisoy at Rancho Chisoy.  Syntype ZMB 112736.

 

  DISTRIBUTION RECORDS.-  GUATEMALA: known only from the type locality.

 

 

Bulimulus inutilis (Reeve, 1850)

 

Bulimus inutilis Reeve, 1850; Conch. Icon.: pl. 86, fig. 639 (shell).

Bulimulus inutilis (Reeve).  Pilsbry, 1897; Man. Conch., 11: 73-74; pl. 11, fig. 37 (shell).-  Breure, 1978; Uitg. Rijkmuseum Nat. Hist. Leiden (164): 144.-  Breure, 1979; Zool. Verhandl. Uit. Rijkmuseum Nat. Hist. Leiden (168): 63.

 

  TYPE LOCALITY.-  Central America.  Lectotype BMNH 1975162 (Breure, 1978; Uitg. Rijkmuseum Nat. Hist. Leiden (164): 144).

 

  DISTRIBUTION.- Unknown.  The taxonomic validity of this form is questionable, and its inclusion in the Central American fauna is doubtful.

 

 

 

Bulimulus sarcodes (Pfeiffer, 1846)

 

Bulimus sarcodes Pfeiffer, 1846a; Proc. Zool. Soc. Lond.: 30.

Bulimus (Leptomerus) sardodes Pfeiffer.  Albers, 1850; Die Heliceen: 166.

Bulimulus (Leptonerus) sarcodes (Pfeiffer).  Von Martens, 1893; Biol. Cent. Amer.: 248.

Bulimulus sarcodes (Pfeiffer).  Pilsbry, 1897; Man. Conch., 11: 56; pl. 10, figs. 74-76 (shell).

 

  TYPE LOCALITY.-  Honduras.  Syntype ZMB 112737 (Kohler, 2007: 135).

 

  DISTRIBUTION RECORDS.-  HONDURAS: not recorded from a specific locality.

 

 

Bulimulus unicolor (Sowerby, 1833)

 

Bulimus unicolor Sowerby, 1833; Proc. Zool. Soc. London, 1833: 72.-  Neubert & Jannsen, 2004; Arch. fur Moll., 123: 233; pl. 10, fig. 109 (syntype).

Bulimulus unicolor (Sowerby).   Pilsbry, 1897: Man. Conch., 11: 53; pl. 10, fig. 73 (shell).-  Bequaert & Clench, 1933; Pub. Carnegie Inst. Wash., (431): 534.-  Branson & McCoy, 1963; Nautilus, 76: 105.-  Thompson, 1967; Bull. Fla. St. Mus., 11: 244-248.

Bulimulus unicolor unicolor (Reeve).  Breure, 1978; Uitg. Rijkmuseum Nat. Hist. Leiden (164): 147-148.

Bulimus umbraticus Reeve, 1849; Conch. Icon.: pl. 77, fig. 550.

Bulimus ignavus Reeve, 1849; Conch. Icon.: pl. 77, fig. 562.

Bulimus Peténensis Morelet, 1852: 10.-  Neubert & Jannsen, 2004; Archiv fur Molluskenkunde, 123: 222; pl. 10, fig. 108 (paralectotype).

Bulimulus unicolor petenensis (Morelet).  Breure, 1978; Uitg. Rijkmuseum Nat. Hist. Leiden (164): 149. 

Bulimulus (Leptomeris) istapensis Fischer & Crosse, 1975: 549; pl. 20, fig. 18.

Bulimulus sanmiguelensis Richards, 1937; Proceedings of the American Philosophical Society, 77: 253-254; pl. 4, fig. 6.

Bulimulus ochraspiris Branson & McCoy, 1965; Neotropica, 11: 97-100; figs. 1a, 1b (shell), fig. 2 (jaw), figs. 3 (radula).-  Thompson, 1969: 106-107.

 

 

  TYPE LOCALITIES.-  Bulimus unicolor: Panama, Perico Island.  Lectotype BMNH 1975411 (Breure, 1978; Uitg. Rijkmuseum Nat. Hist. Leiden (164): 148).  limus umbraticus: Central America.  Bulimus ignavus: Central America.  Lectotype BMNH 1975411 (Breure, 1979; Zool. Verhandl. Uit. Rijkmuseum Nat. Hist. Leiden (168): 63).  Bulimus petenensis: Petén, Guatemala.  Lectotype BMNH 1893.2.4.1176 (Breure, 1979: 64).  Bulimulus istapensis: Istapa, Dept. Escuintla, Guatemala.  Bulimulus sanmiguelensis: near the town of San Miguel, Cozumel Island, Quintana Roo.  Holotype ANSP 167746.  Bulimulus ochraspiris: 17.2 miles south of Chapotón, Campeche.  Holotype: University of Colorado Museum 26373.

 

  DISTRIBUTION RECORDS.-  BELIZE: W of Gales Point; Benque Viejo; Benque Viejo River, 1 mi. from cayo; Belize River (Thompson, 1967).  GUATEMALA, Dept. Petén: Paso Caballo; Flores; Puebla Nueva; Remate; island in Lago Eckixil; Uaxactun (Thompson 1967); Tikal National Park (Basch, 1959; Occ. Pap. Mus. Zool. Univ. Mich., (612): 10).  Dept. Zacapa: around Zacapa (Thompson, 1967).  HONDURAS, Dept. Chiquimula: between Jocotan and Jumusna (Thompson, 1967).   NICARAGUA: Greytown (Pilsbry, 1897).  CAMPECHE: NUMEROUS LOCALITIES (Thompson, 1967).  CHIAPAS.  QUINTANA ROO: Cozumel Island, 1.5 mi. NNE of San Miguel; 7.1 mi. NNW of Xiatil; 2.3 mi. SSE of Xiatil, 4.0 mi. E of Xpujil (Thompson, 1967).  YUCATAN: numerous localities (Pilsbry, 1897; Bequaert & Clench, 1933); Chichen Itza; San Ignacio; Merida; Uxmal; 0.8 mi. NE of Becanchén; 10.0 mi. NE of Becanchén; 7.0 mi. SSE of Uman; 1.0 mi. SSE of Puerto Telchac (Thompson, 1967).  COLIMA: 8 mi. SW of Colima (Branson & McCoy, 1963).

 

 

Genus Drymaeus Albers, 1850

 

Drymaeus is a very large genus consisting of nearly 600 named species.  The genus includes 65 species and 50 subspecies in Mexico and Central America. Since the genus first was proposed, satisfactory division into smaller units has eluded authors. Recognition of subgenera has proven to be unsustainable because of the numerous intermediate species. The arrangement of species groups presented here is modified from the systems employed by Von Martens (1893) and by Pilsbry (1899).  Nothing more satisfactory has been proposed during the past century.  Many of the species have colorful shells, and local varieties frequently have been named.  This has added to confusion when working with such a large group of species.  Many of the varieties have little basis for taxonomic recognition.  They were proposed as descriptors and not as taxonomic units, although subsequent authors have given them formal recognition.  They are included in this account while recognizing that their taxonomic status requires further study.

 

DISTRIBUTION.-  Throughout tropical and subtropical America from Argentina north to Mexico, and throughout the West Indies to Florida.

 

TAXONOMY.-  Within the study area two subgenera are recognized by Breure & Eskens (1981), Drymaeus and Mesembrinus.  They are distinguished by the development of the peristome and by some anatomical differences (Breure. 1979: 103).  The shell of hee subgenus Drymaeus has an expanded to broadly reflected outer lip, whereas in the subgenus Mesembrinus the outer lip is simple to narrowly expanded.  The distinction is arbitrary except when dealing with extreme cases.  Breure (1979) lists nine subgeneric names that have been applied to the subgenus Drymaeus.  Three subgenera have been proposed for species of the Central American fauna.

 

Drymaeus Albers, 1950: 155.  Type species by subsequent designation (Pilsbry, 1898): Helix hygrohylaeus Orbigny, 1835 (Bolivia).

Goniognathus Fischer & Crosse, 1875.  Type species by original designation: Bulimus lattrei Pfeiffer, 1847.

Metadrymaeus Pilsbry, 1926.  Type species by original designation: Bulimus josephus Angus, 1878.

 

Breure (1979) lists five subgeneric names that have been applied to Mesembrinus.  Three names pertain to the Central American fauna.

 

Mesembrinus Albers, 1850: 157.  Type species by subsequent designation, Albers, 1860: Helix virgulata Ferussac, 1821.

Antidrymaeus Germain, 1907.  Type species by subsequent designation, Pilsbry, 1926: Bulimulus inusitatus Fulton, 1900.

Leptodrymaeus Pilsbry, 1946.  Type species by original designation: Bulimus dominicus Reeve, 1850.

 

Drymaeus is divided into species groups on the basis of similar shell traits.  The distinction between species groups sometimes is arbitrary because of intraspecific shell variations.  Solem (1955) divided eastern Mexican species into four groups based on penis sheath morphology.  Breure & Eskens (1981) examined the anatomy of 49 species from throughout the range of the genus.  Their data do not support Solem’s arrangement. 

 

 

Subgenus Drymaeus Albers, 1850

 

     Thirteen species and twenty subspecies occur in the study area.  The following two species groups may be referred to Drymaeus.

 

 

Group of Drymaeus josephus

 

 

Drymaeus (Drymaeus) expansus balboa Pilsbry, 1926

 

Drymaeus expansus balboa Pilsbry, 1926; Proc. Acad. Nat. Sci. Phila., 78: 82-83; pl. 10, fig. 5-7 (shell).

 

  TYPE LOCALITY.-  Rio Puerco, Panama Prov., Panama.  Holotype ANSP 140691.

 

  DISTRIBUTION RECORDS.-  PANAMA: known only from the type locality.

 

 

 

Drymaeus (Drymaeus) josephus josephus (Angas, 1878)

 

Bulimus josephus Angas, 1873; Proc. Zool. Soc. London: 73; pl. 5, figs. 13, 14 (shell).

Otostomus josephus (Angas).  Von Martens, 1893; Biol. Cent. Amer.: 202; pl. 12.

Drymaeus josephus (Angas).  Pilsbry, 1899; Man. Conch., (2), 12: 32; pl. 26, figs. 6-13 (shell).-  Pilsbry, 1920; Proc. Acad. Nat. Sci. Phila, 72: 6.

 

  TYPE LOCALITY.-  San José, Costa Rica.

 

  DISTRIBUTION RECORDS.-  COSTA RICA, Limon Prov.: Talamanca (Von Martens, 1893); Guapiles (Pilsbry, 1920).

 

 

 

Drymaeus josephus concolor (Von Martens, 1893)

 

Otostomus josephus var. concolor Von Martens, 1893; Biol. Cent. Amer.: 202; pl. 12, figs. 8-10 (shell).-  Von Martens, 1901; Biol. Cent. Amer.: 630.-  Köhler, 2007; Mitt. Mus. für Nat. Berlin, Zool., 82:145; fig. 88 (syntype).

Drymaeus josephus var. concolor (Von Martens).  Pilsbry, 1899; Man. Conch., (2), 12: 32; pl. 20, figs. 6, 7, 8, 10, 13 (shell).

Bulimulus (Leptobrysus) zeledoni Dall, 1893; Proc. U. S. Nat. Mus.16: 644; pl. 71, fig. 2 (shell).

 

  TYPE LOCALITIES.-  Otostomus josephus var. concolor: Costa Rica; exact locality not stated.  Bulimulus zeledoni: Costa Rica; holotype USNM 98231.

 

  DISTRIBUTION RECORDS.-  COSTA RICA, Alahuela Prov.: Santa Clara. Cartago Prov.: Turrialba; Tuis.  Limon Prov:  Puerto Viejo.  Heredia Prov.: Rio Sarapiquí; upper Savegal Valley, 1000 m alt.  Puntarenas Prov.: nr. Terraba; Quebrada de Tocori, Rio Paqueta Valley; El Pital, Rio Naranjo Valley (all records from Von Martens, 1893, 1901).

 

 

 

Drymaeus (Drymaeus) josephus errans Pilsbry, 1926

 

Drymaeus (Drymaeus) josephus errans Pilsbry, 1926; Proc. Acad. Nat. Sci. Phila., 78: 84; text-fig. 15a (shell).

 

  TYPE LOCALITY.-  Mono Creek, Bocas del Torro Prov., Panama.  Holotype in the ANSP.

  DISTRIBUTION RECORDS.-  PANAMA, Bocas del Torro Prov.: Senori Creek.  Canal Zone: between Tabernilla and San Pablo.

 

 

 

Drymaeus (Drymaeus) josephus maculosus (Von Martens, 1893)

 

Otostomus josephus var. maculosus Von Martens, 1893; 202; pl. 12, fig. 7 (shell).

Drymaeus josephus var. maculosus (Von Martens).  Pilsbry, 1899; Man. Conch., (2), 12: 32; pl. 26, fig. 9 (shell).

 

  TYPE LOCALITY.-  On the banks of the Rio Pacuare del Sur; about 500 m alt.

 

  DISTRIBUTION RECORDS.-  COSTA RICA: known only from the type locality.

 

 

Drymaeus (Drymaeus) megastomus Parodiz, 1962

 

Drymaeus megastomus Parodiz, 1962; Proc. U. S. Nat. Mus.113: 435-436; pl. 1, fig. 7.

 

  TYPE LOCALITY.-  Costa Rica.  Holotype USNM 98230.

 

  DISTRIBUTION RECORDS.-  COSTA RICA, no specific locality.  PANAMA: Cerro Campaña (Parodiz, 1962). 

 

 

 

Drymaeus (Drymaeus) zhorquinensis (Angas, 1879)

 

Bulimus zhorquinensis Angas, 1879; Proc. Zool. Soc. Lond.: 478; pl. 40, fig. 4 (shell).

Otostomas zhorquinensis (Angas).  Von Martens, 1893; Biol. Cent. Amer.: 202.

Drymaeus zhorquiensis (Angas).  Pilsbry, 1899; Man. Conch., (2), 12: 31; pl. 26, figs. 14, 15.

 

  TYPE LOCALITY.-  SE Costa Rica, middle Zhorquin to Caubre, on low hills and flat ground.

 

  DISTRIBUTION RECORDS.-  COSTA RICA: known only from the type locality.

 

 

Group of Drymaeus serperastrus

 

 

Drymaeus (Drymaeus) castus castus (Pfeiffer, 1846)

 

Bulimus castus Pfeiffer, 1846; Proc. Zool. Soc. Lond.: 112.

Bulimuls (Drymaeus) castus (Pfeiffer).  Fischer & Crosse, 1875; Miss. Sci. Mex.: 485; pl. 24, figs. 11, 11a-d (shell).

Otostomus castus (Pfeiffer).  Von Martens, 1893; Biol. Cent. Amer.: 206; pl. 12, figs. 17-19, 21 (shell).

Drymaeus castus (Pfeiffer).  Pilsbry, 1899; Man. Conch., (2), 12: 43-45; pl. 9, figs. 42-51 (shell).-  Hinkley, 1920; Nautilus, 34: 50.

 

  TYPE LOCALITY.-  Not specified.

 

  DISTRIBUTION RECORDS.-  GUATEMALA, Dept. Alta Verapaz: Cobán; Tamahú (Von Martens, 1893); Chejel (Hinkley, 1950)

 

 

 

Drymaeus (Drymaeus) castus xantholeucus (Von Martens, 1893)

 

Otostomus castus var. xantholeucus Von Martens, 1893; Biol. Cent. Amer.: 206; pl. 12, figs. 16, 16a (shell).- Köhler, 2007; Mitt. Mus. für Nat. Berlin, Zool., 82: 154; fig. 139 (lectotype).

Drymaeus castus var. xantholeucus (Von Martens).  Pilsbry, 1899; Man. Conch., (2), 12: 44; pl. 9, figs. 52, 53 (shell).

 

  TYPE LOCALITY.-  Sabo [Sabob], Polochic Valley, Dept. Alta Verapaz, Guatemala.  Lectotype ZMB 109.941 (Köhler, 2007).

 

  DISTRIBUTION RECORDS.-  GUATEMALA: known only from the type locality.

 

 

 

Drymaeus (Drymaeus) castus porrectus (Von Martens, 1893)

 

Otostomus castus var. porrectus Von Martens, 1893; Biol. Cent. Amer.: 206; pl. 12, fig. 20.-  Köhler, 2007; Mitt. Mus. für Nat. Berlin, Zool., 82: 153; fig. 132 (lectotype).

Drymaeus castus var. porrectus (Von Martens).  Pilsbry, 1899; Man. Conch., (2), 12: 44.

 

  TYPE LOCALITY.-  Vera Paz, Guatemala.  Lectotype ZMB 109.942 (Köhler, 2007).

 

 DISTRIBUTION RECORDS.-  GUATEMALA, Dept. Alta Verapaz: Cobán or Tamahu (Von Martens, 1893).

 

 

 

Drymaeus (Drymaeus) chiapensis chiapensis (Pfeiffer, 1866)

 

Bulimus chiapensis Pfeiffer, 1866; Malak. Blätter, 13: 81.

Bulimulus (Drymaeus) chiapensis (Pfeiffer).  Fischer & Crosse, 1875; Miss. Sci. Mex.: 483.

Bulimulus chiapensis (Pfeiffer).  Strebel, 1882: 70-73; pl. 5, figs. 14a-c; pl. 6, figs 13, 15, 16; pl. 12, figs. 19a-c (shell).

Otostomus chiapensis (Pfeiffer).  Von Martens, 1893; Biol. Cent. Amer.: 205; pl. 12, fig. 15 (shell).

Drymaeus chiapensis (Pfeiffer). Pilsbry, 1899; Man. Conch., (2), 12: 42-43; pl. 8, figs. 27, 29-31 (shell).

 

  TYPE LOCALITY.-  Cumbre de Manzanilla, Chiapas.

 

  DISTRIBUTION RECORDS.-  MEXICO: Cuautitlan (Strebel, 1882).    VERACRUZ: Cordoba; Cerro de Palmas, nr. Cordoba; Orizaba; Coatepec (Von Martens, 1893).

 

 

 

Drymaeus (Drymaeus) chiapensis quadrifasciatus (Von Martens, 1893)

 

Otostomus chiapensis var. quatrifasciatus Von Martens, 1893; Biol. Cent. Amer.: 205; pl. 12, fig. 15.

Drymaeus chiapensis var. quadrifasciatus (Von Martens).  Pilsbry, 1899; Man. Conch., (2), 12: 42; pl. 8, fig. 32 (shell).

 

  TYPE LOCALITY.-  Matlaquihahuitl, Veracruz.

 

  DISTRIBUTION RECORDS.-  VERACRUZ: Cerro de Plumas, nr. Cordoba (Von Martens, 1893).

 

 

 

Drymaeus (Drymaeus) chiapensis nebulosus (Von Martens, 1893)

 

Otostomus chiapensis var. nebulosus Von Martens, 1893; Biol. Cent. Amer.: 205-206.

Drymaeus chiapensis var. nebulosus (Von Martens).  Pilsbry, 1899; Man. Conch., (2), 12: 43; pl. 8, fig. 33 (shell).

 

  TYPE LOCALITY.-  Cuautitlan, Mexico (Breure, 1979)

 

  DISTRIBUTION RECORDS.-  MEXICO: Cuautitlan.  VERACRUZ: Coatepec; Quimistlan (Von Martens, 1893).

 

 

 

Drymaeus (Drymaeus) colimensis (Rolle, 1895)

 

Otostomus colimensis Rolle, 1895; Nach. Deut. Malak. Ges., 32: 130.-  Von Martens, 1901; Biol. Cent. Amer.: 630; pl. 44, fig. 9 (shell).-  Köhler, 2007; Mitt. Mus. für Nat. Berlin, Zool., 82: 144; fig. 86 (lectotype).

Drymaeus colimensis (Rolle).  Pilsbry, 1899; Man. Conch., (2), 12: 47.-  Solem, 1959; Occ. Pap. Mus. Zool. Univ. Mich., (611): 9; pl. 1, fig. 16.-  Solem, 1959; Occ. Pap. Mus. Zool. Univ. Mich., (611): 8; pl. 1, fig. 16.

 

  TYPE LOCALITY.-  Colima.  Lectotype ZMB 47.660a (Köhler, 2007).

 

  DISTRIBUTION RECORDS.-  COLIMA: no specific locality known.

 

 

 

Drymaeus (Drymaeus) dombeyanus (Ferussác, 1842)

 

Bulimus dombeyanus Ferussác, 1842, in Pfeiffer, Symb. Hist. Helic., II: 76.

Bulimulus (Scutalus) dombyanus (Ferussac).  Fischer & Crosse, 1873; Miss. Sci. Mex., I: 515.-  Strebel, 1882: 74; pl. 7, fig. 3a, 3b (shell).

Drymaeus dombyanus (Ferussác).  Pilsbry, 1899; Man, Conch., 12: 33; pl. 6, figs. 1-5 (shell).-  Pilsbry, 1920; Proc. Acad. Nat. Sci. Phila., 71: 218.-  Solem, 1957 Notulae Naturae, (298): 11-12.

Bulimus alcantarae Bernardi, 1853; Jour. de Conchyl, 4: 35; pl. 3, fig. 1.

Bulimulus chaperi Crosse & Fischer, 1893; Jour. de Conchyl, 41: 31; pl. 1, figs. 1, 2 (shell).

 

  TYPE LOCALITIES.-  Bulinus dombeyanus: not stated.  Bulimus  alcantarae: not stated.  Bulimulus chaperi: Mescala Island, Lake Chapala, Jalisco.

 

  DISTRIBUTION RECORDS.-  GUERRERO: Laguna Coyuca, nr. Acapulco.  JALISCO: Presidea Island, Lago de Chapala; Guadalajara.  MICHOACÁN: E side of Cerro de Guzman, Coalcomán, 3200 ft. alt.; W side of Cerro Guzman, Coalcomán, 3500 ft. alt.; Sierra de Coalcomán, 3500-4500 ft. alt.; San José de Purua (Solem, 1957); Venta de Zopilote (Pilsbry, 1899).

 

 

 

 

Drymaeus (Drymaeus) dunkeri dunkeri (Pfeiffer, 1846)

 

Bulimus dunkeri Pfeiffer, 1846, in Philippi; Abbild. neuer conch., 2: 12; pl. 4, fig. 10.

Bulimulus (Scutalus) dunkeri (Pfeiffer).  Fischer & Crosse, 1875; Miss. Sci. Mex.: 511.-  Strebel, 1882: 76.

Otostomus dunkeri (Pfeiffer).  Von Martens, 1893; Biol. Cent. Amer.: 207-208; pl. 13, fig. 1 (shell).  

Drymaeus dunkeri (Pfeiffer).  Pilsbry, 1899; Man. Conch., (2), 12: 45-46; pl. 10, figs. 54-55; pl. 15, fig. 31 (shell).-  Solem, 1857; Notulae Naturae, (298): 13.

Drymaeus (Drymaeus) dunkeri (Pfeiffer).  Breure & Eskens, 1981; Zool. Verh. Rijkmuseum Nat. Hist. Leiden, (216): 20; pl. 6, fig. 2 (lectotype).

 

  TYPE LOCALITY.-  Michoacán, Mexico.  Lectotype BMNH 1975512 (Breure & Eskens, 1981).

 

  DISTRIBUTION RECORDS.-  MICHOACÁN: Patzcuaro; Tzintzuntzan; Morelia; Uruapan; 2 mi. S of Los Reyes (Solem, 1957).  NAYARIT: Tepíc (Pilsbry, 1899).

 

 

 

Drymnaeus (Drymaeus) dunkeri forreri (Mousson, 1883)

 

Bulimulus forreri Mousson, 1883; Jour. de Conchyl., 31: 217; pl. 9, fig. 2 (shell).

Otostomus dunkeri var. forreri (Mousson). Von Martens, 1893; 207; pl. 13, figs. 2, 2a (shell).

Drymaeus dunkeri var. forreri (Mousson).  Pilsbry, 1899; Man. Conch., (2), 12: pl. 10, figs. 56-58 (shell).

.

  TYPE LOCALITY.-  Ventanas, Durango.

 

  DISTRIBUTION RECORDS.-  CHIHUAHUA: no specific locality (Von Martens, 1893).  DURANGO: type locality.

 

 

 

Drymaeus (Drymaeus) fenestratus (Pfeiffer, 1846)

 

Bulimus fenestratus Pfeiffer, 1846; Proc. Zool. Soc. Lond.: 26.

Bulimulus (Scutalus) fenestratus (Pfeiffer).  Fischer & Crosse, 1875; Miss. Sci. Mex.: 528.

Otostomus fenestratus (Pfeiffer).  Von Martens, 1893; Biol. Cent. Amer.: 200; pl. 12. figs. 1-1b (shell).

Drymaeus fenestratus (Pfeiffer).  Pilsbry, 1899; Man. Conch., (2), 12: 34-35; pl. 7, figs. 11-14.-  Solem, 1857; Notulae Naturae, (298): 12.  

Drymaeus (Drymaeus) fenestratus (Pfeiffer).  Breure & Eskens, 1981; Zool. Verh. Rijkmuseum Nat. Hist. Leiden, (216): 22-25; text-figs. 67-76 (reproductive anatomy); pl. 2, fig. 6 (radula); pl. 5, fig. 2 (lectotype shell).

Bulimulus (Otostomus) piescheli Von Martens, 1863; in Monatsber. Akad. Wiss. Berl., 26: 541.-  Köhler, 2007; Mitt. Mus. für Nat. Berlin, Zool., 82: 152-153; fig. 131 (lectotype).

 

  TYPE LOCALITIES.-  Bulimus fenestratus: Mexico.  Lectotype BMNH 1975525 (Breure & Eskens, 1981).  Bulimulus piescheli: Manzanilla, Colima.  Bulimulus piescheli: Manzanillo, Colima; lectotype ZMB 4.581 (Köhler, 2007).

 

  DISTRIBUTION RECORDS.-  COLIMA: Manzanillo (Solem, 1957).  JALISCO: NW slope of Nevada de Colima, 7500 ft. alt. (Solem, 1957); 9.1 mi. SE of San Vicente (Breure & Eskens, 1981).  MICHOACÁN: Sierra de Coalcomán, between Pomare and Maruato; Rancho Quemado, 5600 ft. alt.; between Tecatas and Las Higueritas, 5300 ft.; Barranca de Herradero, S of San Pedro Damian, 1500 ft. alt.; above San José de los Montañas, 4700 ft. alt.; Cerro Barolosa, N of Coalcomán, 7600-9000 ft. alt. (Solem, 1959); 4 mi. S of Capacuano (Breure & Eskens, 1981).

 

 

 

Drymaeus (Drymaeus) lattrei lattrei (Pfeiffer, 1846)

 

Bulimus lattrei Pheiffer, 1846: 112; pl. 4, fig. 11 (shell).

Bulimulus delattrei (Pfeiffer).  Fischer & Crosse, 1785: 481; pl. 20, fig. 3 (animal), fig. 4 (shell); pl. 22, figs. 1-11 (anatomy).  (Name amended).

Otostomus delattrei (Pfeiffer).  Von Martens, 1893; Biol. Cent. Amer.: 204; pl. 12, figs. 11-14 (shell).

Drymaeus lattrei (Pfeiffer).  Pilsbry, 1899; Man. Conch., (2), 12: 41-42 (shell).-  van der Schalie, 1940: 4.

Drymaeus (Drymaeus) lattrei (Pfeiffer).  Breure & Eskens, 1981; Zool. Verh. Rijkmuseum Nat. Hist. Leiden, (216): 28.

Bulimus facillatus Reeve, 1848: pl. 36, fig. 211.

 

 

  TYPE LOCALITY.-  Vera Cruz, Mexico.  Lectotype BMNH 1975555 (Breure & Eskens, 1981).  The type locality was corrected to Vera Paz, Guatemala, by Pilsbry (1899:42).

 

  DISTRIBUTION RECORDS.-  GUATEMALA, Dept. Alta Verapaz: Cobán; trail to Cobán ca. 3 km E of Samac (van der Schalie, 1940).   Dept. Baja Verapaz: San Geronimo (Von Martens, 1893).

 

 

Drymaeus (Drymaeus) lattrei hiabundus (Von Martens, 1893)

 

Otostomus delattrei var. hiabundus Von Martens, 1893; 205; pl. 12; fig. 13.-  Köhler, 2007; Mitt. Mus. für Nat. Berlin, Zool., 82: 146; fig. 94 (lectotype).

Drymaeus lattrei var hiabundus (Von Martens).  Pilsbry, 1899; Man. Conch., (2), 12: 92; pl. 8, fig. 17-19 (shell). 

 

  TYPE LOCALITY.-  Cerro Zunil, [Dept. Quetzaltenango], Guatemala.  Lectotype ZMB 101.812a (Köhler, 2007).

 

  DISTRIBUTION RECORDS.-  GUATEMALA: known only from the type locality.

 

 

Drymaeus (Drymaeus) lilacinus lilacinus (Reeve, 1949)

 

Bulimus lilacinus Reeve, 1849; Conchologica Icanica, V: pl. 74, fig. 532.

Bulimulus lilacinus (Reeve).  Fischer & Crosse, 1875; Miss. Sci. Mex.: 478; pl. 20, figs. 1, 2 (shell).

Otostomus lilacinus (Reeve).  Von Martens, 1893; Biol. Cent. Amer.: 201; pl. 12, figs. 3-6 (shell).

Drymaeus lilacinus (Reeve).  Pilsbry, 1899; Man. Conch., (2), 12: 35-36; pl. 26, figs. 1-5 (shell).-  Haas, 1949; Nautilus, 62: 138.

Drymaeus (Drymaeus) lilacinus (Reeve).  Breure & Eskens, 1981; Zool. Verh. Rijkmuseum Nat. Hist. Leiden, (216): 28.

Bulimus patricius Reeve, 1849: pl. 81, fig. 600.

 

  TYPE LOCALITIES.-  Bulimus lilacinus: not given.  Bulimus patricius: not given; lectotype BMNH 1874.12.11.220 (Breure & Eskens, 1982).

 

  DISTRIBUTION RECORDS.-  GUATEMALA, Dept. Sololá: San Agustin (Von Martens, 1893).  Dept. Esquintla: Zapote (Haas, 1949).

 

 

 

Drymaeus (Drymaeus) lilacinus crossei (Von Martens, 1893)

 

Otostomus lilacinus var. crossei Von Martens, 1893; Biol. Cent. Amer.: 201.

Drymaeus lilacinus var. crossei (Von Martens).  Pilsbry, 1899; Man. Conch., (2), 12: 37; pl. 7, fig. 5, 6 (shell).

 

  TYPE LOCALITY.-  Alta Vera Paz, Guatemala.

 

  DISTRIBUTION RECORDS.-  GUATEMALA: known only from the type locality.

 

 

 

Drymaeus (Drymaeus) lilacinus ictericus (Von Martens, 1893)

 

Otostomus lilacinus var. ictericus Von Martens, 1893; Biol. Cent. Amer.: 202; pl. 12, fig. 5 (shell).-  Köhler, 2007; Mitt. Mus. für Nat. Berlin, Zool., 82: 146 (syntype).  (Not Ototostomus depictus var. ictericus Martens, 1873, a Venezuela shell.) 

Drymaeus lilacinus var. ictericus (Von Martens).  Pilsbry, 1899; Man. Conch., (2), 12: 37; pl. 7, fig. 10 (shell).

 

  TYPE LOCALITY.-  Cerro Zunil, [Dept. Quetzaltenango], Guatemala.  Syntype ZMB 109.937 (146).

 

  DISTRIBUTION RECORDS.-  GUATEMALA: known only from the type locality.

 

 

 

Drymaeus (Drymaeus) lilacinus jansoni (Von Martens, 1893)

 

Otostomus lilacinus var. jansoni Von Martens, 1893; 201; pl. 12, figs. 3-3b.

Drymaeus lilacinus var. jansoni (Von Martens).  Pilsbry, 1899; Man. Conch., (2), 12: 37; pl. 1, figs. 7-9 (shell).

 

  TYPE LOCALITY.- Nicaragua; lectotype BMNH 1901.6.22.951 (Breure & Eskens, 1981).

 

  DISTRIBUTION RECORDS.-  NICARAGUA: not known from a specific locality.

 

 

Drymaeus (Drymaeus) lilacinus undulosus (Von Martens, 1893)

 

Otostomus lilacinus var. undulosus Von Martens, 1893; Biol. Cent. Amer.: 201; pl. 12, fig. 4 (shell).-  Köhler, 2007; Mitt. Mus. für Nat. Berlin, Zool., 82: 148; fig. 110 (lectotype).

Drymaeus lilacinus var. undulosus (Von Martens).  Pilsbry, 1899; Man. Conch., (2), 12: 36; pl. 7, fig. 4 (shell).

 

  TYPE LOCALITY.-  Hacienda de Las Nubes, Cerro Zunil, [Dept. Quetzaltenango], Guatemala; 4000 ft. alt.  Lectotype ZMB 109.940a (Köhler, 2007).

 

  DISTRIBUTION RECORDS.  GUATEMALA: known only from the type locality.

 

 

 

Drymaeus (Drymaeus) lilacinus unicolor (Von Martens, 1893).

 

Otostomus lilacinus var. unicolor Von Martens, 1893; Biol. Cent. Amer.: 201; pl. 12, fig. 6 (shell).-  Köhler, 2007; Mitt. Mus. für Nat. Berlin, Zool., 82: 153, fig. 110 (lectotype).

Drymaeus lilacinus var. unicolor (Von Martens).  Pilsbry, 1899; Man. Conch., (2), 12: 37.

 

  TYPE LOCALITY.-  Cerro Zunil, Dept. Quetzaltemnango, Guatemala.  Lectotype ZMB 109.919 (Köhler, 2007).

 

  DISTRIBUTION RECORDS.-  GUATEMALA, Dept. Huehuetenango: Cholhuitz; Dept. Quetzaltenango: Cerro Zunil, 4000 ft. alt.; El Esposo, 1000 ft. alt. Dept. Sacatepeqeuez: mountain crest between Antigua and Escuintla; Hacienda Buenavista and Hacienda Helvitia (Von Martens, 1893).

 

 

 

Drymaeus (Drymaeus) serperastrus (Say, 1829)

 

Bulimus serperastrus Say, 1829; New Harmony Disseminator, Jan. 1: 25.

Bulimulus serperastrus (Say).  Fischer & Crosse, 1875; Miss. Sci. Mex.: 488; pl. 24, fig. 4.-  Strebel, 1882: 83; pl. 6, fig. 12 (shell).

Otostomus serperastrus (Say).  Von Martens, 1893; Biol. Cent. Amer.: 202-204.

Drymaeus serperastrus (Say).  Pilsbry, 1899; Man. Conch., (2), 12: 37-39; pl. 9, figs. 34-41 (shell).-  Bequaert & Clench, 1933; Pub. Carnegie Inst. Wash., (431): 532.-  Bequaert & Clench, 1938; Pub. Canegie Inst. Wash., (491): 253.-  Harry, 1950; Occ. Pap. Mus. Zool. Univ. Mich., (524): 12.-  Solem, 1955; Occ. Pap. Mus. Zool. Univ. Mich., (566): 15-16; pl. 1, fig. 8; pl. 2, figs. 1, 10 (radula); pl. 3, fig. 2 (jaw); pl. 4, fig. 6 (reproductive anatomy).-  Branson & McCoy, 1963; Nautilus, 76: 106.-  Thompson, 1967; Bull. Fla. St. Mus., 11: 249.-  Breure & Eskens, 1981; Zool. Verh. Rijkmuseum Nat. Hist. Leiden, (216): 87-88; text-figs. 297-298 (reproductive anatomy).

Bulimus liebmanni Pfeiffer, 1846; Zeitshrift für Malak.: 159.

Bulimus paivanus Pfeiffer, 1866; Malak. Blätter, 13; 81.

 

  TYPE LOCALITIES.-   Bulimus serperastrus: On the road between Vera Cruz and Mexico City; holotype ANSP 25909 (H. B. Baker, 1963: 230).  Bulimus liebmanni: Mexico.  Bulimus paivanus: Veracruz.

 

  DISTRIBUTION RECORDS.-  CAMPECHE: 7.1 mi. SW of Cd. Campeche (Thompson, 1967).  QUINTANA ROO: Cozumel Island, 3 km S of San Miguel (Branson & McCoy, 1963).  SINALOA: 22 mi. NNW of Mazatlán (Breure & Eskens, 1981).  YUCATAN: Chichén Itza; Progreso (Bequaert & Clench, 1933); Ebitz Cave, Oxkutzcab (Bequaert & Clench, 1938); San Ignacio (Solem, 1955a); 1.5 mi. S of Libre Union; Piste (Branson & McCoy, 1963).  VERACRUZ: Jalapa; Almolonga; Cordoba; Veracruz (Von Martens, 1893); Tierra Colorado, between Jalapa and Veracruz; Paso de San Juan and Loma de Piedra on the Rio Jamapa; between Paso de Ovejas and Mirador; Bobo (Pilsbry, 1899).  Reported from HIDALGO, and TAMAULIPAS, but without specific localities (Pilsbry, 1899).

 

 

 

Drymaeus (Drymaeus) ziegleri (Pfeiffer, 1846)

 

Bulimulus zieglei Pfeiffer, 1846; PS: 113.-  Binney, 1869: 133; text-fig. 336.

Drymaeus ziegleri (Pfeiffer).  Pilsbry, 1899; Man. Conch., (2), 12: 39-41; pl. 40, figs. 4-6 (shell).

Bulimus californicus Reeve, 1848: pl. 5, fig. 378.

 

  TYPE LOCALITY.-  Not stated.

 

  DISTRIBUTION RECORDS.-  SINALOA: Mazatlán; Altata (Pilsbry, 1899).

 

 

 

Subgenus Mesembrinus Albers

 

  TAXONOMY.-  Fifty-two species and thirty subspecies occur in the study area. The following species groups are referred to Mesembrinus.

 

 

Group of Drymaeus sulcosus

 

Drymaeus (Mesembrinus) aurifluus (Pfeiffer, 1856)

 

Bulimus aurifluus Pfeiffer, 1857: 319; pl. 35, fig. 10 (shell).

Bulimulus aurifluus (Pfeiffer).  Fischer & Crosse, 1875; Miss. Sci. Mex.: 499; pl. 20, figs. 21, 22 (shell).-  Strebel, 1882: 79.

Otostomus aurifluus (Pfeiffer).  Von Martens, 1893; Biol. Cent. Amer.: 213.

Drymaeus (Mesembrinus) aurifluus (Pfeiffer).  Pilsbry, 1899; Man. Conch, 12: 55; pl. 1, figs. 86-87 (shell).-  Solem, 1955; Occ. Pap. Mus. Zool. Univ. Mich., (566): 17-18; pl. 1, fig. 4 (shell); pl. 2, fig. 5 (radula); pl. 3, fig. 7 (jaw); pl. 4, fig. 3 (reproductive anatomy).-  Breure & Eskens, 1982: 53; text-fig. 187 (reproductive anatomy).

 

  TYPE LOCALITY.-  Cordoba, Veracruz.

 

  DISTRIBUTION RECORDS.-  OAXACA: Juquila; Yalalag.  VERACRUZ: Cordoba; Jalapa, Mirsdor; Tuxpan (Von Martens, 1893); Sumidero (Solem, 1955a); 4.7 mi. S of Huatusco (Breure & Eskens, 1982).

 

 

 

Drymaeus (Mesembrinus) botterii (Crosse & Fischer, 1875)

 

Bulimulus (Drymaeus) botterii Crosse & Fischer, 1875; Jour. de Conchyl., 23: 52.-  Fischer & Crosse, 1875; Miss. Sci. Mex.: 487; pl. 24, figs. 10, 10a (shell).-  Strebel, 1882: 63.

Otostomus sulcosus var. botterii (Crosse & Fischer).  Von Martens, 1893; Biol. Cent. Amer.: 208.

Drymaeus botterii (Crosse & Fischer).  Pilsbry, 1899; Man. Conch., (2), 12: 47-48; pl. 15, figs. 34, 35.

 

  TYPE LOCALITY.-  Near Orizaba, Veracruz.

 

  DISTRIBUTION RECORDS.-  VERACRUZ: known only from the type locality.

 

 

 

Drymaeus (Mesembrinus) cucullus (Morelet, 1851)

 

Bulimus cucullus Morelet, 1849; Test Noviss., I: 9.

Bulimulus (Scutalus) cucullus (Morelet).  Fischer & Crosse, 1875; Miss. Sci. Mex.: 519; pl. 20, fig. 11 (shell).

Otostomus cucullus (Morelet).  Von Martens, 1893; Biol. Cent. Amer.: 214.

Drymaeus cucullus (Morelet).  Pilsbry, 1899; Man. Conch., (2), 12: 58; pl. 1, figs. 88, 89.

Bulimulus (Scutalus) cucullus var. gracilior Fischer & Crosse, 1875; Miss. Sci. Mex.: 519; pl. 20, fig. 12.

 

  TYPE LOCALITY.-  Sisal, Yucatan.

 

  DISTRIBUTION RECORDS.-  YUCATAN: known only from the type locality.

 

 

 

Drymaeus (Mesembrinus) ghiesbreghti ghiesbreghti (Pfeiffer, 1866)

 

Bulimus ghiesbreghti Pfeiffer, 1866; Malak. Blätt., 82.

Bulimulus (Scutalus) ghiesbreghti (Pfeiffer).  Fischer & Crosse, 1875; Miss. Sci. Mex.: 522; pl. 23, fig. 4 (shell).-  Strebel, 1882: 60; pl. 6, fig. 14 (shell).

Otostomus ghiesbreghti (Pfeiffer).  Von Martens, 1893; Biol. Cent. Amer.: 209-210; pl. 13, fig. 9 (shell).

Drymaeus ghiesbreghti (Pfeiffer).  Pilsbry, 1899; Man. Conch., (2), 12: 50-52; pl. 15, figs. 32, 33; pl. 1, figs. 79, 80 (shell).

 

  TYPE LOCALITY.-  Chiapas.

 

  DISTRIBUTION RECORDS.-  OAXACA: Tlacolula (Von Martens, 1893).

 

 

 

Drymaeus (Mesembrinus) ghiesbreghti interstitialis (Von Martens, 1893)

 

Otostomus ghiesbreghti var. interstitialis Von Martens, 1893; Biol. Cent. Amer.: 210; pl. 13, fig. 9 (shell).-  Köhler, 2007; Mitt. Mus. für Nat. Berlin, Zool., 82: 151, fig. 120 (lectotype).

Drymaeus ghiesbreghti var. interstitialis (Von Martens).  Pilsbry, 1899; Man. Conch., (2), 12: 51; pl. 1, fig. 75 (shell).

 

  TYPE LOCALITY.-  Cumbres de San Martin, on the NW slope of the Cordillera, Guatemala; lectotype ZMB 112.876 (Köhler, 2007).

 

  DISTRIBUTION RECORDS.-  GUATEMALA: known only from the type locality.

 

 

 

Drymaeus (Mesembrinus) ghiesbreghti iodostylus (Pfeiffer, 1861)

 

Bulimulus iodostylus Pfeiffer, 1861; Proc. Zool. Soc. Lond.: 23

Bulimulus (Scutalus) iodostylus (Pfeiffer).  Fischer & Crosse, 1875; Miss. Sci. Mex.: 539.

Bulimulus iodostylus (Pfeiffer).  Strebel, 1882: 70; pl. 12, figs. 8a, 8b (shell).

Otostomus ghiesbreghti var. iodostylus (Pfeiffer).  Von Martens, 1893; Biol. Cent. Amer.: 210.-  Rolle, 1895: 130.

Drymaeus ghiesbreghti var. iodostylus (Pfeiffer).  Pilsbry, 1899; Man. Conch., (2), 12: 51-52; pl. 1, figs. 79, 80 (shell).

 

  TYPE LOCALITY.-  Santa Efigenia, Isthmus of Tehuantepec, Oaxaca.

 

  DISTRIBUTION RECORDS.-  COLIMA: Colima (Rolle, 1895).  OAXACA: known only from the type locality.

 

 

 

Drymaeus (Mesembrinus) ghiesbreghti stolli (Von Martens, 1887)

 

Bulimulus jonasi var. stolli Von Martens, 1887; Sitz.-Ber Ges. Naturf. Freude Berlin, 1886: 161.

Otostomus ghiesbreghti var. stolli Von Martens, 1893; Biol. Cent. Amer.: 210; pl. 13, figs. 5-8, 10 (shell).

Drymaeus ghiesbreghti var. stolli (Von Martens).  Pilsbry, 1899; Man. Conch., (2), 12: 51; pl. 1, figs. 72, 73, 76-78 (shell).

 

  TYPE LOCALITY.-  Llanos of Quetzaltenango, Dept. Quetzaltenango, Guatemala; 6000-8000 ft. alt.

 

  DISTRIBUTION RECORDS.-  GUATEMALA, Dept. Chilpancingo: Tecpam, 7000 ft. alt.  Dept. Huehuetenango: Hacienda Nubes, slopes of Cerro Zunil.  Dept. Sapatapequez: Antigua. Dept. Sololá: Argueta; Los Encuentros, 8000 ft. alt. (Von Martens, 1893).

 

 

 

Drymaeus (Mesembrinus) hegewischi (Pfeiffer, 1842)

 

Bulimus hegewischi Pfeiffer, 1842; Symbolae ad historiam heliceorum, 2: 46.

Bulimulus (Scutalus) hegewischi (Pfeiffer).  Fischer & Crosse, 1875; Miss. Sci. Mex.: 526; pl. 23, figs. 10, 10a (shell).

Bulimulus hegewischi (Pfeiffer).  Strebel, 1882: 66; pl. 6, fig. 10 (shell).

Otostomus hegewischi (Pfeiffer).  Von Martens, 1893; Biol. Cent. Amer.: 211; pl. 13, fig. 14.

Drymaeus hegewischi (Pfeiffer).  Pilsbry, 1899; Man. Conch., (2), 12: 52-54; pl. 1, figs. 81-84 (shell).-  Pilsbry, 1920: 192.-  Pilsbry, 1926; Proc. Acad. Nat. Sci. Phila., 77: text-fig. 4 (shell).

Drymaeus (Mesembrinus) hegewischi (Pfeiffer).  Breure & Eskens, 1982: 71-73; text-figs. 225-232 (reproductive anatomy), pl. 4, fig. 5 (radula).

 

  TYPE LOCALITY.-  Not stated.

 

  DISTRIBUTION RECORDS.-  GUERRERO: Omilteme.  JALISCO: Lake Chapala (Pilsbry, 1920).  MEXICO: environs of Cd. Mexico; Toluca.  MORELOS: Cuernavaca.  OAXACA: Tenango.  PUEBLA: Puebla; Tehuacán (all above records from Von Martens, 1893); NE of Chietla (Breure & Eskens, 1982).  QUERÉTARO: Querétaro (Pilsbry, 1926).  VERACRUZ: Orizaba.

 

 

 

 

Drymaeus (Mesembrinus) jonasi (Pfeiffer, 1846)

 

Bulimus jonasi Pfeiffer, 1846, in Philippi, Abbild. neuer Conchyl., 2: 125; pl. 5, fig. 4 (shell).

Bulimulus (Scutalus) jonasi (Pfeiffer).  Fischer & Crosse, 1875; Miss. Sci. Mex.: 525; pl. 23, figs 7, 7a (shell).

Otostomus jonasi (Pfeiffer).  Von Martens, 1893; Biol. Cent. Amer.: 212-213; pl. 13, figs. 11-13 (shell).

Drymaeus jonasi (Pfeiffer).  Pilsbry, 1899; Man. Conch., (2), 12: 54-55; pl. 10, figs. 64, 68-71 (shell).-  Hinkley, 1920; Nautilus, 34: 38.-  Haas, 1949; Nautilus, 62: 136, 13.

 

  TYPE LOCALITY.-  America Centralis, Vera Cruz” (Breure, 1979).  This is in error for Vera Paz, Guatemala (see Von Martens, 1893; Biol. Cent. Amer.: 212).  Three syntype BMNH 1975557.

 

  DISTRIBUTION RECORDS.-  GUATEMALA, Dept. Alta Verapaz: Cobán.  Dept. Chimaltenango: Yepocapa (Haas, 1949).  Dept. Guatemala: Amatitlan (Von Martens, 1893); Guatemala (Hinkley, 1920; Haas, 1949).  Dept. Sacatepequez: Antigua; Dueñas (Von Martens, 1893); Finca San Rafael (Haas, 1949).

 

 

 

Drymaeus (Mesembrinus) lineolatus (Conrad, 1855)

 

Bulimulus lineolatus Conrad, 1855; Proc. Acad. Nat. Sci. Phila., 7: 32.

Otosomus recluzianus var. lineolatus (Conrad).  Von Martens, 1893; Biol. Cent. Amer.: 214.

Drymaeus lineolatus (Conrad).  Pilsbry, 1899: 57; pl. 1, figs. 90, 90 (shell).

 

  TYPE LOCALITY.-  Volcán de Cartago, Cartago Prov., Costa Rica.

 

  DISTRIBUTION RECORDS.-  COSTA RICA, Cartago Prov.: Azahar de Cartago, 1600 m alt.  San Jose Prov.: San Jose, 1161 m alt.

 

 

 

Drymaeus (Mesembrinus) lirinus (Morelet, 1851)

 

Bulimus lirinus Morelet, 1851; Test. Noviss., II: 11.

Bulimulus (Drymaeus) lirinus (Morelet).  Fischer & Crosse, 1875; Miss. Sci. Mex.: 494; pl. 20, fig. 10 (shell).

Otostomus lirinus (Morelet).  Von Martens, 1893; Biol. Cent. Amer.: 214.

Drymaeus lirinus (Morelet).  Pilsbry, 1899; Man. Conch., (2), 12: 57-58; pl. 1, fig. 96 (shell).

 

  TYPE LOCALITY.-  San Luis de Petén, Dept. Petén, Guatemala.  Syntypes BMNH 1893.2.4.1954. (Breure, 1979).

 

  DISTRIBUTION RECORDS.-  GUATEMALA: known only from the type locality.

 

 

 

Drymaeus (Mesembrinus) recluzianus recluzianus (Pfeiffer, 1847)

 

Bulimus recluzianus Pfeiffer, 1847; Zeit. Malak.: 82.

Bulimulus (Scutalus) recluzianus (Pfeiffer).  Fischer & Crosse, 1875; Miss. Sci. Mex.: 510.

Bulimulus recluzianus (Pfeiffer).  Strebel, 1882: 68; pl. 6, fig. 8 (shell).

Otostomus recluzianus (Pfeiffer).  Von Martens, 1893; Biol. Cent. Amer.: 213-214.

 

  TYPE LOCALITY.-  Chiapas.

 

  DISTRIBUTION RECORDS.-  CHIAPAS: not known from a specific locality.

 

 

 

Drymaeus (Mesembrinus) recluzianus martensianus Pilsbry, 1899

 

Otostonus recluzianus var. lineolatus Conrad.  Von Martens, 1893; Biol. Cent. Amer.: 214.

Drymaeus recluzianus var. martensianus Pilsbry, 1899; Man. Conch., 12: 56.

Drymaeus (Mesembrinus) recluzianus martensianus Pilsbry.  Breure & Eskens, 1981; Zool. Verh. Rijkmuseum Nat. Hist. Leiden, (216): 83; text-figs. 267-268 (reproductive anatomy).

 

  TYPE LOCALITY.-  Not stated.

 

  DISTRIBUTION RECORDS.-  COSTA RICA, San José Prov.: San José; San Francisco de Dos Rios; La Union, 1100 m  alt. (Von Martens, 1893).

 

 

 

Drymaeus (Mesembrinus) rudis (Anton, 1839)

 

Bulimus rudis Anton, 1839; Verz. Der. Conch. Sammlungen: 43.

Bulimulus (Scutalus) rudis (Anton).  Fischer & Crosse, 1875; Miss. Sci. Mex.: 523; pl. 22, figs. 7-11 (anatomy); pl. 23, figs. 6, 6a (shell).

Bulimulus rudis (Anton).  Strebel, 1882: 63; pl. 5, fig. 3; pl. 5, fig. 9 (shell).

Otostomus rudis (Anton).  Von Martens, 1893; Biol. Cent. Amer.: 209.

Drymaeus rudis (Anton).  Pilsbry, 1899; Man. Conch., (2), 12: 49-50; pl. 15, figs. 43-46; pl. 10, figs. 65-67 (shell).-  Solem, 1955; Occ. Pap. Mus. Zool. Univ. Mich., (566): 16; pl. 1, fig. 5 (shell); pl. 2, fig. 11 (radula); pl. 4, fig. 7 (reproductive anatomy). 

Drymaeus (Drymaeus) rudis (Anton).  Breure & Eskens, 1981; Zool. Verh. Rijkmuseum Nat. Hist. Leiden, (216): 36-37; text-figs. 112-113 (reproductive anatomy).

 

  TYPE LOCALITY.-  Not stated.

 

  DISTRIBUTION RECORDS.-  MEXICO: environs of Cd. Mexico; Chapultepec (Von Martens, 1893).  MICHOACÁN: Angangüeo (Von Martens, 1893).  OAXACA: Oaxaca (Breure & Eskens, 1981).  MEXICO, D. F.: Chapultepec Park (Solem, 1955a).

 

 

 

Group of Drymaeus attenuatus

 

 

Drymaeus (Mesembrinus) attenuatus attenuatus (Pfeiffer, 1851)

 

Bulimus attenuatus Pfeiffer, 1861; Proc. Zool. Soc. Lond.: 258.

Bulimulus (Drymaeus) attenuatus (Pfeiffer).  Fischer & Crosse, 1875; Miss. Sci. Mex.: 491; pl. 23, figs. 1, 1a (shell).

Bulimulus attenuatus (Pfeiffer).  Strebel, 1882: 79-81; pl. 5, figs. 7, 7a, 8a, 8b, 8c (shell); pl. 13, fig. 13 (jaw).

Otostomus attenuatus (Pfeiffer).  Von Martens, 1893; Biol. Cent. Amer.: 215-216; pl. 13, figs. 15-15b (shell).

Drymaeus attenuatus (Pfeiffer).  Pilsbry, 1899; Man. Conch., (2), 12: pl. 2, figs. 6-8, 12-14 (shell).-  Solem, 1955; Occ. Pap. Mus. Zool. Univ. Mich., (566): 11-12; pl. 1, fig. 7 (shell); pl. 2, fig. 2 (radula); pl. 3, fig. 4 (jaw).-  Breure & Eskens, 1981; Zool. Verh. Rijkmuseum Nat. Hist. Leiden, (216): 6-8; figs. 3-6 (reproductive anatomy}.

Bulimus kefersteini Pfeiffer, 1866: 82.

Otostomus attenuatus var. concolor Von Martens, 1893; Biol. Cent. Amer.: 215; pl. 13, figs. 16-16b (shell).-  Köhler, 2007; Mitt. Mus. für Nat. Berlin, Zool., 82: 145; fig. 88 syntype).

 

  TYPE LOCALITIES.-  Bulimus attenuatus: Vera Cruz, Mexico.  Lectotype BMNH 1975458 (Breure & Eskens, 1981). Otostomus attenuatus var. concolor:  Veracruz; syntypes ZMB.

 

  DISTRIBUTION RECORDS.-  COSTA RICA, Cartago Prov.: Cartago (Breure & Eskens, 1981).  VERACRUZ: Cordoba; Orizaba, Atoyac; Mirador (Von Martens, 1893); Sumidero (Solem, 1955a).

 

 

 

Drymaeus (Mesembrinus) attenuatus pittieri (Von Martens, 1893)

 

Otostomus attenuatus var. pittieri Von Martens, 1893; Biol. Cent. Amer.: 216.-  Von Martens, 1901; Biol. Cent. Amer.: 631.-  Köhler, 2007; Mitt. Mus. für Nat. Berlin, Zool., 82: 147; fig. 105 (lectotype).

Drymaeus attenuatus var. pittieri (Von Martens).  Pilsbry, 1899; Man. Conch, 12: 64; pl. 2, fig. 15 (shell).

Drymaeus attenuatus pittieri (Von Martens).  Pilsbry, 1920c; Proc. Acad. Nat. Sci. Phila., 72: 5-6.

 

  TYPE LOCALITY.-  Alto de Mano Tigre, near Terraba, Puntarenus Prov., Costa Rica; 600 m alt.  Lectotype ZMB 48.234 (Köhler, 2007).

 

  DISTRIBUTION RECORDS.-  COSTA RICA, Cartago Prov.: road from Juan Viñas to Rio Reventazón, 3000 ft. alt. (Pilsbry, 1920).  San José Prov: Dota (Von Martens, 1893).

 

 

 

Drymaeus (Mesembrinus) attenuatus varicosus (Pfeiffer, 1851)

 

Bulimus varicosus Pfeiffer, 1851; Proc. Zool. Soc. Lond.: 256.

Otostomus attenuatus var. varicosus (Pfeiffer).  Von Martens, 1893; Biol. Cent. Amer.: 216; pl. 13, figs. 16, 16a, 16b (shell).

Drymaeus attenuatus var. varicosus (Pfeiffer).  Pilsbry, 1899; Man. Conch., (2), 12: 61; pl. 15, figs. 36, 37; pl. 2, figs. 9-11 (shell).

 

  TYPE LOCALITY.-  “Vera Cruz”.

 

  DISTRIBUTION RECORDS.-  COSTA RICA, San Jose Prov.: San Jose (Von Martens, 1901).  VERACRUZ: Cordoba; Orizaba; Atoyac; Mirador (Von Martens, 1893). 

 

 

 

Drymaeus (Mesembrinus) bugabensis (Von Martens, 1893)

 

Otostomus bugabensis Von Martens, 1893; Biol. Cent. Amer.: 218; pl. 13, figs. 21, 21a (shell).

Drymaeus bugabensis (Von Martens).  Pilsbry, 1899; Man. Conch., (2), 12: 64-65; pl. 3, figs. 34, 35 (shell).-  Pilsbry, 1926; Proc. Acad. Nat. Sci. Phila., 78: 86; text-figs. 17a.

Drymaeus (Mesembrinus) bugabensis (Von Martens).  Breure & Eskens, 1982: 54; text-fig. 4 (lectotype).

 

  TYPE LOCALITY.-  Bugaba, Chiriqui Prov., Panama.  Lectotype BMNH 1901.6.22.958 (Breure & Eskens, 1982).

 

  DISTRIBUTION RECORDS.-  PANAMA: known only from the type locality.

 

 

 

Drymaeus (Mesembrinus) chiriquiensis DaCosta, 1901

 

Drymaeus chiriquiensis DaCosta, 1901; Proc. Malac. Soc. London, 4: 238; pl. 24, fig. 1 (shell).-  Pilsbry, 1902; Man. Conch., (2), 14: 162; pl. 48, fig. 47 (shell).-  Pilsbry, 1926; Proc. Acad. Nat. Sci. Phila., 78: 86; text-fig. 17b (shell).

 

  TYPE LOCALITY.-  Boquete, Chiriqui Prov., Panama.

 

  DISTRIBUTION RECORDS.-  PANAMA: known only from the type locality.

 

 

 

Drymaeus (Mesembrinus) costaricensis (Pfeiffer, 1862)

 

Bulimus costaricensis Pfeiffer, 1862; Malak. Blätt., 9: 152.

Otostomus costarricensis (Pfeiffer).  Von Martens, 1893; Biol. Cent. Amer.: 217-218.

Drymaeus costaricensis (Pfeiffer).  Pilsbry, 1899; Man. Conch., (2), 12: 63-64; pl. 2, figs. 19-22 (shell).- Pilsbry, 1920c; Proc. Acad. Nat. Sci. Phila., 72: 5.

Drymaeus (Drymaeus) costaricensis (Pfeiffer). Breure & Eskens, 1981; Zool. Verh. Rijkmuseum Nat. Hist. Leiden, (216): 17; figs. 34-47 (reproductive anatomy).

Bulimus navarrensis Angas, 1875; Proc. Zool. Soc. Lond.: 73; pl. 5, figs. 15, 16 (shell).

 

  TYPE LOCALITY.-  Costa Rica.

 

  DISTRIBUTION RECORDS.-  COSTA RICA, Alahuela Prov.: Alahuela, 900-1000 m alt.  Cartago Prov.: Cartago to Navarro (Von Martens, 1893); Juan Viñas; on road to Reventazón (Pilsbry, 1920).  San José Prov.: San José; San Francisco de Dos Rios NICARAGUA. Dept. Matagalpa: 1.5 mi. E of Matagalpa, 2700 ft. alt. (Breure & Eskens, 1981).

 

 

 

Drymaeus (Mesembrinus) droueti droueti (Pfeiffer, 1856)

 

Bulimus droueti Pfeiffer, 1856: Proc. Zool. Soc.: 319; pl. 35, fig. 12.

Bulimulus droueti (Pfeiffer).  Fischer & Crosse, 1875; Miss. Sci. Mex.: 533; pl. 23, figs. 9. 9a (shell).-  Strebel, 1882: 77; pl. 6, fig. 6; pl. 12, fig. 7b (shell).

Otostomus droueti (Pfeiffer).  Von Martens, 1893; Biol. Cent. Amer.: 218.

Drymaeus droueti(Pfeiffer).  Pilsbry, 1899; Man. Conch., (2), 12: 65-67; pl. 3, figs. 36, 38, 43 (shell).-  Solem, 1955; Occ. Pap. Mus. Zool. Univ. Mich., (566): 12-13; pl. 1, figs. 11, 13, 14 (shell); pl. 2, fig. 4 (radula) pl. 3, fig. 6 (jaw); pl. 4, fig. 4 (reproductive anatomy).

Drymaeus bourgeoisae Rehder, 1943: 28; pl. 6, fig. 10 (shell).

 

  TYPE LOCALITIES.-  Bulimus droueti: Cordoba, Veracruz.    Drymaeus bourgeoisae: near Paraje Nuevo, near Cordoba, Veracruz; holotype USNM 517550.

 

  DISTRIBUTION RECORDS.-  VERACRUZ: Atoyac; Cordoba; Coatepec; Chirimoyo; San José Miahuatlan; Jalapa; Mirador; Orizaba; Tuxpan (Pilsbry, 1899); Potrero to Sumidero (Solem, 1955a).

 

 

 

Drymaeus (Mesembrinus) droueti deletus Solem, 1955

 

Drymaeus droueti deletus Solem, 1955; Occ. Pap. Mus. Zool. Univ. Mich., (566): 13-14; pl. 1, fig. 12 (shell).

 

  TYPE LOCALITY.-  Sumidero, Veracruz.  Holotype UMMZ 181389.

 

  DISTRIBUTION RECORDS.-  VERACRUZ: known only from the type locality.

 

 

Drymaeus (Mesembrinus) droueti sporlederi (Pfeiffer, 1866)

 

Bulimus sporlederi Pfeiffer, 1866; Makakozoologische Blätter, 13: 83.

Bulimulus sporlederi (Pfeiffer).  Fischer & Crosse, 1875; Miss. Sci. Mex.: 535; pl. 21, figs. 5, 5a (shell).-  Strebel, 1882: 78; pl. 12, figs. 7a, 7c; pl. 13, fig. 10; pl. 14, figs. 18a, 18c (shell).

Drymaeus droueti sporlederi (Pfeiffer).  Pilsbry, 1899; Man. Conch., (2), 12: 66; pl. 3, figs. 40-42 (shell).-  Solem, 1955; Occ. Pap. Mus. Zool. Univ. Mich., (566): 14; pl. 1, fig. 10 (shell).

 

  TYPE LOCALITY.-  Mirador, near Veracruz, Veracruz.

 

  DISTRIBUTION RECORDS.-  VERACRUZ: Penuela to Sumidero (Solem, 1955a).

 

 

 

Drymaeus (Mesembrinus) fenestrellus (Von Martens, 1863)

 

Bulimulus (Scutalus) fenestrellus Von Martens, 1863; 541.-  Fischer & Crosse, 1875; Miss. Sci. Mex.: 529.-  Köhler, 2007; Mitt. Mus. für Nat. Berlin, Zool., 82: 145; fig. 90 (lectotype).

Bulimulus fenestrellus (Von Martens).  Strebel, 1882; 64; pl. 5, fig. 10b (shell); pl. 13, fig. 5 (radula); pl. 14, figs. 6a, 6b (reproductive anatomy).

Otostomus fenestrellus (Von Martens).  Von Martens, 1893; Biol. Cent. Amer.: 214-215.

Drymaeus fenestrellus (Von Martens).  Pilsbry, 1899; Man. Conch., (2), 12: 58-60; pl. 2, figs. 1-5 (shell).

Drymaeus (Drymaeus) fenestrellus (Von Martens).  Breure & Eskens, 1981; Zool. Verh. Rijkmuseum Nat. Hist. Leiden, (216): 26; text-figs. 56-57 (reproductive anatomy).

Bulimus gealei H. Adams, 1867; Proc. Zool. Soc. Lond., 309; pl. 19, fig. 21.

Otostomus fenestrellus var. subunicolor Von Martens, 1893; Biol. Cent. Amer.: 215.

 

  TYPE LOCALITY.-  Bulimulus (Scutalus) fenestrellus: Patzcuaro, Michoacán; lectotype ZMB 4.457a (Köhler, 2007; Mitt. Mus. für Nat. Berlin, Zool., 82: 145).  Ototstomus fenestrellus var. subunicolor: Puebla; lectotype ZMB 18.767a (Köhler, 2007).

 

  DISTRIBUTION RECORDS.-  MICHOACÁN: nr. El Tigre (Breure & Eskens, 1981).  PUEBLA: Matamoros Izucar; Puebla (Von Martens, 1893).

 

 

 

Drymaeus (Mesembrinus) hepatostomus (Pfeiffer, 1861)

 

Bulimus hepatostomus Pfeiffer, 1861; Proc. Zool. Soc. Lond.: 23; pl. 3, fig. 4 (shell).

Bulimulus (Drymaeus) hepatostomus (Pfeiffer).  Fischer & Crosse, 1875; Miss. Sci. Mex.: 493; pl. 21, figs. 2, 2a (shell).

Otostomus hepatostomus (Pfeiffer).  Von Martens, 1893; Biol. Cent. Amer.: 217; pl. 13, figs. 18-20 (shell).

Drymaeus hepatostomus (Pfeiffer).  Pilsbry, 1899; Man. Conch., (2), 12: 62-63; pl. 2, figs. 23-27 (shell).

Drymaeus (Drymaeus) hepatostomus (Pfeiffer).  Breure & Eskens, 1981; Zool. Verh. Rijkmuseum Nat. Hist. Leiden, (216): 28; pl. 8, fig. 10 (lectotype).

 

  TYPE LOCALITY.-  Tepenistlahuaca, Oaxaca.  Lectotype BMNH 1975571 (Breure & Eskens, 1981).

 

  DISTRIBUTION RECORDS.-  OAXACA: Juquila (Von Martens, 1893).

 

 

 

Drymaeus (Mesembrinus) inglorius inglorius (Reeve, 1848)

 

Bulimus inglorius Reeve, 1848; Conchologica Iconica, 5: pl. 55, fig. 368 (shell).

Otostomus inglorius (Reeve).  Von Martens, 1893; Biol. Cent. Amer.: 219.

Drymaeus inglorius (Reeve).  Pilsbry, 1899; Man. Conch., (2), 12: 67-68; pl. 3, 43 (shell).

Drymaeus (Mesembrinus) inglorius (Reeve).  Breure & Eskens, 1981; Zool. Verh. Rijkmuseum Nat. Hist. Leiden, (216): 73-74; figs. 233-234 (reproductive anatomy).

 

  TYPE LOCALITY.-  Juquila, Oaxaca.  Lectotype BMNH 1975536 (Breure & Eskens, 1981).

 

  DISTRIBUTION RECORDS.-  OAXACA: Oaxaca (Breure & Eskens, 1981)

 

 

 

Drymaeus (Mesembrinus) inglorius heynemanni (Pfeiffer, 1866)

 

Bulinus heynemanni Pfeiffer, 1866; Malak. Blätter, 13: 89.

Bulimulus (Scutalis) heynemanni (Reeve).  Fischer & Crosse, 1875; Miss. Sci. Mex.: 527. 

Bulimulus heynemanni (Reeve).  Strebel, 1882: 68; pl. 6, fig. 7.

Otostomus inglorius var. heynemanni (Reeve).  Von Martens, 1893; Biol. Cent. Amer.: 220.

Drymaeus inglorius heynemanni (Reeve).  Pilsbry, 1899; Man. Conch., (2), 12: 68; pl. 3, figs. 44, 45, 47-52 (shell).

 

  TYPE LOCALITY.-  Orizaba, Veracruz.

 

  DISTRIBUTION RECORDS.-  OAXACA: Cerro San Antonio de la Cal; Tlacolula.  PUEBLA: Tecomavaca, SE of Tehuacán (Von Martens, 1893).  VERACRUZ: type locality. 

 

 

 

Drymaeus (Mesembrinus) necaxanus Solem, 1955

 

Drymaeus necaxanus Solem, 1955; Occ. Pap. Mus. Zool. Univ. Mich., (566): 14-15; pl. 1, fig. 9 (shell); pl. 2, fig. 6 (radula); pl. 5, fig. 5 (jaw); pl. 4, fig. 5 (reproductive system).

 

  TYPE LOCALITY.-  Pine slope near Salto Grande, Necaxa, Puebla.  Holotype UMMZ 181377.

 

  DISTRIBUTION RECORDS.-  PUEBLA: known only from the type locality.

 

 

 

Drymaeus (Mesembrinus) pluvialis (Pfeiffer, 1862)

 

Bulimus pluvialis Peiffer, 1862; Malak. Blätt., 9: 153.

Otostomus pluvialis (Pfeiffer).  Von Martens, 1893; Biol. Cent. Amer.: 218.

Drymaeus pluvialis (Pfeiffer).  Pilsbry, 1899; Man. Conch, 12: 64; pl. 3, figs. 28, 29 (shell).

 

  TYPE LOCALITY.-  Costa Rica.

 

  DISTRIBUTION RECORDS.-  COSTA RICA: not known from a specific locality.

 

 

 

Drymaeus (Mesembrinus) rFLMNHescens pinchoti Pilsbry, 1930

 

Drymaeus rFLMNHescens pinchoti Pilsbry, 1930; Proc. Acad. Nat. Sci. Phila., 82: 252-255; text-figs. 10a (reproductive anatomy), 10b, 10c (jaw and radula); pl. 18, fig. 13-24 (shells).

Drymaeus (Mesembrinus) rFLMNHescens pinchoti Pilsbry.  Breure & Eskens, 1981; Zool. Verh. Rijkmuseum Nat. Hist. Leiden, (216): 83-84; text-figs. 269-277; pl. 1, fig. 2.  (reproductive anatomy).

 

  TYPE LOCALITY.- COLOMBIA, Old Providence Island: about half a mile inland from South Bay.  Holotype ANSP 150852.

 

  DISTRIBUTION RECORDS.-  COLOMBIA: known only from Old Providence Island.

 

 

Drymaeus (Mesembrinus) sargi sargi (Crosse & Fischer, 1875)

 

Bulimulus sargi Crosse & Fischer, 1875; Jour. de Conchyl., 23: 52.

Bulimulus (Scutalus) sargi Crosse & Fischer.  Fischer & Crosse, 1875; Miss. Sci. Mex.: 534; pl. 24, figs. 6, 6a (shell).

Otostomus sargi (Crosse & Fischer).  Von Martens, 1893; Biol. Cent. Amer.: 218.

Drymaeus sargi (Crosse & Fischer).  Pilsbry, 1899; Man. Conch., (2), 12: 65; pl. 3, figs. 30, 31 (shell).

 

  TYPE LOCALITY.-  Tamahú, Dept. Alta Verapaz, Guatemala.

 

  DISTRIBUTION RECORDS.-  GUATEMALA: known only from the type locality.

 

 

Drymaeus (Mesembrinus) sargi motaguae (Von Martens, 1893)

 

Otostomus sargi var. motaguae Von Martens, 1893; Biol. Cent. Amer.: 218; pl. 14, figs. 2, 2a (shell). 

Otostomus sargi ar. montagua Breure, 1979.-  Köhler, 2007; Mitt. Mus. für Nat. Berlin, Zool., 82: 152; fig. 126 (lectotype).

Drymaeus sargi var. motaguae (Von Martens).  Pilsbry, 1899; Man. Conch., (2), 12: 65; pl. 3, figs. 32, 33 (shell).

 

  TYPE LOCALITY.-  Rio Motagua Valley, Dept. Baja Verapaz, Guatemala.  Lectotype ZMB 112.877a (Köhler, 2007).

 

  DISTRIBUTION RECORDS.-  GUATEMALA: known only from the type locality.

 

 

Drymaeus (Mesembrinus) sulcosus (Pfeiffer, 1841)

 

Bulimus sulcosus Pfeiffer, 1841, Symbolae Hist. Heliciorum, 1: 43.

Bulimulus sulcosus (Pfeiffer).  Fischer & Crosse, 1875; Miss. Sci. Mex.: 523; pl. 23, figs. 6, 6a (shell); pl. 22, figs. 7-11 (anatomy).-  Strebel, 1882: 63; pl. 5, fig. 3; pl. 6, fig. 9; pl. 11, figs. 12-15 (shell).

Otostomus sulcosus (Pfeiffer).  Von Martens, 208; pl. 13. figs. 3, 3a, 4 (shell).

Drymaeus (Drymaeus) sulcosus (Pfeiffer).  Pilsbry, 1899; Man. Conch, 12: 48-49; pl. 10, figs. 59-83.-  Solem, 1955; Occ. Pap. Mus. Zool. Univ. Mich., (566): 16-17; pl. 1, fig. 6 (shell); pl. 3, fig. 1 (jaw); pl. 4, fig. 1 (reproductive anatomy).-  Breure & Eskens, 1981; Zool. Verh. Rijkmuseum Nat. Hist. Leiden, (216): 42-44; text-figs. 141-149 (reproductive anatomy).

 

  TYPE LOCALITY.-  Tacubaya, Mexico.  There are six states that have a place named Tacubaya: Distrito Federal, Oaxaca, Jalisco, Guanajuato, Chihuahua and Yucatan.

 

  DISTRIBUTION RECORDS.-  GUERRERO: Soledad; mountains near Chilpancingo, 9500 ft. alt. (Von Martens, 1893); Cerro Muerto, 2600 ft. alt. (Breure & Eskens, 1981).  MEXICO: nr. Cd. Mexico; summit de los Aguas Escondidas (Von Martens, 1893); Santa Rosa to El Desierto de los Leones (Solem, 1955a).  VERACRUZ: Tuxpán (Strebel, 1882).

 

 

 

Drymaeus (Mesembrinus) trimarianus (Von Martens, 1893)

 

Otostomus trimarianus Von Martens, 1893; Biol. Cent. Amer.: 216; pl. 13, fig. 17 (shell).

Drymaeus trimarianus (Von Martens).  Pilsbry, 1899; Man. Conch., (2), 12: 62; pl. 2, figs. 17, 18 (shell).-  Jacobson, 1958: 11.

Drymaeus (Mesembrinus) trimarianus (Von Martens).  Breure, 1979; Zool. Verhandl. Uit. Rijkmuseum Nat. Hist. Leiden (168): 24.-  Köhler, 2007; Mitt. Mus. für Nat. Berlin, Zool., 82: 153; fig. 135 (syntype).

 

  TYPE LOCALITY.-  Tres Marias Islands, Nayarit.  Lectotype BMNH 1901.6.22.950 (Breure, 1979).

 

  DISTRIBUTION RECORDS.  NAYARIT, Tres Marias Islands: Maria Madre Island (Jacobson, 1958).

 

 

 

Group of Drymaeus tripictus

 

 

Drymaeus (Mesembrinus) irazuensis (Angas, 1878)

 

Bulimus irazuensis Angas, 1878; Proc. Zool. Soc. Lond.: 73; pl. 5, figs. 17-21 (shell).

Otostomus irazuensis (Angas).  Von Martens, 1893; Biol. Cent. Amer.: 224; pl. 14, figs. 12, 12a, 13, 13a (shell).-  Von Martens, 1901; Biol. Cent. Amer.: 631.

Drymaeus irazuensis (Angas).  Pilsbry, 1899; Man. Conch., (2), 12: 68-69; pl. 6, figs. 16-20, 24, 25 (shell).

 

  TYPE LOCALITY.-  Volcán de Irazú, Cartago Prov., Costa Rica.

 

  DISTRIBUTION RECORDS.-  COSTA RICA, Cartago Prov.: Tierra Blanca, on the south slope of Volcán Irazu, 1800 m alt. (Von Martens, 1893); Estrella de Cartago (Von Martens, 1901).

 

 

 

Drymaeus (Mesembrinus) tripictus tripictus (Albers, 1857)

 

Bulimus tripictus Albers, 1857; Malak. Blätt., 3: 97.

Otostomus tripictus (Albers).  Von Martens, 1893; Biol. Cent. Amer.: 225.

Drymaeus tripictus (Albers).  Pilsbry, 1899; Man. Conch., (2), 12: 69-70; pl. 6, figs. 12-15.

Bulimulus rhodotrema Von Martens, 1868; Malak. Blätt., 15: 156.-  Köhler, 2007; Mitt. Mus. für Nat. Berlin, Zool., 82: 153, fig. 133 (holotype).

 

  TYPE LOCALITY.-  Bulimus tripictus: unknown.  Bulimulus rhodotrema: Costa Rica; holotype ZMB 14.410 (Köhler, 2007).

 

  DISTRIBUTION RECORDS.-  COSTA RICA: not known from a specific locality.

 

 

 

Drymaeus (Mesembrinus) tripictus hoffmanni (Von Martens, 1893)

 

Otostomus tripictus var. hoffmanni Von Martens, 1893; Biol. Cent. Amer.: 225; pl. 14, figs. 11, 11a (shell).-  Von Martens, 1901; Biol. Cent. Amer.: 631.  

Drymaeus tripictus var. hoffmanni (Von Martens).  Pilsbry, 1899; Man. Conch., (2), 12: 70; pl. 6, figs. 5. 6 (shell).-  Pilsbry, 1926; Proc. Acad. Nat. Sci. Phila., 78: 85-86; text-figs. 16a-d (shell).

Drymaeus prestoni DaCosta, 1906; Proc. Malac. Soc. London, 7: 9; pl. 1, fig. 9 (shell).

Drymaeus prestoni var. cancellata DaCosta, 1906; Proc. Malac. Soc. London, 7: 9; pl. 1, fig. 10.

 

  TYPE LOCALITIES.-  Drymaeus tripictus var. hoffmanni: Woods of San Lorenzo de Dota, San José Prov., Costa Rica; 1300 m alt.  Drymaeus prestoni: Chiriqui, Panama.  Holotype BMNH 1975478 (Breure, 1979).  Drymaeus prestoni cancellata: Chiriqui, Panama; holotype BMNH 1907.11.21.13 (Breure, 1979).

 

  DISTRIBUTION RECORDS.-  COSTA RICA, Heredia Prov.: Heredia (Von Martens, 1893); slope of Volcán de Barba, 1600 m alt.  San Jose Prov.: Tarbaca, 1700 m alt.  PANAMA, Chiriqui Prov.: Chiriqui (DaCosta, 1906).

 

 

 

Drymaeus (Mesembrinus) gabbi (Angas, 1879)

 

Bulimus gabbi Angas, 1879; Proc. Zool. Soc. Lond.: 477; pl. 40, figs. 3, 3a (shell).

Otostomus gabbi (Angas). Von Martens, 1893; Biol. Cent. Amer.: 207.

Drymaeus gabbi (Angas).  Pilsbry, 1899; Man. Conch., (2), 12: 70-71; pl. 6, figs. 7-11 (shell).

 

  TYPE LOCALITY.-  Central Costa Rica, upon the flanks of Pico Blanco, at an altitude of about 3000 to 6000 ft.  Lectotype BMNH 1879.7.22.23 (Breure, 1979).

 

  DISTRIBUTION RECORDS.-  COSTA RICA, Heredia Prov.: La Paz, on the road to the Rio Sarapiquí (Von Martens, 1893).

 

 

 

Drymaeus pilsbryi Zetek, 1933

 

Drymaeus pilsbryi Zetek, 1933; Nautilus, 47; pl. 13, figs. 1 (shell).

 

  TYPE LOCALITY.-  Barro Colorado Island, Canal Zone, Panama.  Holotype ANSP 162124.

 

  DISTRIBUTION RECORDS.-  PANAMA, Canal Zone: Pedro Miguel.  San Blas Com.: San Blas; Ustupu [Ustupo] (Zetek, 1933).

 

 

 

Group of Drymaeus totonacus

 

 

Drymaeus (Mesembrinus) albostriatus (Strebel, 1882)

 

Bulimulus albostriatus Strebel, 1882: 94; pl. 6, fig. 3 (shell).

Drymaeus alsostriatus (Strebel).  Pilsbry, 1899; Man. Conch., (2), 12: pl. 12, figs. 25, 26 (shell).

 

  TYPE LOCALITY.-  Tehuantepec, Oaxaca.

 

  DISTRIBUTION RECORDS.-  OAXACA: known only from the type locality.

 

 

 

Drymaeus (Mesembrinus) championi (Von Martens, 1893)

 

Otostomus championi Von Martens, 1893; Biol. Cent. Amer.: 222; pl. 14, fig. 5 (shell).

Drymaeus championi (Von Martens).  Pilsbry, 1899; Man. Conch., (2), 12: 73; pl. 5, fig. 10 (shell).

Drymaeus (Drymaeus) championi (Von Martens).  Breure & Eskens, 1981; Zool. Verh. Rijkmuseum Nat. Hist. Leiden, (216): 14.

 

  TYPE LOCALITY.-  Hacienda de Las Nubes, Cerro Zunil, [Dept. Quetzaltenango], Guatemala.  Lectotype BMNH 1901.6.22.451 (Breure & Eskens, 1981).

 

  DISTRIBUTION RECORDS.-  GUATEMALA: known only from the type locality.

 

 

 

Drymaeus (Mesembrinus) dominicus (Reeve, 1850)

 

Bulimus dominicus Reeve, 1850; Conch. Icon.: pl. 88, fig. 659.

Bulimulus dominicus (Reeve).  Fischer & Crosse, 1875; Miss. Sci. Mex.: 540.-  Strebel, 1882: 94.

Drymaeus dominicus (Reeve).  Pilsbry, 1899; Man. Conch., (2), 12: 3-7; pl. 20, figs. 30-32 (shell).-  Baker, 1922b: 10.-  Bequaert & Clench, 1933; Pub. Carnegie Inst. Wash., (431): 532.-  Branson & McCoy, 1965: 8.-  Thompson, 1967; Bull. Fla. St. Mus., 11: 249.-  Pérez & López, 2002: 216-218.

Drymaeus (Leptodrymaeus) dominicus (Reeve).  Pilsbry, 1946; Land Moll. N. Amer., 2: 24-26; text-figs. 14a-14d (shell).

Drymaeus (Mesembrinus) dominicus (Reeve).  Breure & Eskens, 1942: 55-56; text-figs. 188-189 (reproductive anatomy).

 

  TYPE LOCALITY.-  Florida, United States of America.  Lectotype BMNH 1975199 (Breure & Eskens, 1981.

 

  DISTRIBUTION RECORDS.-  Widely distributed from Florida to Cuba and Hispaniola, and from eastern Mexico south to Nicaragua.  NICARAGUA: San Nicolas (Pilsbry, 1899).  Depts. Leon, Rivas and Chontales (Pérez & López, 2002).  CAMPECHE: 11 mi. E of Cd. Campeche (Branson & McCoy, 1965); 19.2 mi. E of Silvituc (Thompson, 1967).  QUINTANA ROO: 4.0 mi. E of Xpujil (Thompson, 1967).  TABASCO.  VERACRUZ: Mirador; Callejon de Zamarana, nr. Veracruz (Pilsbry, 1899).  YUCATAN: Chichen Itza (Bequaert & Clench, 1933); Labna (Pilsbry, 1899).

 

 

 

Drymaeus (Mesembrinus) emeus (Say, 1829)

 

Bulimus emeus Say, 1829a; New Harmony Disseminator of Useful Knowledge: 25-26.

Otostomus emeus (Say).  Von Martens, 1893; Biol. Cent. Amer.: 222,; pl. 14, figs. 6, 6a, 8, 8a (shell).

Drymaeus emeus (Say).  Pilsbry, 1899; Man. Conch., 12: 73-75; ; pl. 4, figs. 52-61 (shell).-  Correa-Sandoval, 1993; Rev. Biol. Trop., 41: 675.-  Correa-Sandoval, 1999; Rev. Biol. Trop., 47:-  Correa-Sandoval, Gutiérrez & Reza, 1998; Acta Zool.Mex., (73):: 17.-  Correa-Sandoval, 2000; Acta Zool.Mex., (79):: 9.-  Correa-Sandoval & Castro, 2002; Acta Zool. Mex., (86): 238.

Drymaeus (Mesembrinus) emeus (Say).  Breure & Eskens, 1981; Zool. Verh. Rijkmuseum Nat. Hist. Leiden, (216): 66-68; text-figs. 213-221 (reproductive anatomy).

Bulimulus palpaloensis Strebel, 1882: 85-87; pl. 5, figs. 12a-c, 16 (shell).-  Neubert & Jannsen, 2004; Archiv fur Molluskenkunde, 123: 221; pl. 16, fig. 192 (syntype).

Otostomus emeus var. hypozonus Von Martens, 1893; Biol. Cent. Amer.: 223.-  Köhler, 2007; Mitt. Mus. für Nat. Berlin, Zool., 82: 150; fig. 116 (lectotype).

Otostomus emeus var. albivaricosus Von Martens, 1893; Biol. Cent. Amer.: 223; pl. 14, figs. 8, 8a.-  Kohler, 2007: 143; fig. 82 (letotype).

 

  TYPE LOCALITIES.-  Bulimus emeus: road from Veracruz to Mexico City.  Bulimulus palpaloensis: not specified among three localities near Misantly, Veracruz.  Otostomus emeus var. hypozonus: Jalapa, Veracruz; lectotpe ZMB 109.947a (Köhler, 2007).  Otostomus emeus var. albivaricosus: Playa Vicente, Veracruz; lectotype ZMB 109946a (Köhler, 2007).

 

  DISTRIBUTION RECORDS.-  Reported from northeastern Mexico south to Espirito Santo, Brazil (Breure & Eskens, 1982).  NUEVO LEON: Santiago (Correa-Sandoval, 1993).  PUEBLA: nr. Necaxa (Solem, 1955a). SAN LUIS POTOSÍ: numerous localities (Correa-Sandoval, et al. 1998).  TAMAULIPAS: numerous localities (Correa-Sandoval & Castro, 2002).  VERACRUZ: base of Volcán San Martin (Brere & Eskens, 1982); San Juan Cuajinampa (21°11’53” N, 97°30’00” W); El Bajío, Carr. Naranjos-Tuxpan (20°57’17” N, 97°25’57” W); Carr. Tuxpan- Poza Rica, km 234 (20°49’11” N, 97°30’00” W); Barra de Cazones, 2 km al N (20°44’51” N, 97°12’06” W); El Cedral, Carr. Poza Rica-Tajín (20°29’11” N, 97°25’23” W); Ruinas El Tajín (20°26’29” N, 97°22’30” W) (Correa-Sandoval, 2000).

 

 

 

Drymaeus (Mesembrinus) intrapictus Pilsbry, 1930

 

Drymaeus intrapictus Pilsbry, 1930; Proc. Acad. Nat. Sci. Phila., 82: 340; pl. 30, fig. 8 (shell).

 

  TYPE LOCALITY.-  Tonosi, Los Santos Prov., Panama.  Holotype ANSP 140834.

 

  DISTRIBUTION RECORDS.-  PANAMA: known only from the type locality.

 

 

 

Drymaeus (Mesembrinus) perductorum Rehder, 1943

 

Drymaeus perductorum Rehder, 1943; Nautilus, 57: 20; pl. 6, figs. 6, 7 (shell).-  Breure & Eskens, 1981; Zool. Verh. Rijkmuseum Nat. Hist. Leiden, (216): 81; figs. 257-266 (reproductive anatomy); pl. 4, fig. 5. (radula).

 

  TYPE LOCALITY.-  Near Las Grutas de Cacahuamilpa, Guerrero.  Holotype USNM 517552.

 

  DISTRIBUTION RECORDS.-  GUERRERO: know only from the immediate vicinity of the type locality.

 

 

 

Drymaeus (Mesembrinus) semimaculatus Pilsbry, 1898

 

Bulimus maculatus Lea, 1839; Trans. Amer. Philos. Soc., 6: 86; pl. 23, fig. 112. (not Bulimus maculatus Bruguiere, 1798).

Otostomus maculatus (Lea).  Von Martens, 1893; Biol. Cent. Amer.: 220-221; pl. 14, fig. 3.

Drymaeus semimaculatus Pilsbry, 1898; Man. Conch., (2), 11: 297-297; pl. 5, figs. 8, 9 (shell).-  Pilsbry, 1926; Proc. Acad. Nat. Sci. Phila., 78: 84-85.-  Pilsbry, 1930; Proc. Acad. Nat. Sci. Phila., 82: 340.-  Morrison, 1946; Smiths. Misc. Coll., 106: 43.

Drymaeus (Mesemnbrinus) semimaculatus Pilsbry.  Breure & Eskens, 1981; Zool. Verh. Rijkmuseum Nat. Hist. Leiden, (216): 84-86; text-figs. 278-287 (reproductive anatomy).

 

  TYPE LOCALITY.-  Cartagena, Colombia.

 

  DISTRIBUTION RECORDS.-  Colombia north to Guatemala.  PANAMA, Pearl Islands: San José Island (Morrison, 1946).  Canal Zone: Barro Colorado Island (Pilsbry, 1930d).  Chiriqui Prov.: Chiriqui.  Panama Prov.: Garachino.  COSTA RICA, Cartago Prov.: Tapantí (Breure & Eskens, 1982).  NICARAGUA: San Nicolas (Martens, 1893).  GUATEMALA, Dept. Petén: Dolores; San Luis.  Dept. Retalhuleu: Champerico (Von Martens, 1893).

 

 

 

Drymaeus (Mesembrinus) totonacus (Strebel, 1882)

 

Bulimulus totnacus Strebel, 1882: 84; pl. 5, figs. 13, 13a (shell); pl. 13, figs. 11, 11a (radula); pl. 14, figs. 9a-f, 10e (anatomy).

Otostomus totonacus (Strebel).  Von Martens, 1893; Biol. Cent. Amer.: 221-222.

Drymaeus totonacus (Strebel).  Pilsbry, 1899: 71-72; pl. 5, figs. 11-13 (shell).

 

  TYPE LOCALITY.-  Not stated.

 

  DISTRIBUTION RECORDS.-  VERACRUZ: Rancho Quilate, nr. Misantla; Agua Caliente, nr. Misantla (Strebel, 1892).

 

 

 

Drymaeus (Mesembrinus) tryoni tryoni (Fischer & Crosse, 1875)

 

Bulimus mexicanus (Lamarck). Reeve, 1850; Conch. Icon. Bulimus: pl. 40, fig 244.

Bulimulus (Thaumastus) tryoni Fischer & Crosse, 1875; Miss. Sci. Mex.: 565.

Otostomus tryoni (Fischer & Crosse).  Von Martens, 1893; Biol. Cent. Amer.: 232-233.

Drymaeus tryoni (Fischer & Crosse).  Pilsbry, 1899; Man. Conch., (2), 12: 75-76; pl. 3, fig. 53 (shell).

 

  TYPE LOCALITY.-  Sinaloa.

 

  DISTRIBUTION RECORDS.-  SINALOA: not known from a specific locality.

 

 

 

Drymaeus (Mesembrinus) tryoni pochutlensis (Crosse & Fischer, 1875)

 

Bulimulus tryoni pochutlensis Crosse & Fischer, 1975, in Fischer & Crosse, 1875; Miss. Sci. Mex., I: 565; pl. 24, figs. 3, 3a (shell).

Otostomus tryoni var. pochutlensis (Crossse & Fischer).  Von Martens, 1893; Biol. Cent. Amer.: 233.

Drymaeus tryoni var pochutlensis (Crosse & Fischer).  Pilsbry, 1899; Man. Conch., (2), 12: 76; pl. 3, 54-55 (shell).

 

  TYPE LOCALITY.-  Pochutla, near Chilapa, Guerrero.

 

  DISTRIBUTION RECORDS.-  GUERRERO: known only from the type locality.

 

 

 

Group of Drymaeus sulphureus

 

 

Drymaeus (Mesembrinus) moricandi moricandi (Pfeiffer, 1846)

 

Bulimus moricandi Pfeiffer, 1846; Proc. Zool. Soc. Lond.: 113.

Ulimulus (Drymaeus) moricandi (Pfeifffer).  Fischer & Crosse, 1875; Miss. Sci. Mex.: 497; pl. 24, figs 9, 9a (shell).

Drymaeus moricandi (Pfeiffer).  Pilsbry, 1899; Man. Conch., 12: 78-79; pl. 4, figs. 62-64 (shell).

Drymaeus (Mesembrinus) moricandi (Pfeiffer).  Breure & Eskens, 1981; Zool. Verh. Rijkmuseum Nat. Hist. Leiden, (216): 78; text-figs. 247-248 (reproductive anatomy).

 

  TYPE LOCALITY.-  Mount Cobán, Central America.  Lectotype BMNH 1975212 (Breure, 1979).

 

  DISTRIBUTION RECORDS.-  VERACRUZ: above Rio Metlac, 19 mi. SW and 0.8 mi. N of Fortín (Breure & Eskens, 1981).

 

 

 

 

Drymaeus moricandi hyalinoalbidus (Fischer & Crosse, 1875)

 

Bulimulus (Drymaeus) moricandi var. hyalino-albida Fischer & Crosse, 1875; Miss. Sci. Mex.: 498.

Otostomus moricandi var. hyalino-albida (Fischer & Crosse).  Von Martens, 1893; Biol. Cent. Amer.: 227.

Drymaeus moricandi var. hyalino-albidus (Fischer & Crosse).  Pilsbry, 1899; Man. Conch., 12: 79.

 

  TYPE LOCALITY.-  Chiapas.

 

  DISTRIBUTION RECORDS.-  GUATEMALA, Dept. Izabal: Izabal (Von Martens, 1893).  CHIAPAS: no specific locality.

 

 

 

Drymaeus (Mesembrinus) sulfureus (Pfeiffer, 1856)

 

Bulimus sulfureus Pfeiffer, 1857; Proc. Zool. Soc. Lond.: 318; pl. 35, fig. 11 (shell).

Bulimulus sulfureus (Pfeiffer).  Fischer & Crosse, 1875; Miss. Sci. Mex.: 495; pl. 23, figs. 3, 3a (shell).-  Strebel, 1882: 87; pl. 5, figs. 11a-d; pl. 13, figs. 15, 15a, 16; pl. 15, figs. 2a-c (shell).

Otostomus sulfureus (Pfeiffer).  Von Martens, 1893; Biol. Cent. Amer.: 225; pl. 14, figs. 14-17 (shell).-  Von Martens, 1901; Biol. Cent. Amer.: 631-632.

Drymaeus sulfureus (Pfeiffer).  Pilsbry, 1899; Man. Conch., (2), 12: 76-78; pl. 4, figs. 65-68 (shell).-  Hinkley, 1920; Nautilus, 34: 48, 51.-  Pilsbry, 1920; Proc. Acad. Nat. Sci. Phila., 72: 5.-  Van der Schalie, 1940; Occ. Pap. Mus. Zol. Univ. Mich., (413): 4.-  Haas, 1949; Nautilus, 62: 138.-  Solem, 1955; Occ. Pap. Mus. Zool. Univ. Mich., (566): 19; pl. 1, fig. 3 (shell); pl. 2, figs. 6, 9, 12 (radula); pl. 3, fig. 3 (jaw); pl. 4, fig. 2 (reproductive anatomy).-  Branson & McCoy, 1963; Nautilus, 76: 106.-  Correa-Sandoval, et al, 1998: 17.-  Correa-Sandoval, 2000; Acta Zool.Mex., (79):: 9.-  Correa-Sandoval, 2002; Acta Zool. Mex., (86): 238.

Otostomus sulfurous var. albidus Von Martens, 1893; Biol. Cent. Amer.: 226.-  Köhler, 2007; Mitt. Mus. für Nat. Berlin, Zool., 82: 148; fig. 112 (lectotype).

Otostomus sulfurous var. gracilior Von Martens, 1893; Biol. Cent. Amer.: 226; pl. 14, fig. 14 (shell).-  Köhler, 2007; Mitt. Mus. für Nat. Berlin, Zool., 82: 150, fig. 115 (lectotype).

Bulimus citronellus Angas, 1879; Proc. Zool. Soc. Lond.: 479; pl. 40, fig. 5 (shell).

Otostomus sulfureus var. obesus Von Martens, 1893; Biol. Cent. Amer.: 226; pl. 14, fig. 18 (shell).

 

  TYPE LOCALITY.-  Bulimus sulfureus: Cordoba, Veracruz. Bulimus citronellus: Uren to Lipurio, Costa Rica; lectotype BMNH 1879.7.22.19 (Breure, 1979; Zool. Verhandl. Uit. Rijkmuseum Nat. Hist. Leiden (168): 117).  Otostomus sulfurous var. albidus: Teapa, Tabasco; lectotype ZNB 109.947a (Köhler, 2007).  Otostomus sulfurous var. gracilior: Senahu, Dept. Alta Verapaz, Guatemala; lectotype ZMB 109.887a (Köhler, 2007).  Otostomus sulfureus var. obesus: Huatusco, Veracruz.

 

  DISTRIBUTION RECORDS.-  COSTA RICA, Alahuela Prov.: San Mateo.  Cartago Prov.: Cartago; Rio Reventazón Valley, nr. Juan Viñas, 2500 ft. alt.; Turrialba; Tuis; Santa Clara.  Heredia Prov.: San Juan, Sarapiquí, 100-200 m alt.  Limon Prov.: nr. Guapiles, 980 ft. alt.  Puntarenas Prov.: Bonnefil Farms, Rio Surubres, 700 ft. alt. (Pilsbry, 1920c).  San Jose Prov.: San Jose, 1161 m alt.  NICARAGUA, Dept. Chontales: La Libertad.  GUATEMALA, Dept. Alta Verapaz: north of Cobán; Chiacam; San Joaquin; Chacoj; Senahu; Panzos (Pilsbry, 1899); Chamá (Hinkley, 1920); Samac; Panzamala (van der Schalie, 1940).  Dept. Izabal: Jocolo (Hinkley, 1920).  Dept. Zacapa: Santa Clara (Haas, 1949).  CAMPECHE: Cd. de Carmen; 16 mi. E of Cd. Campeche (Branson & McCoy, 1963). SAN LUIS POTOSI: entrance to Las Pozas (Xilitla), 540 m alt. (21°23’39” N, 98°59’44” W); Carr. Tamazunchale-Chapulhuacán, km 15, 140 m alt. (21°12’40” N, 98°51’34” W); Carr. Tamazunchale- Chapulhuacán, 15.5 km from Tamazunchale, 140 m. alt. (21°12’26” N, 98°53’25” W); Vega Larga, 5 km from Tamazunchale, 120 m. alt. (21°14’23” N, 98°50’31” W); Cueva “El Salitre”, Xilitla (21°22’55” N, 98°57’53” W) Correa-Sandoval, et al. 1998).  TABASCO: Teapa.  TAMAULIPAS: (Correa-Sandoval, 2002).  VERACRUZ: Atoyac; Cordoba; Orizaba; Consolapa; Soncoantla; Jalapa (Pilsbry, 1899); Sumidero (Solem, 1955a); Rancho El Sol, Naranjos (21°20’00” N, 97°43’6” W); San Juan Cuajinampa (21°11’53” N, 97°30’00” W); El Bajío, Carr. Naranjos- Tuxpan (20°57’17” N, 97°25’57” W); Carr. Tuxpan- Poza Rica, km 234  (20°49’11” N, 97°30’00” W); El Cedral, Carr. Poza Rica-Tajín (20°29’11” N, 97°25’23” W); La Ordeña, Papantla (20°29’43” N, 97°18’27” W); Papantla (20°25’24” N, 97°17’18” W) (Correa-Sandoval, 2000).  YUCATAN: 1.5 mi. S of Libre Union (Branson & McCoy, 1963).

 

 

 

Group of Drymaeus multilineatus

 

 

Drymaeus (Mesembrinus) discrepans (Sowerby, 1833)

 

Bulimus discrepans Sowerby, 1833; Proc. Zool. Soc. Lond.: 72.

Bulimulus (Liostracus) discrepans (Sowerby).  Fischer & Crosse, 1875; Miss. Sci. Mex.: 503.

Otostomus discrepans (Sowerby).  Von Martens, 1893; Biol. Cent. Amer.: 230.

Drymaeus discrepans (Sowerby).  Pilsbry, 1899; Man. Conch., (2), 12: 81-82; pl. 12, figs. 18, 19 (shell).- Pérez & López, 2002: 213-215.

 

  TYPE LOCALITY.-  Central America.

 

  DISTRIBUTION RECORDS.-  COSTA RICA, Guanacaste Proc.: Salinas Bay.  NICARAGUA, Dept. Boaco: Masapa.  Dept. Grenada: Grenada.  Dept. León: San Nicolas.  Pacific versant, numerous localities (Pérez & López, 2002).  EL SALVADOR, Dept. La Union: Conchagua.  GUATEMALA, Dept. Baja Verapaz: Salama; San Geronimo, nr. Salama.  Dept. Zacapa: Zacapa. (All records from Von Martens, 1893).

 

 

 

 

Drymaeus (Mesembrinus) heterogeneus (Pfeiffer, 1866)

 

Bulimus heterogeneus Pfeiffer, 1866; Malak. Blätt., 8: 118.

Bulimulus (Liostracus) heterogeneus (Pfeiffer).  Fischer & Crosse, 1875; Miss. Sci. Mex.: 506.

Bulimulus heterogeneus (Pfeiffer).  Strebel, 1882: 92.

Otostomus heterogeneus (Pfeiffer). Von Martens, 1893; Biol. Cent. Amer.: 235.

Drymaeus heterogeneus (Pfeiffer).  Pilsbry, 1899; 85; pl. 12, figs. 22, 23 (shell).

 

  TYPE LOCALITY.-  Vera Cruz.

 

  DISTRIBUTION RECORDS.-  NAYARIT: Tepíc (Von Martens, 1893).    VERACRUZ: Mirador; Palo Gacho, Rinconada; Jalapa (Von Martens, 1893).

 

 

 

Drymaeus (Mesembrinus) inusitatus (Fulton, 1900)

 

Bulimulus (Drymaeus) inusitatus Fulton, 1900.87.-  Köhler, 2007; Mitt. Mus. für Nat. Berlin, Zool., 82: 151; fig. 121 (syntype?).

Drymaeus inusitatus (Fulton).  Pilsbry, 1902; Man. Conch., (2), 14: 162-163; pl. 26, fig. 43.-  Pilsbry, 1926; Proc. Acad. Nat. Sci. Phila., 78: 85; text-fig. 15b (shell).

Drymaeus (Mesembrinus) inusitatus (Fultn).  Breure & Eskens, 1981; Zool. Verh. Rijkmuseum Nat. Hist. Leiden, (216): 75-76; text-figs. 245-246 (reproductive anatomy).

 

  TYPE LOCALITY.-  Costa Rica; holotype BMNH 1901.4.25.28.

 

  DISTRIBUTION RECORDS.-  COSTA RICA, Limón Prov. 28.3 mi. W of Limón (Breure & Eskens, 1981).  PANAMA, Chiriqui Prov.: nr. Chiriqui Lagoon (Pilsbry, 1926).

 

 

 

Drymaeus (Mesembrinus) livescens (Pfeiffer, 1842)

 

Bulimus livescens Pfeiffer, 1842; Symbolae ad historiam heliceorum, II: 48.

Bulimulus (Mesembrinus) livescens (Pfeiffer).  Fischer & Crosse, 1875; Miss. Sci. Mex.: 543.

Bulimulus livescens (Pfeiffer,).  Strebel, 1882: 91; pl. 6, fig. 4 (shell).

Otostomus livescens (Pfeiffer).  Von Martens, 1893; Biol. Cent. Amer.: 228; pl. 15, figs. 7, 8 (shell).

Drymaeus livescens (Pfeiffer).  Pilsbry, 1899; Man. Conch., (2), 12: 80-81; pl. 4, figs. 76-81 (shell).

 

  TYPE LOCALITY.-  Tehuacán, Puebla.

 

  DISTRIBUTION RECORDS.-  GUERRERO: Chilpancingo.  PUEBLA: SE of Tehuacán; Tecomavaca (Von Martens, 1893).

 

 

 

Drymaeus (Mesembrinus) moritinctus (Von Martens, 1893)

 

Otostomus moritinctus Von Martens, 1893; Biologia Centrali  Americana: 228; pl. 14, figs. 9-10.-  Neubert & Jannsen, 2004; Archiv für Mollusk., 123: pl. 16, fig. 191 (paralectotype).

Drymaeus moritinctus (Von Martens).  Pilsbry, 1899; Man. Conch., (2), 12: 79-80; pl. 6, figs. 26-29 (shell). 

Drymaeus (Mesembrinus) moritinctus (Von Martens).  Breure & Eskens, 1981; Zool. Verh. Rijkmuseum Nat. Hist. Leiden, (216): 78.-  Köhler, 2007; Mitt. Mus. für Nat. Berlin, Zool., 82: 152; fig. 128 (paralectotype). 

 

  TYPE LOCALITY.-  Chilpancingo, Guerrero; 4600 ft. alt.  Lectotype BMNH 1901.6.22.841 (Breure & Eskens, 1981).

 

  DISTRIBUTION RECORDS.-  GUERRERO: Known only from the vicinity of the type locality.

 

 

 

Drymaeus (Mesembrinus) multilineatus (Say, 1825)

 

Bulimus multilineatus Say, 1825; Journ. Journ. Acad. Nat. Sci. Phila., 5: 120.

Drymaeus multilineatus (Say).  Pilsbry, 1899; Man. Conch., (2), 12: 27; pl. 11, figs. 27-33.-  Hinkley, 1907; Nautilus, 21: 72.-  Pilsbry, 1946; Land Moll. N. Amer., 2: 26-28; figs. 15a-15d.-  Branson & McCoy, 1965; Univ. Colorado Stud, Biol., (13): 8.-  Correa-Sandoval, Gutiérrez & Reza, 1998; Acta Zool.Mex., (73):: 17.-  Correa-Sandoval & Castro, 2002; Acta Zool. Mex., (86): 238.- Pérez & López, 2002: 218-220.

Drymaeus (Mesembrinus) multilineatus (Say).  Breure & Eskens, 1981; Zool. Verh. Rijkmuseum Nat. Hist. Leiden, (216): 78-80; text-figs. 249-256 (reproductive anatomy).)

Bulimus sisalensis Morelet, 1849; Test Noviss., I: 9.-  Neubert & Jannsen, 2004; Archiv fur Molluskenkunde, 123: 230; pl. 16, fig. 193 (paralectotype).

 

 

  TYPE LOCALITY.-  Bulimus multilineatus: Miami, Dade Co., Florida.  Neotype ANSP 12309a (Pilsbry, 1946).  Bulimus sisalensis: Sisal, Yucatan.

 

  DISTRIBUTION RECORDS.-  South Florida, Colombia, Venezuela, Curaçao and Mexico.  NICARAGUA, Depts. Managua, Chontales (Pérez & López, 2002).  CAMPECHE: 11 mi. E of Cd. Campeche (Branson &McCoy, 1965: 8).  OAXACA: 4.0 mi. WNW of Tehantepec (Breure & Eskens, 1981).  SAN LUIS POTOSÍ: Valles (Hinkley, 1907); Rancho El Rodeo; Ejido Buenavista, km 18, Carr. Cd. Valles, 500 m alt. (22°11’13” N, 99°00’00” W); Cd. del Maíz-El Naranjo, S.L.P. km 10 (22°30’00” N, 99°22’06” W); Cd. del Maíz-El Naranjo, S.L.P. km 35 (22°30’00” N, 99°22’06” W); carr. Cd. del Maíz-El Naranjo, La Cortina (22°30’58” N, 99°20’15” W) (Correa-Sandoval- Sandoval, et al, 1998).  TAMAULIPAS: numerous localities (Correa-Sandoval & Costro, 2002).  YUCATAN Sisal.

 

 

 

 

Drymaeus (Mesembrinus) semipellucidus (Tristram, 1861)

 

Bulimus semipellucidus Tristram, 1861, Proc. Zool. Soc. Lond.: 230; pl. 26, fig. 8 (shell).

Bulimulus (Liostracus) semipellucidus (Tristram).  Fischer & Crosse, 1875; Miss. Sci. Mex.: 507.

Otostomus semipellucidus (Tristram).  Von Martens, 1893; Biol. Cent. Amer.: 236; pl. 15, fig. 10 (shell).

Drymaeus semipellucidus (Tristram).  Pilsbry, 1899; Man. Conch., (2), 12: 82-83; pl. 4, figs. 71-73 (shell).

Drymaeus (Mesembrinus) semipellucidus (Tristram).  Breure & Eskens, 1981; Zool. Verh. Rijkmuseum Nat. Hist. Leiden, (216): 86-87; text figs. 288-296 (reproductive anatomy).

 

  TYPE LOCALITY.-  Guatemala.

 

  DISTRIBUTION RECORDS.-  COSTA RICA, Alahuela Prov.: Alahuela.  Guanacaste Prov.: Salinas Bay.  Heredia Prov.: Puerto Viejo, Rio Sarapiquí (Von Martens, 1893).  NICARAGUA, Dept. Masaya: 2.4 mi. N of Masatepe (Breure & Eskens, 1981).

 

 

 

Drymaeus (Mesembrinus) shattucki Bequaert & Clench, 1931

 

Drymaeus shattucki Bequaert & Clench, 1931: 424.-  Bequaert & Clench, 1933; Pub. Carnegie Inst. Wash., (431): 533-534; pl. 68, figs. 4-7.-  Bequaert & Clench, 1936; Pub. Canegie Inst. Wash., (457): 64.-  Bequaert & Clench, 1938; Pub. Canegie Inst. Wash., (491): 258.-  Basch, 1959; Occ. Pap. Mus. Zool. Univ. Mich., (612): 10.-  Branson & McCoy, 1963; Nautilus, 76: 106.

 

  TYPE LOCALITY.-  Chichen Itza, Yucatan.  Holotype MCZ 79396.

 

  DISTRIBUTION RECORDS.-  GUATEMALA, Dept. Petén: Tikal National Park (Basch, 1959).  QUINTANA ROO: 3 km S of San Miguel, Cozumel Island (Branson & McCoy, 1963).  YUCATAN: Tabi; Progreso (Bequaert & Clench, 1933); Ebitz Cave, Oxkutzcab (Bequaert & Clench, 1938). 

 

 

 

Drymaeus (Mesembrinus) tropicalis (Morelet, 1849)

 

Bulimus tropicalis Morelet, 1849; Test Noviss., I: 9.-  Neubert & Jannsen, 2004; Archiv fur Molluskenkunde, 123: 233; pl. 16, fig. 194 (paralectoype).

Bulimulus (Liostracus) tropicalis (Morelet).  Fischer & Crosse, 1875; Miss. Sci. Mex.: 504; pl. 20, figs. 7, 8 (shell).

Bulimulus tropicalis (Morelet).  Strebel, 1882: 95.-  Pilsbry, 1891; Proc. Acad. Nat. Sci. Phila.: 315.

Otostomus tropicalis (Morelet).  Von Martens, 1893; Biol. Cent. Amer.: 233.

Drymaeus tropicalis (Morelet).  Pilsbry, 1899; Man. Conch., (2), 12: 85-86; pl. 6, figs. 21-23 (shell).-  Bequaert & Clench, 1933; Pub. Carnegie Inst. Wash., (431): 533.-  Bequaert & Clench, 1938; Pub. Canegie Inst. Wash., (491): 258.-  Basch, 1959; Occ. Pap. Mus. Zool. Univ. Mich., (612): 10.-  Branson & McCoy, 1963; Nautilus, 76: 106.-  Thompson, 1967; Bull. Fla. St. Mus., 11: 248-249.

Drymaeus (Mesembrinus) tropicalis (Morelet).  Breure & Eskens, 1981; Zool. Verh. Rijkmuseum Nat. Hist. Leiden, (216): 89.

 

  TYPE LOCALITY.-  Mexico, Campeche, civic plaza of Cd. Campeche.  Lectotype BMNH 1893.2.4.210 (Breure & Eskens, 1981).

 

  DISTRIBUTION RECORDS.-  GUATEMALA, Dept. Petén: Tikal National Park (Basch, 1959).  CAMPECHE: 5-11 mi. E of Cd. Campeche; 32 mi. S of Cd. Campeche (Branson & McCoy, 1963); 5.1 mi. NNW of Dzibalchén; 3.6 mi. S of Hopelchén; 5.7 mi. E of Cd. Campeche (Thompson, 1967).  QUINTANA ROO: 4.0 mi. E of Xpujil (Thompson, 1967).  YUCATAN: Labna; Tabi, S of Merida (Pilsbry, 1891); Ebitz Cave, Oxkutzcab (Bequaert & Clench, 1938); 0.8 mi. NE of Becanchén; 19.1 mi. SSE of Uman (Thompson, 1967). 

 

 

 

Drymaeus (Mesembrinus) uhdeanus uhdeanus (Von Martens, 1893)

 

Bulimulus (Mesembrinus) uhdeanus Von Martens, 1863; in Monatsber. Akad. Wiss. Berl., 26: 541.-  Fischer & Crosse, 1875; Miss. Sci. Mex.: 530; pl. 21, figs. 4, 4a (shell).-  Strebel, 1882: 90; pl. 11, figs. 11a, 11b (shell).-  Köhler, 2007; Mitt. Mus. für Nat. Berlin, Zool., 82: 153; fig. 136 (lectotype).

Otostomus uhdeanus (Von Martens).  Von Martens, 1893; Biol. Cent. Amer.: 233; pl. 15, figs. 1-3 (shell).

Drymaeus uhdeanus (Von Martens).  Pilsbry, 1899; Man. Conch., (2), 12: 83-84; pl. 15, figs. 47, 49, 50 (shell).-  Solem, 1955; Occ. Pap. Mus. Zool. Univ. Mich., (566): 11.

Drymaeus (Mesembrinus) uhdeanus (Von Martens).  Breure & Eskens, 1981; Zool. Verh. Rijkmuseum Nat. Hist. Leiden, (216): 89; text-figs. 299-300 (reproductive anatomy).

Drymaeus herrerae Bartsch, 1907; Proc. U. S. Nat. Mus., 32: 119, text-fig. 1 (shell).

 

  TYPE LOCALITY.-  Bulimulus uhdeanus: Mexico; lectotype ZMB 4.575b (Köhler, 2007).  Drymaeus herrerae: Bonanza Zimapan, Hidalgo; holotype USNM 192997.

 

  DISTRIBUTION RECORDS.-  HIDALGO: Zimapan.  JALISCO: Sayula.  VERACRUZ: Aculcingo, south of Orizaba (Pilsbry, 1899); 8.5 mi. SW of Cd. Mendoza, 5200 ft. alt. (Breure & Eskens, 1981).

 

 

Drymaeus (Mesembrinus) uhdeanus borealis (Von Martens, 1893)

 

Otostomus uhdeanus var. borealis Von Martens, 1893; Biol. Cent. Amer.: 234; pl. 15, fig. 6 (shell).-  Köhler, 2007; Mitt. Mus. für Nat. Berlin, Zool., 82: 150; fig. 114 (lectotype).

Drymaeus uhdeanus borealis (Von Martens).  Pilsbry, 1899; Man. Conch., (2), 12: 84; pl. 15, fig. 53 (shell).

 

  TYPE LOCALITY.-  Ventanas, Durango; 2000 ft. alt.  Lectontype ZMB 101.817a (Köhler, 2007).

 

  DISTRIBUTION RECORDS.-  DURANGO: known only from the type locality.

 

 

 

Drymaeus (Mesembrinus) uhdeanus cuernovacensis (Crosse & Fischer, 1874)

 

Bulimulus (Scutalus) cuernovacensis Crosse & Fischer, 1874; Jour. Conchyl., 22: 283.-  Fischer & Crosse, 1875; Miss. Sci. Mex.: 532; pl. 23, figs. 11, 11a (shell).-  Strebel, 1882: 65; pl. 12, fig. 17 (shell).

Otostomus uhdeanus var. cuernovacensis (Crosse & Fischer).  Von Martens, 1893; Biol. Cent. Amer.: 234; pl. 15, fig. 4 (shell).

Drymaeus uhdeanus cuernovacensis (Crosse & Fischer).  Pilsbry, 1899; Man. Conch., (2), 12: 84; pl. 15, figs. 42, 51 (shell).-  Solem, 1955; Occ. Pap. Mus. Zool. Univ. Mich., (566): 11.

Drymaeus herrerae veracruzensis Bartsch, 1907: 120; text-fig. 2.

 

  TYPE LOCALITY.-  Bulimulus cuernovaensis: Cuernovaca, Morelos.  Drymaeus herrerae veracruzensis: Cordoba, Veracruz.

 

  DISTRIBUTION RECORDS.-  MORELOS: Cuernovaca.  VERACRUZ: Orizaba; Maltrata (Pilsbry, 1899).

 

 

 

 

Drymaeus (Mesembrinus) uhdeanus tepicensis (Von Martens, 1893)

 

Otostomus uhdeanus var. tepicensis Von Martens, 1893; Biol. Cent. Amer.: 234; pl. 15, fig. 5.

Drymaeus uhdeanus tepicensis (Von Martens).  Pilsbry, 1899; Man. Conch., (2), 12: 84; pl. 15, fig. 52 (shell).

 

  TYPE LOCALITY.-  Tepíc, Jalisco [Nayarit].

 

  DISTRIBUTION RECORDS.-  NAYARIT: Maria Madre Island, Tres Marias Islands (Dall, 1926).

 

 

 

Group of Drymaeus translucens

 

 

Drymaeus (Mesembrinus) cozumelensis Richards, 1937

 

Drymaeus shattucki cozumelensis Richards, 1937; Proceedings of the American Philosophical Society, 77: 253; pl. 4, fig. 4.-  Rehder, 1966; Proc. Biol. Soc. Wash., 79: 285-286; fig. 7.

 

  TYPE LOCALITY.-  On trail from San Miguel to Santa Rita, Cozumel Island, Quintana Roo. Holotype ANSP 167744.

 

  DISTRIBUTION RECORDS.-  QUINTANA ROO: known only from Cozumel Island.

 

 

Drymaeus (Mesembrinus) hondurasanus (Pfeiffer, 1846)

 

Bulimus hondurasanus Pfeiffer, 1846; Proc. Zool. Soc. Lond.: 29.

Bulimulus (Liostracus) hondurasanus (Pfeiffer): Fischer & Crosse, 1875; Miss. Sci. Mex.: 503.

Otostomus hondurasanus (Pfeiffer).  Von Martens, 1893; Biol. Cent. Amer.: 232.

Drymaeus hondurasanus (Pfeiffer).  Pilsbry, 1899; Man. Conch., (2), 12: 88-89; pl. 15, fig. 41 (shell).-  Rehder, 1966; Proc. Biol. Soc. Wash., 79: 286; figs. 5-6 (lectotype shell).

Drymaeus (D.) alternans honduranus (Pfeiffer).  Haas & Solem, 1960; Nautilus, 73: 131.

 

  TYPE LOCALITY.-  Honduras.  Lectotype in the BMNH 1975265 (Rehder, 1966).

 

  DISTRIBUTION RECORDS.-  BELIZE: Rio Frio Cave; Kate’s Lagoon (Haas & Solem, 1960).  HONDURAS: not known from a specific locality.

 

 

Drymaeus (Mesembrinus) mayaorum Rehder, 1966

 

Drymaeus mayaorum Rehder, 1966; Proceedings of the Biological Society of Washington, 79: 287; figs. 3, 4.

 

  TYPE LOCALITY.-  Isla Mujeres, Quintana Roo, Mexico.  Holotype USNM 251656.

 

  DISTRIBUTION RECORDS.-  QUINTANA ROO: Isla Mujeres; near San Miguel, Cozumel Island (Rehder 1966).

 

 

 

 

Drymaeus (Mesembrinus) translucens translucens (Broderip, 1832)

 

Bulimus translucens Broderip, 1832; Proc. Zool. Soc. Lond.: 31.

Otosomus translucens (Broderip).  Von Martens, 1893; Biol. Cent. Amer.: 250.

Drymaeus translucens (Broderip).  Pilsbry, 1899; Man. Conch., (2), 12: 89; pl. 24, figs. 28, 29 (shell).- Pilsbry, 1930; Proc. Acad. Nat. Sci. Phila., 82: 340; pl. 30, figs. 13, 14. (shell).-  Morrison, 1946; Smiths. Misc. Coll., 106: 42-43.-  Pérez & López, 2002: 221.

 

  TYPE LOCALITY.-  King and Saboga Islands, Panama.

 

  DISTRIBUTION RECORDS.-  PANAMA, Pearl Islands: King Island; Saboga Island (Pilsbry, 1926); San José Island (Morrison, 1946). NICARAGUA, Dept. Boaco (Pérez & López, 2002)?

 

 

 

Drymaeus (Mesembrinus) translucens alternans (Beck, 1837)

 

Bulimus (Bulimulus)alternans Beck, 1837; Index molluscorum.: 65.

Bulimulus(Liostracus) alternans (Beck).  Fischer & Crosse, 1875; Miss. Sci. Mex.: 500; pl. 23, figs. 5 (shell).

Otostomus alternans (Beck).  Von Martens, 1893; Biol. Cent. Amer.: 230-231.-  Von Martens, 1901; Biol. Cent. Amer.: 632.

Drymaeus alternans (Beck).  Pilsbry, 1899; Man. Conch., (2), 12: 86-88; pl. 15, figs. 38-40 (shell).-  Pilsbry, 1926; Proc. Acad. Nat. Sci. Phila., 78: 84; text-figs. 15d, 15e (shell).-  Hinkley, 1920; Nautilus, 34: 38.-  Pérez & López, 2002: 213.

Drymaeus translucens alternans (Beck).  Pilsbry, 1930; Proc. Acad. Nat. Sci. Phila., 82: 340; pl. 20, fig. 4 (shell).

Drymaeus (Mesembrinus) alternans (Beck).  Breure & Eskens, 1981; Zool. Verh. Rijkmuseum Nat. Hist. Leiden, (216): 47-50; text-figs. 160-173 (reproductive anatomy); pl. 3, fig. 7 (radula).

 

  TYPE LOCALITY.-  Saboga Island, Panama.

 

  DISTRIBUTION RECORDS.-  COSTA RICA, Alahuela Prov.: Alahuela (Von Martens, 1893).  Cartago Prov.: Cartago, 1142 m alt.  Puntarenas Prov.: San Diquis Valley, nr. Terraba, 100 m alt.  San José Prov.: San José; La Uruca, 100 m alt. (Von Martens, 1893, 1901).  GUATEMALA, Dept. Baja Verapaz: San Géronimo, nr. Salama (Von Martens, 1893).  Dept. Guatemala: Cd. Guatemala (Hinkley, 1920).  Dept. Sacatepequez: Capetillo, nr. Antigua (Von Martens, 1901).  NICARAGUA, Dept. Matagalpa: 1 mi. E of Matagalpa (Breure & Eskens, 1981).  Dept. Leon (Pérez & López, 2002).  PANAMA, Pearl Islands: King’s Island; Saboga Island (Pilsbry, 1926).

 

 

 

Drymaeus (Mesembrinus) translucens juquilensis (Von Martens, 1893)

 

Otostomus alternans var. juquilensis Von Martens, 1893; Biol. Cent. Amer.: 231.

Drymaeus alternans var. juquilensis (Von Martens).  Pilsbry, 1899; Man. Conch., (2), 12: 88; pl. 15, fig. 40 (shell).

 

  TYPE LOCALITY.-  Juquila, Oaxaca.

 

  DISTRIBUTION RECORDS.-  OAXACA: Juquila.  GUATEMALA: “Vera Paz” (Von Martens, 1893).

 

 

 

Drymaeus (Mesembrinus) translucens misellus Pilsbry, 1926

 

Drymaeus translucens form misellus Pilsbry, 1926; Proc. Acad. Nat. Sci. Phila., 78: 83; text-fig. 14b (shell).

 

  TYPE LOCALITY.-  Tonosi, Los Santos Prov., Panama.  Holotype ANSP 140281.

 

  DISTRIBUTION RECORDS.-  PANAMA: known only from the type locality.

 

 

 

Drymaeus (Mesembrinus) translucens pachecensis Pilsbry, 1930

 

Drymaeus translucens form pachenensis Pilsbry, 1930d; Proc. Acad. Nat. Sci. Phila., 82: 34; pl. 30, fig. 11 (shell).

 

  TYPE LOCALITY.-  Pacheco Island, Pearl Islands, Panama.  Holotype ANSP 151302.

 

  DISTRIBUTION RECORDS.-  PANAMA: known only from the type locality.

 

 

 

Drymaeus (Mesembrinus) translucens panamensis (Broderip, 1833)

 

Bulimus panamensis Broderip, 1833; Proc. Zool. Soc. Lond.: 105.

Otostomus panamensis (Broderip).  Von Martens, 1893; Biol. Cent. Amer.: 236.

Drymaeus panamensis (Broderip).  Pilsbry, 1899; Man. Conch., (2), 12: 90; pl. 24, fig. 30 (shell).

Drymaeus translucens form panamensis (Broderip).  Pilsbry, 1930; Proc. Acad. Nat. Sci. Phila., 82: 340; pl. 30, fig. 12.

 

  TYPE LOCALITY.-  San Miguel, King Island, Panama.

 

  DISTRIBUTION RECORDS.-  PANAMA: King Island, Saboga Island (Pilsbry, 1926).

 

 

 

Drymaeus (Mesembrinus) translucens sororcula Pilsbry, 1926

 

Drymaeus translucens sororcula Pilsbry, 1926; Proc. Acad. Nat. Sci. Phila., 78: 83; text-fig. 14c (shell).

 

  TYPE LOCALITY.-  Taboga Island, Panama.  Holotype ANSP 45238.

 

  DISTRIBUTION RECORDS.-  PANAMA: known only from the type locality.

 

 

 

Drymaeus (Mesembrinus) translucens subfloccosus Pilsbry, 1899

 

Drymaeus translucens var. subfloccusus Pilsbry, 1899; Man. Conch., (2), 12: 90; pl. 24, figs. 26, 27 (shell).-  Breure, 1979; Zool. Verhandl. Uit. Rijkmuseum Nat. Hist. Leiden (168): 124.

 

  TYPE LOCALITY.-  Nicaragua.  Lectotype ANSP 25040a (Breure, 1979).

 

  DISTRIBUTION RECORDS.-  NICARAGUA: no specific locality known.

 

 

 

Drymaeus (Mesembrinus) translucens tonosiesis Pilsbry, 1930

 

Drymaeus alternans (Beck).  Pilsbry, 1926; Proc. Acad. Nat. Sci. Phila., 78: 84; text-fig. 15E.

Drymaeus translucens tonosiesis Pilsbry, 1930: 340. 

 

  TYPE LOCALITY.-  Tonosi, Los Santos Prov., Panama.  Holotype ANSP 151288.

 

  DISTRIBUTION RECORDS.-  PANAMA: known only from the type locality.

 

 

Genus Naesiotus Albers, 1850

 

Naesiotus Albers, 1850: 162.-  Breure, 1979; Zool. Verhandl. Uit. Rijkmuseum Nat. Hist. Leiden (168): 64-72.-    Hoffman, 1988; Veliger, 30: 117-120.-  Smith, Miller, Christensen & Roth, 1990; Proc. Calif. Acad. Sci., 47: 112-119.

Sonorina Pilsbry, 1896: Nautilus, 9: 114.

Puritanina Jacobson, 1958; Amer. Mus. Nov., (1899): 7.

Hannarabdotus Emerson & Jacobson, 1965; Trans> San Diego Soc. Nat. Hist., 13: 325.

 

  TYPE SPECIES.-  Naesiotus: Bulimus nux Broderip, 1832.  Sonorina: Bulimulus spirifer Gabb, 1868.  Puritanina: Bulimulus montezuma Dall, 1893.  Hannarabdotus: Bulimulus slevini Hanna, 1923.

 

 

  DISTRIBUTION.-  South America, the Galapagos Islands, the Windward Islands of the West Indies, northwestern Mexico and Arizona (Breure, 1978; Hoffman, 1988).

 

  TAXONOMY.-  The generic synonymy given above applies only to the Mexican fauna.  Breure (1979: 67-72) lists over 160 names that have been proposed for species of Naesiotus.  Twenty species are recognized in the study area.  An Arizona species is included.

 

 

 

Naesiotus altus (Dall, 1893)

 

Naesiotus altus (Dall).  Hoffman, 1988; Veliger, 30: 420.-  Smith, Miller, Christensen & Roth, 1990; Proc. Calif. Acad. Sci., 47: 112; text-fig. 16 (map).

Bulimulus (Leptobrysus) subspirifer Mabille, 1895; Bulletin de la Société Philomathique de Paris, (8), 7: 67.-  Christensen & Miller, 1976; Western Soc. Malac. Ann. Rep., 9:: 51.

 

  TYPE LOCALITIES.-  Bulimulus altus: Sierra Laguna, Baja California Sur; 3000 ft. alt. (fide Christensen, 1978). Bulimulus subspirifer: Lower California.

 

  DISTRIBUTION RECORDS.-  BAJA CALIFORNIA SUR: 2.3 km from main hwy. on road to microwave station San Bartolo, 430 m alt.; ca. 1 km SE of San Bartolo, 240 m alt.; 2.1 km SE of San Bartolo (Smith, et al, 1990).

 

 

 

Naesiotus beldingi (Cooper, 1892)

 

Bulimulus inscendens var. beldingi Cooper, 1892; Proc. Calif. Acad. Sci., (2), 3: 209.

Bulimulus (Orthotomium) beldingi Cooper.   Dall, 1895; Proc. U. S. Nat. Mus.19: 357.-  Pilsbry, 1898; Man. Conch., 11: 149; pl. 25, fig. 56 (shell).

Naesiotus beldingi (Cooper).  Hoffman, 1988; Veliger, 30: 420.-  Smith, Miller, Christensen & Roth, 1990; Proc. Calif. Acad. Sci., 47: 112-113; text-fig. 16 (map).

Bulimulus (Leptobrysus) dismenicus Mabille, 1895; Bulletin de la Société Philomathique de Paris, (8), 7: 67.-   Emerson & Jacobson, 1964; Trans. San Diego Soc. Nat. Hist., 16: 51.  

 

  TYPE LOCALITIES.-  Bulimulus beldingi: Sierra Laguna, Punta Arena, near San Jose del Cabo, Baja California Sur.  Bulimulus dismenicus: Sierra de Puna, Baja California Sur, 1800 m alt.

 

  DISTRIBUTION RECORDS.-  BAJA CALIFORNIA SUR: several localities in the Sierra Laguna at 610-2070 m alt. (Smith, et al, 1990).

 

 

 

Naesiotus christenseni (Miller & Reeder, 1984)

 

Rabdotus christenseni Miller & Reeder, 1984; Bulletin of the Southern California Academy of Sciences, 8: 106-109; figs. 1 center, 1 right (shell), text-fig. 2 (reproductive anatomy).

Naesiotus christenseni (Miller & Reeder).  Hoffman, 1988; Veliger, 30: 417.

 

  TYPE LOCALITY.-  Arizona, Pima Co.; north end of the Santa Rita Mountains, in a large rockslide on the left bank of south fork of Sycamore Creek, ca. 100 m upstream from Sycamore Spring; 31º52.9’ N, 110º45.7’ W; ca. 1300 m alt.  Holotype CAS 37525.

 

  DISTRIBUTION RECORDS.-  ARIZONA: known only from the type locality.

 

 

 

Naesiotus cosmicus (Mabille, 1895)

 

Bulimulus (Scutalus) cosmicus Mabille, 1895; Bulletin de la Société Philomathique de Paris, (8), 7: 68.

Bulimulus (Orthotomium) cosmicus Mabille.  Pilsbry, 1897; Man. Conch., 11: 144.

Naesiotus cosmicus (Mabille).  Hoffman, 1988; Veliger, 30: 420.-  Smith, Miller, Christensen & Roth, 1990; Proc. Calif. Acad. Sci., 47: 113; text-fig. 17 (map).

 

  TYPE LOCALITY.-  “La Sierra du Sud dela Presqu’ile”, Baja California Sur.

 

  DISTRIBUTION RECORDS.-  BAJA CALIFORNIA SUR: stations on the NE side of the Sierra Laguna (Smith, et al, 1990).

 

 

 

Naesiotus dentifer dentifer (Mabille, 1895)

 

Bulimulus (Leptobrysus) dentifer Mabille, 1895; Bulletin Societe Philomath., (8) 7: 67. 

Bulimulus (Sonoribna) dentifer Mabille.  Pilsbry, 1898; Man. Conch, 11: 161-162.-  Hanna, 1923; Proc. Calf. Acad. Sci., (4), 12: 492-494; pl. 8, figs. 16-20 (shell); text-figs. 1 (reproductive anatomy, jaw).

Rabdotus (Leptobrysus) dentifer (Mabille).  Emerson & Jacobson, 1964; Trans. San Diego Soc. Nat. Hist., 16: 319-321.

Naesiotus dentifer dentifer (Mabille).  Hoffman, 1988; Veliger, 30: 420.-  Smith, Miller, Christensen & Roth, 1990; Proc. Calif. Acad. Sci., 47: 113; text-fig. 18 (map).

 

TYPE LOCALITY.-  Isla Tortuga, Baja California Sur.

 

DISTRIBUTION RECORDS.-  BAJA CALIFORNIA SUR: known only from Isla Tortuga (Smith, et al, 1990).

 

 

 

Naesiotus dentifer johnstoni (Hanna, 1923)

 

Bulimulus johnstoni Hanna, 1923; Proc. Calif. Acad. Sci., 12: 491; pl. 7, figs. 1-6 (shell); pl. 11, fig. 3 (embryonic sculpture).

Rabdotus (Hannarabdotus) johnstoni (Hanna).  Emerson & Jacobson, 1964; Trans. San Diego Soc. Nat. Hist., 16: 326.

Naesiotus dentifer johnstoni (Hanna).  Smith, Miller, Christensen & Roth, 1990; Proc. Calif. Acad. Sci., 47: 114; text-fig. 18 (map).

 

 

  TYPE LOCALITY.- Isla Santa Catalina, Baja California Sur.  Holotype CAS 1024.

 

  DISTRIBUTION RECORDS.-  BAJA CALIFORNIA SUR: known only from Santa Catalina Island (Smith, et al, 1990).

 

 

 

Naesiotus dentifer lamellifer (Pilsbry, 1897)

 

Bulimulus lamellifer Pilsbry, 1897; Nautilus, 10: 103. 

Bulimulus (Sororina) lamellifer Pilsbry, 1898; Man. Conch., 11: 160; pl. 21, figs. 94-99 (shell).-   Hanna, 1923; Proc. Calf. Acad. Sci., (4), 12: 495-497.

Rabdotus (Leptobrysus) lamellifer lamellifer (Pilsbry).  Emerson & Jacobson, 1964; Trans. San Diego Soc. Nat. Hist., 16: 321-321; figs. 2a-k (shell).

Naesiotus dentifer lamellifer (Pilsbry).  Smith, Miller, Christensen & Roth, 1990; Proc. Calif. Acad. Sci., 47: 114-115; text-fig. 18 (map).

Bulimulus spirifer var. orthelasmus Pilsbry, 1898; Man. Conch., 11: 159-160.-   Christensen & Miller, 1976; Western Soc. Malac. Ann. Rep., 9: 51.

Bulimulus ximenez Hanna, 1923; Proceedings of the California Academy of Science, (4), 12: 497- 499; pl. 8, figs. 4-9 (shell).

Rabdotus (Leptobrysus) ximenez (Pilsbry).  Emerson & Jacobson, 1964; Trans. San Diego Soc. Nat. Hist., 16: 323-324; figs. 3a-e (shell).

Bulimulus bakeri Hanna, 1923; Proc. Calf. Acad. Sci., (4), 12: 500-501; pl. 7, figs. 7-10 (shell).

Bulimulus sanmarcosensis Pilsbry & Lowe, 1932; Nautilus, 46: 49-50; unfigured.

Bulimulus carmen Pilsbry & Lowe, 1932; Nautilus, 46: 50-51; unfigured.-  Pilsbry, 1935; Proc. Acad. Nat. Sci. Phila., 87: 2; pl. 1, fig. 7 (shell).

 

  TYPE LOCALITIES.-  Bulimulus lamellifer: Lower California.  Bulimulus spirifer var. orthelasmus: not given.  Bulimulus ximinez: Marquer Bay, Isla Carmen.  Bulimulus bakeri: Punta San Antonio, south of Bahia San Nicolas, Baja California Sur; holotype CAS 1017.  Bulimulus sanmarcosensis: Isla San Marcos.  Bulimulus carmen: Salinas Bay, Isla Carmen.

 

  DISTRIBUTION RECORDS.-  BAJA CALIFORNIA SUR: northern peninsular Baja California Sur and Isla San Marcos, Isla Carmen, Isla Danzante, Isla Coronados and Isla San Jose (Smith, et al, 1990).

 

 

 

Naesiotus dentifer santacruzensis (Hanna, 1923)

 

Bulimulus santacruzensis Hanna, 1923; Proc. Calif. Acad. Sci., 12: 487; pl. 7, figs. 12-15 (shell).

Bulimulus slevini Hanna, 1923; Proc. Calif. Acad. Sci., 12: 488-489; pl. 7, figs. 16-19 (shell).

Rabdotus (Hannarebdotus) slevini (Hanna).  Emerson & Jacobson, 1964; Trans. San Diego Soc. Nat. Hist., 16: 325-326; figs. 4a-f (shell).

Naesiotus dentifer slevini (Hanna).  Smith, Miller, Christensen & Roth, 1990; Proc. Calif. Acad. Sci., 47: 115; text-fig. 18 (map).

 

  TYPE LOCALITIES.- Bulimulus santacruzensis: Isla Santa Cruz, Baja California Sur; holotype CAS 1030.  Bulimulus dentifer slevini: Isla Monserrate, Baja California Sur; holotype CAS 1034. 

 

  DISTRIBUTION RECORDS.-  BAJA CALIFORNIA SUR: known only from Isla Monserrate and Isla Santa Cruz (Smith, et al, 1990).

 

  TAXONOMY.-  Smith, et al (1990) used the name slevini for this subspecies, but the name santacruzensis has page priority.

 

 

Naesiotus durangoanus (Von Martens, 1893)

 

Bulimulus durangoanus Von Martens, 1893; Biol. Cent. Amer.: 246; pl. 15, figs. 11, 11a (shell).-  Pilsbry, 1897; Man. Conch, 11: 127-128; pl. 18, figs. 32, 33 (shell).

 

  TYPE LOCALITY.-  Villa Lerdo, Durango.

 

  DISTRIBUTION EECORDS.-  DURANGO: reported only from the type locality.

 

  TAXONOMY.-  There are many samples of this and/or closely related species in the Florida Museum of Natural History from the states of Chihuahua, Durango, Coahuila and Nuevo León.  The sculpture of the juvenile shells indicates that they are properly placed in Naesiotus.  Further study is needed to determine their specific status.

 

 

Naesiotus excelsus (Gould, 1853)

 

Bulimus excelsus Gould, 1853; Jour. Boston Soc. Nat. Hist., 6: 376; pl. 14, fig. 3 (shell).

Bulimulus (Orthotomium) excelsus (Gould).  Pilsbry, 1897; Man. Conch., 11: 141-142; pl. 20, figs. 69-71.

Naesiotus excelsus (Gould).  Hoffman, 1988; Veliger, 30: 420.-  Smith, Miller, Christensen & Roth, 1990; Proc. Calif. Acad. Sci., 47: 115; text-fig. 17 (map).

Bulimulus elatus Gould, 1853; Jour. Boston Soc. Nat. Hist., 6:: 406.

Bulimulus (Scutatus) cacotycus Mabille, 1895; Bulletin de la Société Philomathique de Paris, (8), 7: 69.-  Emerson & Jacobson, 1964; Trans. San Diego Soc. Nat. Hist., 16: 51.

Bulimulus excelsus var. sinaloae Pilsbry, 1897; Man. Conch., 11: 142.

 

  TYPE LOCALITIES.-  Bulimus excelsus: “California”.  Bulimus elatus: not given.  Bulimulus cacotycus: Les Sierras du Sud de la Presqu’ile [Baja California Sur].  Bulimulus excelsus var. sinaloae: Sinaloa (doubtful, Hanna, 1923; Proc. Calf. Acad. Sci., (4), 12: 487; Smith, et al, 1990: 115).

 

  DISTRIBUTION RECORDS.-  BAJA CALIFORNIA SUR: La Paz; N slope of La Calavera Mt. Above Playa Coromuel; Playa Coromuel; 20.6 km E of La Paz on rd. to Las Cruces; granite cliff nr. Rancho Viniamos, 2.6 km E of La Paz (Smith, et al, 1990). 

 

 

 

Naesiotus gabbi (Crosse & Fischer, 1872)

 

Bulimulus (Scutalus) gabbi Crosse & Fischer, 1872; Jour. de Conchyl., 20: 223.-  Fischer & Crosse, 1873; Miss. Sci. Mex., I: 517; l. 20, fig. 19 (shell).

Bulimulus (Orthotomium) gabbi Crosse & Fischer.  Pilsbry, 1898; Man. Conch., 11: 147-148; pl. 19, figs. 58, 59; pl. 33, figs. 34, 35 (shell).

Naesiotus gabbi (Crosse & Fischer). Hoffman, 1988; Veliger, 30: 420.-  Smith, Miller, Christensen & Roth, 1990; Proc. Calif. Acad. Sci., 47: 115; text-fig. 19 (map).

 

  TYPE LOCALITY.-  “California Mexicana”.

 

  DISTRIBUTION RECORDS.-  BAJA CALIFORNIA SUR: rock outcrop E of rd. to La Paz, 4.2 km NE of junct. with rd. between El Gonzaga and El Obispo, 180 m alt.; along rd. between El Obispo and Rancho Tinajitas; arroyo nr. Microwave Station El Rifle, 120 m. alt.; arroyo 1.0 km NE of the trans-peninsular hwy. at km 77, N of La Paz (Smith, et al, 1990).

 

 

 

Naesiotus gigantensis (Christensen & Miller, 1977)

 

Rabdotus gigantensis Christensen & Miller, 1977; Nautilus, 91: 131-132; figs. 1-3 (shell).

Naesiotus gigantensis (Christensen & Miller). Hoffman, 1988; Veliger, 30: 420.-  Smith, Miller, Christensen & Roth, 1990; Proc. Calif. Acad. Sci., 47: 116; text-fig. 19 (map).

 

  TYPE LOCALITY.-  In a large lava rockslide immediately south of the mission at San Javier, Baja California Sur; 350-450 m alt.  Holotype CAS 57937.

 

  DISTRIBUTION RECORDS.-  BAJA CALIFORNIA SUR: nearly the entire length of the Sierra de la Giganta, numerous localities (Smith, et al, 1990).

 

 

Naesiotus hannai (Pilsbry, 1927)

 

Bulimulus hannai Pilsbry, 1927; Proc. Calif. Acad. Sci., (4), 17: 183-184; pl. 11, figs. 16-20 (shell); pl. 12, fig. 3 (sculpture).

Naesiotus hannai (Pilsbry). Hoffman, 1988; Veliger, 30: 420.-  Smith, Miller, Christensen & Roth, 1990; Proc. Calif. Acad. Sci., 47: 116; text-fig. 19 (map).

 

  TYPE LOCALITY.-  Isla Margarita, under stones within 1 mi. west and south of the village near the center of the east side of the island, Baja California Sur.  Holotype CAS 2621.

 

  DISTRIBUTION RECORDS.-  BAJA CALIFORNIA SUR: known only from Isla Margarita.  Ravine below Mt. Izabal; in rocky outcrops in hills just W of rd. from Puerto Alcatraz to Puerto Cortez (Smith, et al, 1990).

 

 

 

Naesiotus harribaueri (Jacobson, 1958)

 

Bulimulus (Puritanina) harribraueri Jacobson, 1958; Amer. Mus. Nov., (1899): 7-11; fig. 1 (shell), fig. 1 left (sculpture).

Naesiotus haribaueri (Jacobson). Hoffman, 1988; Veliger, 30: 420.-  Smith, Miller, Christensen & Roth, 1990; Proc. Calif. Acad. Sci., 47: 116; text-fig. 16 (map).

 

  TYPE LOCALITY.-  Sand dunes along Bahia de los Frailes, Baja California Sur.  Holotype AMNH 74003.

 

  DISTRIBUTION RECORDS.-  BAJA CALIFORNIA SUR: several localities along the east coast of Cabo San Lucas (Smith, et al, 1990).

 

 

 

Naesiotus laevapex (Christensen & Miller, 1977)

 

Rabdotus laevapex Christensen & Miller, 1977; Nautilus, 91: 132-133; figs. 4-7 (shell), fig. 7 (reproductive anatomy).

Naesiotus laevapex (Christensen & Miller).  Hoffman, 1988; Veliger, 30: 420.-  Smith, Miller, Christensen & Roth, 1990; Proc. Calif. Acad. Sci., 47: 116; text-fig. 16 (map).

 

  TYPE LOCALITY.-  In a narrow arroyo approximately 0.5 km inland of the beach at El Limoña, on the west side of Isla Cerrralvo, Baja California Sur; 50-100 m alt.  Holotype CAS 57942.

 

  DISTRIBUTION RECORDS.-  BAJA CALIFORNIA SUR: known only from the type locality.

 

 

 

Naesiotus milleri (Hoffman, 1987)

 

Rabdotus milleri Hoffman, 1987; Veliger, 20: 419-423; figs. 4c, 4d (shell), text-fig. 1 (map), text-fig. 3 (reproductive anatomy).

Naesiotus milleri (Hoffman).  Hoffman, 1988; Veliger, 30: 420.

 

  TYPE LOCALITY.-  Sonora, Mexico; 5.4 km west of the Yaqui River bridge at La Estrella on road to Sahuaripa, in a canyon extending north from road (28º57.1’ N, 109º36.7’ W); 350 m alt.  Holotype SBMNH 3490.

 

  DISTRIBUTION RECORDS.-  SONORA: known only from the vicinity of the type locality (Hoffman, 1987).

 

 

 

Naesiotus montezuma (Dall, 1893)

 

Bulimus proteus Binney, 1869; Smithsonian Miscellaneous Collections, 8: 207; fig. 358 (shell). (Not Bulinus proteus Broderip, 1832.)

Bulimulus (Scutalus) montezuma Dall, 1893; Proc. U. S. Nat. Mus.16: 16, 640; pl. 72, fig. 1.

Bulimulus (Orthotomium) montezuma Dall.  Pilsbry, 1898; Man. Conch., 11: 144-145; pl. 19, figs. 56, 57 (shell).

Bulimulus (Puritanina) montezuna Ball.  Jacobson, 1958: 10-11.

Naesiotus montezuma (Dall).  Hoffman, 1988; Veliger, 30: 420.-  Smith, Miller, Christensen & Roth, 1990; Proc. Calif. Acad. Sci., 47: 116-117; text-fig. 17 (map).

Bulimulus (Scutalus) acholus Mabille, 1895; Bulletin de la Société Philomathique de Paris, (8), 7: 68.-  Emerson & Jacobson, 1964; Trans. San Diego Soc. Nat. Hist., 16: 51.

 

  TYPE LOCALITIES.-  Bulimulus montezuma: Lower California, mostly from the mountainous region.  Lectotype USNM 8564 (Jacobson, 1958: 11).  Bulimulus acholus: “montagnes de la Base Californie”.

 

  DISTRIBUTION RECORDS.-  BAJA CALIFORNIA SUR: Cabo San Lucas, numerous localities from 60-2070 m alt. (Smith, et al, 1990).

 

 

Naesiotus nigromontanus (Dall, 1897)

 

Bulimulus (alternatus var. ?) nigromantanus Dall, 1897; Proc. U. S. Nat. Mus., 19: 357-358.

Rabdotus nigromontanus (Dall).  Hoffman, 1987: 419; text-fig. 1 (map); text-fig. 2 (reproductive anatomy), figs. 4a, 4b (shell).

Naesiotus nigromontanus (Dall, 1897).  Hoffman, 1988; Veliger, 30: 420.

 

  TYPE LOCALITY.-  Summit of Black Mountain, 12 mi. S of Monument 77 of International Boundary, right bank of San Bernardino River, Sonora [La Loma Colorado, 31º14.2’ N, 109º17.5’ W].  Holotype USNM 129993.

 

  DISTRIBUTION RECORDS.-  SONORA: La Loma Colorado; Angostura; Magdalena; Alamos (Hoffman, 1987).  ARIZONA, Santa Cruz Co.: Pena Blanca Canyon, Pajaritos Mts, ca. 4000 ft. alt. (Bequaert & Miller, 1973: 137).

 

 

 

Naesiotus pallidior (Sowerby, 1833)

 

Bulimus pallidior Sowerby, 1833; Proc. Zool. Soc. Lond.: 72.

Bulimulus (Orthotomium) pallidior (Sowerby).  Pilsbry, 1897; Man. Conch., 11: 142-143; pl. 51, figs. 53-55 (shell).

Naesiotus pallidior (Sowerby).  Hoffman, 1988; Veliger, 30: 420.-  Smith, Miller, Christensen & Roth, 1990; Proc. Calif. Acad. Sci., 47: 117-118; text-fig. 20 (map).

Bulimulus vegetus Gould, 1853: Jour. Boston Soc. Nat. Hist., 6: 375; pl. 14, fig. 2.

Bulimulus pallidior var. striatulus Dall, 1893: 640.-  Pilsbry, 1927; Proc. Calif. Acad. Sci., 16: 184-185; pl. 12, figs. 1, 7, 8 (shell).

Bulimulus vegetus vegexspiza Cooper, 1894; Proc. Calif. Acad. Sci., (2), 4: 134.

 

  TYPE LOCALITY.-  Not stated.

 

  DISTRIBUTION RECORDS.-  BAJA CALIFORNIA SUR: Cabo San Lucas, numerous localities from 60-760 m alt. (Smith, et al, 1990).

 

 

Naesiotus rimatus (Pfeiffer, 1846)

 

Bulimus rimatus Pfeiffer, 1846; Proc. Zool. Soc. London: 112.

Bulimulus (Sonorina) rimatus (Pfeiffer).  Pilsbry, 1898; Man. Conch., 11: 157; pl. 21, figs. 1-4 (shell).-  Hanna, 1923; Proc. Calf. Acad. Sci., (4), 12: 495.

Naesiotus rimatus (Pfeiffer).  Hoffman, 1988; Veliger, 30: 420.-  Smith, Miller, Christensen & Roth, 1990; Proc. Calif. Acad. Sci., 47: 117-118; text-fig. 20 (map).

Bulimulus (Mesembrinus) inscendens var. bryanti Cooper, 1891; Proc. Calif. Acad. Sci., Ser. II, 3: 340; pl. 13, figs. 4a-c (shell).

Bulimulus (Leptobrysus) bryanti (Cooper).  Dall, 1893; Proc. U. S. Nat. Mus., 16: 645; pl. 71, fig. 3 (shell).

Bulimulus (Leptobrysus) inscendens var, monticola Dall, 1893; Proc. U. S. Nat. Mus., 16: 643

 

  TYPE LOCALITIES.-  Bulimus rimatus: not given.  Bulimulus inscendens var. bryanti: San Jose del Cabo, Baja California Sur (restricted by Christensen, 1978).  Bulimulus inscendens monticolus: Sierra Laguna, Baja California Sur; 900 m alt. (restricted by Christensen, 1978).

 

  DISTRIBUTION RECORDS.-  BAJA CALIFORNIA SUR: numerous localities on the southern end of Cabo San Lucas below 150 m alt. (Smith, et al, 1990).

 

 

Naesiotus spirifer (Gabb, 1868)

 

Bulimus spirifer Gabb, 1868; Amer. Jour. Conch., 3: 236; pl. 16, fig. 5.

Bulimulus (Sonorina) spirifer (Gabb).  Pilsbry, 1898; Man. Conch., 11: 158-159; pl. 2, figs. 87-91 (shell).

Naesiotus spirifer (Gabb).  Hoffman, 1988; Veliger, 30: 420.-  Smith, Miller, Christensen & Roth, 1990; Proc. Calif. Acad. Sci., 47: 118-119; text-fig. 21 (map).

Bulimulus lapidivagus Mabille, 1895; Bulletin de la Société Philomathique de Paris, (8), 7: 66.-  Emerson & Jacobson, 1964; Trans. San Diego Soc. Nat. Hist., 16: 51.

 

  TYPE LOCALITIES.-  Bulimus spirifer: in the mountains, among rocks from San Antonio, below La Paz, to near San Borja, and in the mountains perhaps even farther north.  Bulimulus lapidivagus: Sierra de la Cachila au Sud de la Paz.

 

  DISTRIBUTION RECORDS.-  BAJA CALIFORNIA SUR: Cabo San Lucas, from various localities in the Sierra Cachila and near by areas; 430-690 m alt. (Smith, et al, 1990).

 

 

Naesiotus veseyianus (Dall, 1893)

 

Bulimulus (Leptobrysus) veseyianus Dall, 1893; Proc. U. S. Nat. Mus.16: 645; pl. 71, figs. 4, 5 (shell). 

Bulimulus (Sonorina) veseyianus Dall.  Pilsbry, 1898; Man. Conch., 11: 160; pl. 21, figs. 92, 93 (shell).-  Hanna, 1923; Proc. Calf. Acad. Sci., (4), 12: 499-500; pl. 8, figs. 1-3 (shell).

Rabdotus (Leptobrysus) veseyianus (Dall).  Emerson & Jacobson, 1964; Trans. San Diego Soc. Nat. Hist., 16: 324.

Naesiotus veseyianus (Dall).  Hoffman, 1988; Veliger, 30: 420.-  Smith, Miller, Christensen & Roth, 1990; Proc. Calif. Acad. Sci., 47: 119; text-fig. 19 (map).

 

  TYPE LOCALITY.-  Isla Espiritu Santo, Baja California Sur.  Holotype USNM 34122.

 

  DISTRIBUTION RECORDS.-  BAJA CALIFORNIA SUR: known only from Isla Espiritu Santo (Smith, et al. 1990).

 

 

Naesiotus xantusi (Binney, 1861)

 

Bulimus xanttusi W. G. Binney, 1861; Proc. Acad. Nat. Sci. Phila.: 331; fig. (shell).

Bulimulus (Orthotomium) xantusi (Binney).   Pilsbry, 1898; Man. Conch., 11: 148; pl. 19, figs. 60-62; pl. 33, fig. 36 (shell).

Naesiotus xantusi (Binney).  Hoffman, 1988; Veliger, 30: 420.-  Smith, Miller, Christensen & Roth, 1990; Proc. Calif. Acad. Sci., 47: 119; text-fig. 22 (map).

 

  TYPE LOCALITY.-  Cabo San Lucas, Baja California Sur.

 

  DISTRIBUTION RECORDS.-  BAJA CALIFORNIA SUR: various localities on the Cabo San Lucas, from 90-910 m alt. (Smith, et al, 1990). 

 

 

Genus Rabdotus Albers, 1850

 

Rabdotus Albers, 1850; Die Heliceen: 164.-  Pilsbry, 1946; Land Moll. N. Amer., 2: 4.

 

  DISTRIBUTION.-  Central Mexico from Puebla north to Texas, the central-southern United States, and Baja California Sur.

 

  TAXONOMY.-  Two subgenera and sixteen species are recognized.

 

 

Subgenus Rabdotus (Albers, 1850

 

Rabdotus Albers, 1850; Die Heliceen: 164.

Odontostomus Fischer & Crosse, 1874; Miss. Sci. Mex., I: 473.

Globulinus Fischer & Crosse, 1874; Miss. Sci. Mex., I: 475.

 

  TYPE SPECIES.-  Rabdotus: Helix dealbatus Say, 1821.  Odontostomus: Bulimus sFLMNHflatus Gould, 1859.  Globulinus: Bulimus sFLMNHflatus Gould, 1859.

 

  DISTRIBUTION.-  The same as for the genus.

 

  TAXONOMY.-  Eleven species and one subspecies occur in the study area.

 

 

Group of Rabdotus alternatus

 

 

Rabdotus (Rabdotus) alternatus alternatus (Say, 1830)

 

Bulimus alternatus Say, 1830; New Harmony Disseminator: 25.

Bulimulus alternatus alternatus (Say, 1830).  Pilsbry, 1928: 117.-  Pilsbry, 1946; Land Moll. N. Amer., 2: 15-16; fig. 6e.-  Branson & McCoy, 1963; Nautilus, 76: 105.

Rabdotus alternatus (Say).  Correa-Sandoval, 1993; Rev. Biol. Trop., 41: 675.-  Correa-Sandoval, Gutierrez & Reza, 1998: 17.-  Correa-Sandoval, 2000; Acta Zool.Mex., (79):: 9.-  Correa-Sandoval & Castro, 2002; Acta Zool. Mex., (86): 238.

Bulimus lactarius Menke, 1846; in Pfeiffer, Symbolae ad Histori Heliceorum, 3: 85.

 

  Type locality.-  Monterrey, Nuevo León (restricted by Pilsbry, 1946).  Holotype ANSP 25694.

 

  DISTRIBUTION RECORDS.-  NUEVO LEÓN: Santa Caterina Valley opposite and above the Obispada (Pilsbry, 1946); Santiago (Correa-Sandoval-Sadoval, 1993).  SAN LUIS POTOSI: 0.1 km Ejido Buenavista km 18 Carr. Mante-Cd. Valles (22°30’43” N, 99°01’50” W, 320 m alt.); Km 92, W of Cd. del Maíz (22°25’36” N, 99°38’09” W, 1260 m alt.); Ejido Buena Vista, 2 km S of Carr. Antiguo Morelos-Cd. Valles (22°16’49” N, 99°02’22” W, 300 m alt.); km 169 on Carretera Río Verde-San Luis Potosí (21°59’30” N, 100°11’50” W); Las Cascadas, Tamasopo (21°56’05” N, 99°25’00” W) (Correa-Sandoval, et al, 1998); 2 mi. SE of Charro Blanco Branson & McCoy, 1963).  TAMAULIPAS: Mesa de Solis, nr. La Lajilla, between Padilla and Jimenez (Pilsbry, 1928); numerous records (Correa-Sandoval & Castro, 2002).  VERACRUZ: La Ordeña, Papantla (20° 29’ 43”N, 97° 18’ 27” W) (Correa-Sandoval, 2000).

 

 

 

Rabdotus (Rabdotus) alternatus mariae (Albers, 1850)

 

Bulimus mariae Albers, 1850; Die Heliceen: 162.

Bulimulus alternatus mariae (Albers). Pilsbry & Ferriss, 1906; Proc. Acad. Nat. Sci. Phila., 58: 139; pl.7, figs. 13-30.

Bulimulus (Rabdotus) alternatus mariae (Albers).  Pilsbry, 1946; Land Moll. N. Amer., 2: 14-15; figs. 6a-d, f (shell).

 

  TYPE LOCALIY.-  Brownsville, Texas (restricted by Pilsbry, 1946).  Syntype ZMB 112669 (Köhler, 2007).

 

  DISTRIBUTION RECORDS.-  Many localities in northern Nuevo León and Tamaulipas (Pilsbry, 1946).

 

 

 

Rabdotus (Rabdotus) dealbatus dealbatus (Say, 1821)

 

Helix dealbaus ay, 1821; Journ. Acad. Nat. Sci. Phila., 2: 150.

Bulimulus dealbatus (Say).  W. G. Binney, 1878; Terrestrial Mollusks of North America, 5: 393; fig. 269 (shell); pl. 8, figs. E (jaw, radula).

Bulimulus (Rabdotus) dealbatus (Say).  Pilsbry, 1946; Land Moll.N. Amer., 2: 7-9; figs. a-d.

Rabdotus dealbatus dealbatus (Say).  Correa-Sandoval, 1993; Rev. Biol. Trop., 41: 675.-   Correa-Sandoval & Castro, 2002; Acta Zool. Mex., (86): 238.

Bulimus liquabilis Reeve, Conch. Icon., 5: pl. 57, fig. 384 (shell).

Bulimus confinis Reeve, 1850; Conch. Icon., 5: pl. 86, fig. 643 (shell).

 

   TYPE LOCALITIES.-  Helix dealbatus: Missouri and Alabama.  Holotype in the ANSP (Pilsbry, 1946).  Bulimus liquabilis: Texas; lectotype BMNH 1975422 (Breure, 1979; Zool. Verhandl. Uit. Rijkmuseum Nat. Hist. Leiden (168): 76).  Bulimus confinis: Texas.

 

   DISTRIBUTION RECORDS.-  Widely distributed in the eastern United States from Illinois south to Alabama and west to eastern Texas and Oklahoma.  Mexican records are as follow.  NUEVO LEÓN: Santiago (Correa-Sandoval, 1993; Rev. Biol. Trop., 41:675).  SAN LUÍS POTOSÍ: Valles (Hinkley, 1907).  TAMAULIPAS: numerous localities (Correa-Sandoval & Castro, 2002).

 

  TAXONOMY.-  Six subspecies are recognized, only one of which occurs in Mexico.

 

 

 

Rabdotus (Rabdotus) pilsbryi (Ferriss, 1925)

 

Bulimulus pilsbryi Ferriss, 1925; Nautilus, 30: 25.

Bulimulus (Rabdotus) pilsbryi (Ferriss).  Pilsbry, 1946; Land Mollua of North America, 2: 20-21, figs. 10a-c (shell).

 

  TYPE LOCALITY.-  Sanderson, Terrell Co., Texas.  Holotype ANSP 175851.

 

  DISTRIBUTION RECORDS.-  Southwestern Texas, Terrell County.  In Mexico this species has been collected at the following localities.  CHIHUAHUA: near Coyame (29º44.5’ N, 104º54.0’ W), 1100 m alt. (FLMNH 251521); pass at NW end of Sierra Matasagnas, ca. 50 km W of Ojinaga, 9.6 km E of turn-off to Alamos on hwy, 16 ((29º02.1’ N, 104º47.6’ W), 1000 m alt. (FLMNH 317157).  COAHUILA: 5 KM NW of Quatro Cienegas, 800 m slt. (FLMNH 267670); limestone hill 18.7 km E of Quatro Cienegas, 800 m alt. (FLMNH 178584); 3 km N of Quatro Cienegas, 870 m alt. (FLMNH 39968); NW foothills of the Sierra la Madera, 9.3 km SE, 14.7 km SW of Ocampo, 1200 m alt. (27º11.6’ N, 102º25.4’ W ) (FLMNH 267667); ridge 2.5 km E of Paredón, 780 m alt. (26º56.3’ N, 100º55,0’ W) (FLMNH 267692); Cerro Sesteadero 11 km S of San Miguel, 1000 m alt. (28º32.9’ N, 102º56.1’ W) (FLMNH 268362).  NUEVO LEÓN: canyon 6.5 km by road N of Rinconada, 1050 m alt. (25º42.5’ N, 100º43.1’ N) (FLMNH 268303).

 

 

 

Rabdotus (Rabdotus) schiedeanus schiedeanus (Pfeiffer, 1841)

 

Bulimus schiedeanus Pfeiffer, 1841; Symbolae ad Histori Heliceorum, 1: 43.

Bulimulus schiedeans (Pfeiffer).  Von Martens, 1893; Biol. Cent. Amer.: 239; pl. 15, figs. 12-23 (shell).-  Pilsbry, 1897; Man. Conch., 11: 131.-  Hinkley, 1907; Nautilus, 21: 72.

Bulimulus (Rabdotus) schiedeanus (Pfeiffer).  Pilsbry, 1946; Land Moll. N. Amer., 2: 16-17; figs. 7a-d (shell).-  Solem, 1954: 4.-  Branson & McCoy, 1963; Nautilus, 76: 105.

Bulimus patriarcha W. G. Binney, 1858; Manual of North American land shells: 396; fig. 431 (shell).

Bulimulus alternatus hesperius Pilsbry & Ferriss, 1924; Nautilus, 38: 40.

 

  TYPE LOCALITIES.-  Bulimus schiedeanus:  Mexico.  Bulimus patriarcha: Buena Vista, [south of Saltillo, Coahuila].  Bulimulus alternatus hesperius: east of the Pecos at the High Bridge, Val Verde Co., Texas; holotype in the ANSP.

 

  DISTRIBUTION RECORDS.-  Texas, counties along the Rio Grande from Val Verde County to Presidio County; widely distributed but sparse in Mexico from the state of Puebla north to Chihuahua and Coahuila.  CHIHUAHUA: La India, 7 mi. E of Escalón (Branson & McCoy, 1963).  COAHUILA: around Saltillo (Pilsbry, 1946).  DURANGO:  Villa Lerdo (Pilsbry, 1946).  Sierra de Tlahualilo, 25 mi. N of Tlahualilo (Solem, 1954: 4).  JALISCO: Laguna de Chapala (Pilsbry, 1946).  PUEBLA: Tahuacan (Pislbry, 1946). TAMAULIPAS: Tampico, in river drift (Hinkley, 1907)

 

  TAXONOMY.-  Two subspecies are recognized.  Rabdotus schiedeanus pecosensis Pilsbry & Ferriss, 1906, is confined to southwest Texas.

 

 

 

Rabdotus (Rabdotus) fonsecanus (Haas, 1961)

 

Bulimulus (Rabdotus) fonsecanus Haas, 1961; Fieldiana, Zoology, 44: 20-21; fig. 11 (shell).

 

  TYPE LOCALITY.-  Gulf of Fonseca, El Salvador or Nicaragua.  Holotype: FMNH 106702.

 

  DISTRIBUTION RECORDS.-  Known only from the type locality.  The generic affinities of this species require further investigation.

 

 

Group of Rabdotus sFLMNHflatus

 

 

Rabdotus (Rabdotus) baileyi (Dall, 1893)

 

Bulimulus (Scutalus) baileyi Dall, 1893; Proc. U. S. Nat. Mus.16: 640-641; pl. 71, fig. 1 (shell).

Bulimulus (Orthotomium) baileyi Dall.  Pilsbry, 1898; Man. Conch., 11: 145-146; pl. 19, figs. 65-67.

Bulimulus baileyi Dall.  Hanna, 1923; Proc. Calf. Acad. Sci., (4), 12: 487.

 

  TYPE LOCALITY.-  Cabo San Lucas, Baja California Sur.  Holotype in the USNM.  (Doubtful locality, fide Hanna, 1923; Proc. Calf. Acad. Sci., (4), 12: 487.)

 

  DISTRIBUTION RECORDS.-  SONORA: Guaymas; Hermosillo; Cerrro Tordilla, between Guaymas and San Marciel; Ortiz (Pilsbry, 1898).

 

 

 

Rabdotus (Rabdotus) ceralboensis (Hanna, 1923)

 

Bulimulus ceralboensis Hanna, 1923; Proc. Calif. Acad. Sci., 12: 490; pl. 7, fig. 11; pl. 11, figs. 2, 4 (shell).

Rabdotus (Rabdotus) ceralboensis  (Hanna).  Emerson & Jacobson, 1964; Trans. San Diego Soc. Nat. Hist., 16: 327.-  Smith, Miller, Christensen & Roth, 1990; Proc. Calif. Acad. Sci., 47: 120; text-fig. 20 (map).

 

  TYPE LOCALITY.-  West of Ranch house, Cerralvo Island, Baja California Sur.  Holotype CAS 1021.

 

  DISTRIBUTION RECORDS.-  BAJA CALIFORNIA SUR: known only from Cerralvo Island (Smith, et al, 1990).

 

 

 

Rabdotus (Rabdotus) chamberlini (Hanna, 1923)

 

Bulimulus chamberlini Hanna, 1923; Proc. Calif. Acad. Sci., 12: 494; pl. 8, fig. 21 (shell).

Rabdotus (Leptobrysus) chamberini (Hanna).  Emerson & Jacobson, 1964; Trans. San Diego Soc. Nat. Hist., 16: 318-319; figs. 1a-f (shell).

Rabdotus (Rabdotus) chamberlini (Hanna, 1923).  Smith, Miller, Christensen & Roth, 1990; Proc. Calif. Acad. Sci., 47: 120; text-fig. 23 (map).

 

  TYPE LOCALITY.-  San Diego Island, Baja California Sur.  Holotype CAS 1023.

 

  DISTRIBUTION RECORDS.-  BAJA CALIFORNIA SUR: known only from Isla San Diego (Smith, et al, 1990).

 

 

 

Rabdotus (Rabdotus) levis (Dall, 1893)

 

Bulimulus xantusi var. levis Dall, 1893; Proc. U. S. Nat. Mus., 16: 642.

Bulimulus levis Dall.  Dall, 1895; Proc. U. S. Nat. Mus., 18: 5.

Bulimulus (Orthotomium) levis Dall.  Pilsbry, 1897: Man. Conch., 11: 140-141; pl. 19, fig. 64 (shell).

Rabdotus (Rabdotus) levis (Dall).  Smith, Miller, Christensen & Roth, 1990; Proc. Calif. Acad. Sci., 47: 120-121; text-fig. 22 (map).

Bulimulus (Thaumastus) digueti Mabille, 1895.

 

 TYPE LOCALITIES.-  Bulimulus levis: not specified.                        Bulimulus digueti: Sierra de La Victoria, Baja California Sur.

  DISTRIBUTION RECORDS.-  BAJA CALIFORNIA SUR: various localities near southern tip of peninsula (Smith, et al, 1990).

 

 

 

Rabdotus (Rabdotus) pilula (Binney, 1861)

 

Bulimulus pilula W. G. Binney, 1881; Proc. Acad. Nat. Sci. Phila., 332; fig. (shell).

Bulimulus (Orthotomium) pilula Binney.  Pilsbry, 1897; Man. Conch., 11: 138-139; pl. 25, fig. 57 (shell).

Rabdotus (Rabdotus) pilula (Binney, 1861).  Smith, Miller, Christensen & Roth, 1990; Proc. Calif. Acad. Sci., 47: 121; text-fig. 22 (map).

Bulimulus cooperi Dall, 1896; Proc. U. S. Nat. Mus.18: 5.

Bulimulus (Orthotomium) decipiens Cooper, 1895; Proc. Calif. Acad. Sci. (2), 5: 164.-  Dall, 1896; Proc. U. S. Nat. Mus.19: 358-459.

 

  TYPE LOCALITIES.-  Bulimulus pilula: Todos Santos Mission, Baja California Sur (Binney & Bland, 1869).  Bulimulus decipiens: Sierra San Lazaro, Cabo San Lucas.  Bulimulus cooperi: San Jose del Cabo.

 

  DISTRIBUTION RECORDS.-  BAJA CALIFORNIA SUR: along the southern tip of the peninsula from Cabo San Lucas to San Jose del Cabo (Smith, et al, 1990).

 

 

 

Rabdotus (Rabdotus) sFLMNHflatus  (Gould, 1859)

 

Bulimus sFLMNHflatus Gould, 1859; Journal of the Boston Society of Natural History, 6: 375; pl. 14, fig. 1 (shell).

Bulimulus (Orthotomium) sFLMNHflatus (Gould).  Pilsbry, 1897; Man. Conch, 11: 136-137; pl. 18, figs. 38-44 (shell).

Rabdotus(Rabdotus) sFLMNHflatus (Gould, 1859).  Smith, Miller & Christensen, 1990: 121-122; text-fig. 23 (map).

Bulimus vesicalis Gould, 1853 (not Bulimus vesicalis Pfeiffer, 1859.

Bulimulus juarezi Pfeiffer, 1966; Proc. ool. Soc. London (1865): 832.

Bulimulus sFLMNHflatus var. insularis Cooper, 1892; Proc. Calif Acad. Sci., (2), 3: 208, 212.

Bulimulus insularis var. chinchesis Cooper, 1894; Proc. Calif. Acad. Sci., (2), 4: 140.

Bulimulus (Globulus) recognitus Mabille, 1895; Bulletin de la Société Philomathique de Paris, (8), 7: 69.-  Emerson & Jacobson, 1964; Trans. San Diego Soc. Nat. Hist., 16: 51.-  Breure, 1976: 1148 (measurements of holotype).

 

  TYPE LOCALITIES.-  Bulimus sFLMNHflatus: not stated.  Bulimulus juarezi: Mexico; syntype in the BMNH (Breure, 1979; Zool. Verhandl. Uit. Rijkmuseum Nat. Hist. Leiden (168): 76).  Bulimulus sFLMNHflatus var. insularis:  Espiritu Santo Island, Baja California Sur.  Bulimulus insularis chinchensis: El Chinche Mts., Baja California Sur. Bulimulus (Globulus) recognitus: Mexico, Baja California; lectotype in the MNHN (Breure, 1979; Zool. Verhandl. Uit. Rijkmuseum Nat. Hist. Leiden (168): 76).

 

  DISTRIBUTION RECORDS.-  BAJA CALIFORNIA SUR: numerous localities on the south end of the peninsula (Smith, et al, 1990).

 

 

 

Subgenus Plicolumna Cooper, 1895

 

Plicolumna Cooper, 1895; Proc. Calif. Acad. Nat. Sci. (2), 5: 164.-  Smith, Miller, Christensen & Roth, 1990; Proc. Calif. Acad. Sci., 47: 101, 122-124.

 

  TYPE SPECIES.-  Rhodea californica ramentosa Cooper, 1895.

 

  DISTRIBUTION.-  Endemic to the Cape region of Baja California Sur.

 

  TAXONOMY.-  Five species are recognized (Smith, et al, 1990).

 

 

 

 

Rabdotus (Plicolumna) abbreviatus (Cooper, 1892)

 

Columna ramentosa var. abbreviata Cooper, 1892; Proc. Calif. Acad. Sci., (2), 3: 215.

Bulimulus (Plicolumna) abbreviata (Cooper).  Pilsbry, 1898; Man. Conch., 11: 153; pl. 20, figs. 80, 81 (shell).

Rabdotus (Plicolumna) abbreviatus (Cooper).  Smith, Miller, Christensen & Roth, 1990; Proc. Calif. Acad. Sci., 47: 122; text-fig. 24 (map).

 

  TYPE LOCALITY.-  Sierra Laguna, Baja California Sur.

 

  DISTRIBUTION RECORDS.-  BAJA CALIFORNIA SUR: known from three localities in the Cape region (Smith, et al, 1990).

 

 

 

Rabdotus (Plicolumna) artemisia (Binney, 1861)

 

Bulimus artemisia W. G. Binney, 1861; Proc. Acad. Nat. Sci. Phila.: 33; fig. (shell).

Bulimulus artemesia (Binney).   Dall, 1893; Proceedings of the U.S. National Museum, 16: 642-643; pl. 72; fig. 5 (shell).-  Dall, 1896: 360-363; pl. 31, fig. 6; pl. 32, fig. D (shell).

Bulimulus (Plicolumna) artemisia (Binney).  Pilsbry, 1898; Man. Conch., 11: 152-153; pl. 20, figs. 77-79; pl. 25, fig. 54 (shell).

Rabdotus (Plicolumna) artemisia (Binney, 1861).  Smith, Miller, Christensen & Roth, 1990; Proc. Calif. Acad. Sci., 47: 122-123; text-fig. 25 (map).

 

  TYPE LOCALITY.-  Cabo San Lucas, Baja California Sur.

 

  DISTRIBUTION RECORDS.-  BAJA CALIFORNIA SUR: Cape region of the peninsula (Smith, et al, 1990).

 

 

 

Rabdotus (Plicolumna) inscendens (Binney, 1861)

 

Bulimus inscendens W. G. Binney, 1861; Proc. Acad. Nat. Sci. Phila.: 332; fig. (shell).

Bulimulus (Orthotomium) inscendens (Binney).  Pilsbry, 1898; Man. Conch., 11: 150-151; pl. 20, figs. 72-74 (shell).

Rabdotus (Plicolumna) inscendens (Binney).  Emerson & Jacobson, 1964; Trans. San Diego Soc. Nat. Hist., 16: 44, 45; text-fig. 4 (reproductive anatomy).-  Smith, Miller, Christensen & Roth, 1990; Proc. Calif. Acad. Sci., 47: 123-124; text-fig. 26 (map).

 

 

  TYPE LOCALITY.-  Not stated.

 

  DISTRIBUTION RECORDS.-  BAJA CALIFORNIA SUR: numerous localities in the Cape region (Smith, et al, 1990).

 

 

 

Rabdotus (Plicolumna) perhirsutus Miller, Christensen & Roth, 1990

 

Rabdotus (Plicolumna) perhirsutus Miller, Christensen & Roth, 1990, in Smith, Miller, Christensen & Roth. 1990: 101-102, 124; figs. 1-3 (shell), fig. 4 (reproductive anatomy), text-fig. 25 (map).

 

  TYPE LOCALITY.-  Along Todos Santos-Cabo San Lucas road 2.2 mi. (2.2 km) N of Saucito, ca. 1500 ft. (460 m) alt. Holotype SBMNH 35110.

 

  DISTRIBUTION RECORDS.-  BAJA CALIFORNIA SUR: known only from the vicinity of the type locality (Smith, et al, 1990).

 

 

 

Rabdotus (Plicolumna) ramentosus (Cooper, 1991)

 

 

Rhodea californica subsp. ? ramentosa Cooper, 1891; Proc. Calif. Acad. Sci., (2), 3: 102.

Bulimulus (Plicolumna) ramentosus (Cooper).  Pilsbry, 1898; Man. Conch., 11: 153-154; pl. 20, figs. 82-85 (shell).

Rabdotus (Plicolumna) ramentosus (Cooper).  Christensen & Miller, 1975: 45; fig. 7 (shell).-  Smith, Miller, Christensen, & Roth, 1990: 124; text-fig. 24 (map).

 

  TYPE LOCALITY.-  Mountains north of San Jose del Cabo, Baja California Sur.

 

  DISTRIBUTION RECORDS.-  BAJA CALIFORNIA SUR:  known from five localities in the Cape region (Smith, et al, 1990).

 

 

 

Genus Spartocentrum Dall, 1895

 

Spartocentrum Dall, 1895; Nautilus, 9: 51.-  Pilsbry, 1902; Man. Conch., 15: 15.-  Christensen & Miller, 1975; Nautilus, 89:  44-46.

Teneritia Mabille, 1897; Bulletin de la Société Philomathique de Paris, (8), 9: 78.

 

  TYPE SPECIES.-  Spartocentrum: Cylindrella irregulare Gabb, 1868.  Teneritia: Berendtia digueti Mabille, 1895,

 

  DISTRIBUTION.-  Endemic to the mainland of Baja California Sur and adjacent islands in the Gulf of California.

 

  TAXONOMY.-  Five species are recognized.

 

 

 

Spartocentrum digueti (Mabille, 1895)

 

Bulimulus (Thaumastus) digueti Mabille, 1895; Bulletin de la Société Philomathique de Paris, (8), 7: 70.

Bulimulus (Orthotomium) digueti Mabille.   Pilsbry, 1898; Man. Conch., 11: 148-149.

Spartocentrum digueti (Mabille).  Hanna & Smith, 1968: 396.-  Breure, 1978; Uitg. Rijkmuseum Nat. Hist. Leiden (164): 155-158; figs. 256-258 (shell).-  Smith, Miller, Christensen & Roth, 1990; Proc. Calif. Acad. Sci., 47: 125.

 

  TYPE LOCALITY.-  Plateau de San Javier, [Baja California Sur] (ca 25º50’ N, 111º30’ W).  Lectotype in the MNHN (Hanna & Smith, 1968; Breure, 1978; Uitg. Rijkmuseum Nat. Hist. Leiden (164): 155).

 

  DISTRIBUTION RECORDS.-  BAJA CALIFORNIA SUR: not known from a definite locality (Smith, et al, 1990).

 

 

 

Spartocentrum eisenianum (Pilsbry, 1900)

 

Coelocentrum eisenianum Pilsbry, 1900; Proc. Acad. Nat. Sci. Phila., 52; 553,; fig. 2 (shell).-  Pilsbry, 1903; Man. Conch., 15: 55-56; pl. 13, figs. 20, 21, 22; pl. 17, fig. 25 (shell).

Spartocentrum eisenianum (Pilsbry).  Smith, Miller, Christensen & Roth, 1990; Proc. Calif. Acad. Sci., 47: 125. 

Coelocentrum eiseni Bartsch, 1907: 119.

 

  TYPE LOCALITY.-  Lower California.  Holotype ANSP 77888.

 

  DISTRIBUTION RECORDS.-  BAJA CALIFORNIA SUR: not known from a definite locality (Smith, et al, 1990).

 

 

 

Spartocentrum insulare (Hanna, 1923)

 

Coelocentrum insulare Hanna, 1923; Proc. Calif. Acad. Sci., 12: 509-511; pl. 9, figs. 9-18 (shell); pl. 111, fig. 5 (juvenile shell).

Spartocentrum insulare (Hanna).  Smith, Miller, Christensen & Roth, 1990; Proc. Calif. Acad. Sci., 47: 125; text-fig. 28 (map).

Coelocentrum oweni Hanna, 1923; 511-512; pl. 9, figs. 3-8 (shell); pl. 11, fig. 8 (juvenile shell).-  Emerson & Jacobson, 1964; Trans. San Diego Soc. Nat. Hist., 16: 328-329.

 

  TYPE LOCALITIES.-  Coelocentrum insulare: Puerto Ballandra, Carmen Island, Baja California Sur.  Holotype CAS 1061.  Coelocentrum oweni: Agua Grande, Isla Carmen, Baja California Sur.  Holotype CAS 1055.

 

  DISTRIBUTION RECORDS.-  BAJA CALIFORNIA SUR: Isla Carmen; Isla Montserrate; Isla Danzante; Isla Santa Catalina (Smith, et al, 1990).

 

 

 

Spartocentrum irregularis (Gabb, 1868)

 

Cylindrella (Urocoptis) irregularis Gabb, 1868: 234; pl. 16, fig. 4 (shell).

Coelocentrum irregularis (Gabb).  Pilsbry, 1903; Man. Conch., 15: 51; pl. 13, figs. 15, 16 (shell), pl. 17, fig. 24 (shell).-  Hanna, 1923; Proc. Calf. Acad. Sci., (4), 12: 508; pl. 9, figs. 1, 2 (shell).

Spartocentrum irregulare (Gabb).  Breure, 1979; Zool. Verhandl. Uit. Rijkmuseum Nat. Hist. Leiden (168): 79.-  Smith, Miller, Christensen & Roth, 1990; Proc. Calif. Acad. Sci., 47: 125-126; text-fig. 28 (map).

Coelocentrum minorinum gabbi Pilsbry, 1900; Proc. Acad. Nat. Sci. Phila., 551.-  Pilsbry, 1903; Man. Conch., 15: 54; pl. 13, figs. 17-19 (shell).

Berendtia minorina Mabille, 1895; Bulletin de la Société Philomathique de Paris, (8), 7: 70.

Spartocentrum minorina (Mabille).  Hanna & Smith, 1968; Proc. calif. Acad. Sci., (4), 30.-  Breure, 1978; Uitg. Rijkmuseum Nat. Hist. Leiden (164): 158; figs. 259, 260 (shell).

Coelocentrum clavigeroi Hanna, 1923; Proc. Calf. Acad. Sci., (4), 12: 512-513; pl. 9, figs. 20-30 (shell); pl. 1, fig. 6 (juvenile shell).

 

  TYPE LOCALITIES.-  Cylindrella irregularis: table lands in the interior of Lower California, especially above Moleje [Mulege]; lectotype ANSP 25075 (H. B. Baker, 1963: 223).  Berendtia minorina: “Au dessus de l’arroyo de Purisima”, [Baja California sur] (ca. 26º15’ N, 112º W); lectotype in the MNHN (Hanna & Smith, 1968).  Coelocentrum minorinum gabbi: table lands in the interior of Lower California, especially above Moleje [Mulege]; holotype ANSP 25077a (Baker, 1963: 222).  Coelocentrum clavigeroi: Agua Verde Bay, Baja California Sur; holotype CAS 1082.

 

  DISTRIBUTION RECORDS.-  BAJA CALIFORNIA SUR: numerous localities in the central third of the peninsula (Smith, et al, 1990).

 

 

 

Spartocentrum vanduzeei (Hanna, 1923)

 

Coelocentrum vanduzeei Hanna, 1923; California Academy of Sciences, 12: 508-509; pl. 9, figs. 31-34 (shell); pl. 11, fig. 7 (juvenile shell).

Spartocentrum vanduzeei (Hanna).  Christensen & Miller, 1975; Nautilus,89: 44-46; text-fig. 2 (reproductive anatomy), fig. 6 (shell).-  Smith, Miller, Christensen & Roth, 1990; Proc. Calif. Acad. Sci., 47: 126-127; text-fig. 28 (map).

 

  TYPE LOCALITY.-  Puerto Escondido, Baja California Sur.  Holotype CAS 1085.

 

  DISTRIBUTION RECORDS.-  BAJA CALIFORNIA SUR: Puerto Escondido; Juncalito; 1.6 km S of Chunque (Smith, et al, 1990.

 

 

 

Genus Simpulopsis Beck, 1838

 

Simpulopsis Beck, 1838; Index Molluscorum.: 100.-  Von Martens, 1897; Biol. Cent. Amer.: 252.-  Pilsbry, 1899; Man. Conch., 12: 212.

 

  TYPE SPECIES.-  Helix (Cochlohydra) sulculosa Ferussac.

 

  DISTRIBUTION.- South America, Trinidad and eastern Mexico.

 

  TAXONOMY.-   Three species occur in the study area.  The taxonomic affinities of the Middle American species are uncertain.

 

 

 

Simpulopsis aenea Pfeiffer, 1861

 

Simpulopsis aenea Pfeiffer, 1861; Proc. Zool. Soc. London: 27.-  Fischer & Crosse, 1877; Miss. Sci. Mex., I: 580; pl. 24, figs. 12, 12a, 12b (shell).-  Von Martens, 1897; 253.

Simpulopsis (Platysuccinea) aenea Pilsbry, 1899; Man. Conch., 12: 225-226; pl. 63, figs. 58, 59, 60 (shell).

 

  TYPE LOCALITY.-  Parada, Oazaca.

 

  DISTRIBUTION RECORDS.-  OAXACA: known only from the type locality.

 

 

Simpulopsis cumingi Pfeiffer, 1861

 

Simpulopsis cumingi Pfeiffer, 1861; Proc. Zool. Soc. London: 27; pl. 3, fig. 2 (shell).-  Von Martens, 1897; Biol. Cent. Amer.: 253.-  Pilsbry, 1899; Man. Conch., 12: 220; pl. 63, figs. 61, 62 (shell).

 

  TYPE LOCALITY.-  Mexico.

 

  DISTRIBUTION RECORDS.-  MEXICO (?).

 

 

Simpulopsis simula (Morelet, 1951)

 

Bulimus simula Morelet, 1951; Test. Noviss.: 11.

Simpulopsis simula (Morelet).  Fischer & Crosse, 1877; Miss. Sci. Mex., I: 576; pl. 24, figs. 13, 13a (shell).-  Von Martens, 1897; Biol. Cent. Amer.: 252.-  Pilsbry, 1899; Man. Conch., 12: 219-220; pl. 63, figs. 56, 57 (shell).-  Bequaert, 1957: 221-222.-  Breure, 1979; Zool. Verhandl. Uit. Rijkmuseum Nat. Hist. Leiden (168): 134.

 

  TYPE LOCALITY.-  Petén, Guatemala.

 

  DISTRIBUTION RECORDS.-  HONDURAS, Dept. Olancho: vicinity of Magua Cave, 940 m alt. (Breure, in press).  GUATEMALA, Dept.Peten: known only from the type locality.  Dept. Alta Verapaz: 15 km by road north of Cobán, 1050 m alt.; 3 km W of Pajál, 1130 m alt.; 11 km W of Lanquín, 1000 m alt. (Breure, in press).  CHIAPAS: Ocotal [15°39'00" N, 92°00'00" W] to El Censo (on the border with Guatemala), 700-1000 m alt. (Bequaert, 1957).

 

 

 

 

Family UROCOPTIDAE Pilsbry, 1898

 

Subfamily EUCALODIINAE Fischer & Crosse, 1963

 

  DISTRIBUTION.-  Mexico, Gatemala, Honduras and Belize. 

 

  TAXONOMY.-  Five genera are included in the Eucalodiinae.  Archegocoptis from Hispaniola may belong here also.

 

 

Genus Eucalodium Crosse & Fischer, 1868

 

Eucalodium Crosse & Fischer, 1868; Jour. de Conchyl., 16: 88.

 

  TYPE SPECIES.-  Pupa decollata Nyst, 1841.

 

  DISTRIBUTION RECORDS.-  El Salvador and Guatemala north and west in México to Jalisco along the west coast and San Luís Potosí along the east coast.

 

  TAXONOMY.-  Four subgenera, 24 species and 9 subspecies are recognized.

 

 

Subgenus Eucalodium Crosse & Fischer, 1868

 

  DISTRIBUTION.-  Tabasco, Chiapas and Oaxaca, México; Guatemala.

 

  TAXONOMY.-  Four species and five subspecies occur in the study area.

 

 

Eucalodium (Eucalodium) compactum Pilsbry, 1893

 

Eucalodium compactum Pilsbry, 1893.  Proc. Acad. Nat. Sci. Phila.: 338; pl. 14, fig. 4.-  Pilsbry, 1902; Manual of Conchology, 15:  5-6; pl. 7, figs. 1-3.

 

  TYPE LOCALITY.-  Tabasco, México. Holotype ANSP 63388a (H. B. Baker, 1963).

 

  DISTRIBUTION RECORDS.- TABASCO: known only from the type locality.

 

 

Eucalodium (Eucalodium) decollatum decollatum (Nyst, 1841)

 

Pupa decollata Nyst, 1841; Bulletin de l’Académie Royal de Bruxelles, 8: 345.

Cylindrella decollata (Nyst).  Pfeiffer, 1856; Mon. helic. viv., VIII: 421.

Eucalodiun decollatum (Nyst).  Fischer & Crosse, 1972; Miss. Sci. Mex., I: 362; pl. 14, figs. 3-3a (shell).

Cylindrella gheisbreghti Pfeiffer, 1856; Proc. Zool. Soc. Lond.: 380; pl. 36, fig. 1.

Eucalodium ghiesbreghti (Pfeiffer).  Crosse & Fischer, 1868; Jour. de Conchyl., 14: 88.-  Fischer & Crosse, 1872;  Miss. Sci. Mex., I: 368; pl. 14, figs. 4-4a (shell); pl. 16, figs. 14-21 (anatomy).-  Strebel, 1880; Beitrag; IV: 63; pl. 5, fig. 20 (shell); pl. 14, figs. 11-11b (shell).

Eucalodium decollatum var. gheisbreghti (Pfeiffer).  Pilsbry, 1902; Man. Conch., 15: 3-4;  pl. 1, fig. 1; pl. 4, fig. 9.—  Thompson, 1968; Bull. Fla. State Mus., 12: 125-183; fig. 1-29.

 

  TYPE LOCALITIES.-  Pupa decollata: State of Tabasco. Cylindrella gheisbreghti: Chiapas, in SE México.  The type locality for both nominate taxa is here restricted to limestone hills 5 km E of Teapa, Tabasco, México.

 

  DISTRIBUTION RECORDS.-  TABASCO: known for certain only from limestone hills immediately east of Teapa, Tabasco, México.  Records for Coban and Vera Paz, Guatemala require confirmation.

 

 

Eucalodium (Eucalodium) decollatum guatemalensis Bartsch, 1906

 

Eucalodium decollatum guatemalensis Bartsch, 1906.  Bull. U. S. Nat. Mus., 31: 110; pl. 3, fig. 9. 

 

  TYPE LOCALITY.-  Guatemala.  Holotype USNM 162307.

 

  DISTRIBUTION RECORD.-  GUATEMALA: known only from the type locality.

 

 

Eucalodium (Eucalodium) mexicanum mexicanum (Pfeiffer, 1860)

 

Cylindrella mexicanum Pfeiffer, 1860.  Proc. Zool. Soc. (London): 139.

Eucalodium mexicanum mexicanum (Pfeiffer).  Fischer & Crosse, 1873; Miss. Sci. Mex., I: 369-371.—  Strebel, 1880; Beitrag, IV: 62; pl. 6, fig. 7; pl. 11, figs. 3 (radula), 11 (jaw), 16 (anatomy).-  Pilsbry, 1902; Man. Conch., 15: 6-8; pl. 1, figs. 2-3; pl. 7, figs. 8-10.

 

  TYPE  LOCALITY.-  Republic of México.

 

  DISTRIBUTION RECORDS.-  OAXACA: Juquila, Oaxaca (Fischer & Crosse, 1873).  Earlier records list "Tabasco" and "Chiapas".

 

  TAXONOMY.-  Three subspecies are listed.  Two were orIginally described from unlocalized sources, but they were later recorded from the same locality in Oaxaca.  A third subspecies was described from Guatemala.  It is unlikely that the latter is conspecific with the form(s) from Oaxaca.

 

 

Eucalodium (Eucalodium) mexicanum major Fischer & Crosse, 1873

 

Eucalodium mexicanum major Fischer & Crosse, 1873; Miss. Sci. Mex., I:.  Miss. Sci. Mex., I: 369.-  Pilsbry, 1902; Man. Conch., 15: 8. 

 

  TYPE LOCALITY.-  Here restricted to woods between Tactíc and Tamahu, Dept. Alta Verapaz, Guatemala.

 

  DISTRIBUTION RECORDS.-  GUATEMALA, Dept. Alta Verapaz: woods between Tactíc and Tamahu.

    

 

Eucalodium (Eucalodium) mexicanum minor (Pfeiffer, 1860)

 

Cylindrella mexicanum minor Pfeiffer, 1860.  Novitat. Conch., III: 435; pl. 97, figs. 16-17.

Eucalodium mexicanum minor (Pfeiffer).  Fischer & Crosse, 1873; Miss. Sci. Mex., I: 370.-  Strebel, 1880; Beitrag, IV: 63; pl. 5, fig. 11; pl. 11, fig. 16.-   Pilsbry, 1902; Man. Conch., 15: 8; pl. 17, figs. 11-14 (shell).

 

  TYPE LOCALITY.-  Not given.

 

  DISTRIBUTION RECORDS.-  OAXACA: Juquila (Fischer & Crosse, 1873).

 

 

Eucalodium (Eucalodium) otoides Thompson, 1968

 

Eucalodium otoides Thompson, 1968; Bull. Fla. St. Mus., 12: 180-182; figs. 29a-b. 

 

  TYPE LOCALITY.-  Forested ravine 3.5 mi by road S of Rayón, Chiapas; 5500 ft. alt.  Holotype FLMNH 19050.

 

  DISTRIBUTION RECORDS.-  CHIAPAS: known only from the immediate vicinity of Rayón.

 

 

 

 

Subgenus Oligostylus Pilsbry, 1895

 

Oligostylus Pilsbry, 1895; Nautilus, 9: 51.

 

  TYPE SPECIES.-  Eucalodium blandianum Crosse & Fischer, 1868.

 

  DISTRIBUTION.-  El Salvador, Guatemala and México in the states of Chiapas, Oaxaca, Veracruz, Guerrero, Michoacán and Jalisco.

 

  TAXONOMY.-  Thirteen species are recognized.

 

 

Eucalodium (Oligostylus) australis Thompson, 1963

 

Eucalodium australis Thompson, 1963; Proc. Biol. Soc. Wash., 76: 22-27; pl. 1, figs. 5-5 (shell); text fig. 2 (anatomy).

 

  TYPE LOCALITY.-  Hacienda Monte Cristo, Metapan, Santa Ana Province, El Salvador.  Holotype SMF 161266.

 

  DISTRIBUTION RECORDS.-  EL SALVADOR: known only from the type locality.

 

 

Eucalodium (Oligostylus) blandianum Crosse & Fischer, 1868

 

Eucalodium blandianum Crosse & Fischer, 1868.  Jour. Conchyl., 16: 276.-  Fischer & Crosse, 1873; Miss. Sci. Mex., I:; Miss. Sci. Mex., 1: 374; pl. 14, figs. 5 (shell); pl. 16, figs. 11-13 (radula).-  Strebel, 1880; Beitrag, IV: 65: pl. 7, figs. 2a-k, 5 (shell); pl. 11, figs. 4, 5, 12 (shell); pl. 12, figs. 2a-o (anatomy); pl. 14, figs. 10a-b.-  Pilsbry, 1902; Man. Conch., 15: 9-10; pl. 5, figs. 23-26; pl. 8, fig. 21.

 

  TYPE LOCALITY.-  Mountains near Orizaba, Veracruz.

 

  DISTRIBUTION RECORDS.-  VERACRUZ: near Orizaba.

  

 

Eucalodium (Oligostylus) decurtatum (H. Adams, 1872)

 

Cylindrella decurtatum H. Adams, 1872; Proc. Zool. Soc. (London): 13; pl. 3, fig. 20.

Eucalodium decurtatum (H. Adams).  Fischer & Crosse, 1873; Miss. Sci. Mex., I: 385; pl. 15, figs. 5, 5a.-  Pilsbry, 1902; Man. Conch., 15: 15; pl. 1, fig. 5; pl. 6, figs. 27-29; pl. 8, figs. 15-16.

 

  TYPE LOCALITY.-  Putla, Oaxaca.

 

  DISTRIBUTION RECORDS.-  OAXACA: Putla.

 

 

Eucalodium (Oligostylus) grande (Pfeiffer, 1860)

 

Cylindrella  grande Pfeiffer, 1860; Proc. Zool. Soc. (London): 139; pl. 50, fig. 3.

Eucalodium grande (Pfeiffer). Fischer & Crosse, 1873; Miss. Sci. Mex., I: 371; pl. 15, fig. 4.-  Strebel, 1880; Beitrag, IV: 68; pl. 5, fig. 19.-  Pilsbry, 1902; Man. Conch., 15: 13-14; pl. 6, figs. 30-32, 36-37; pl. 8, fig. 20. 

 

  TYPE LOCALITY.-  Juquila, Oaxaca.

 

  DISTRIBUTION RECORDS.-  OAXACA: known only from the type locality.

 

 

Eucalodium (Oligostylus) hegewischi (Bartsch, 1947)

 

Bulinus truncatus Pfeiffer, 1841; Symb. Hist. Helc., I: 43.  (Not Bulimus truncatus Brugnoière, 1792).

Cylindrella truncata Von Martens, 1865; Malak. Blätt., 12: 13.

Eucalodium truncatum (Pfeiffer).  Fischer & Crosse, 1873; Miss. Sci. Mex., I: 392.-  Strebel, 1880; Beitrag, IV: 73; pl. 11, fig. 8 (radula), 14 (jaw); pl. 12, fig. 3 (genitalia); pl. 13, fig. 13 (shell).-  Von Martens, 1877; Biol. Cent. Amer.: 264; pl. 16, figs. 309.

Oligostylus hegewischi Bartsch, 1947; Jour. Wash. Acad. Sci., 37: 286.-  Bartsch, 1948; Jour. Wash. Acad. Sci., 38: 350-351.

 

  TYPE LOCALITY.-  Angangueo, Michoacán; type lost.

 

  DISTRIBUTION RECORDS.-  MICHOACÁN: known from the area near Angangueo, 9000 ft. alt. (Thompson, 1968).  Records of this species from Omilteme, Guerrero (Von Martens, 1897: 264) almost certainly pertain to Eucalodium mariae (Bartsch, 1947).

 

 

Eucalodium (Oligostylus) hippocastaneum Dall, 1897

 

Eucalodium hippocastaneum Dall, 1897; Nautilus, 11: 61.-   Pilsbry, 1902; Man. Conch., 15: 15-16.

 

  TYPE LOCALITY.-  San Sabastian, Jalisco.  Holotype USNM 186137.

 

  DISTRIBUTION RECORDS.-  MICHOACÁN: known only from the type locality.

 

 

Eucalodium (Oligostylus) insigne Crosse & Fischer, 1872

 

Eucalodium insigne Crosse & Fischer, 1872; Jour. Conchyl., 22: 301.-  Fischer & Crosse, 1873; Miss. Sci. Mex., I: pl. 14, figs. 7, 7a.-  Pilsbry, 1902; Man. Conch., 15: 12-13; pl. 4, figs. 6-7.

 

  TYPE LOCALITY.-  “Southern Mexico”.

 

  DISTRIBUTION RECORDS.-  Unknown.

 

 

Eucalodium (Oligostylus) mariae (Bartsch, 1947)

 

Oligostylus mariae Bartsch, 1947; Jour. Wash. Acad. Sci., 37: 285-286; fig. 1.

 

  TYPE LOCALITY.-  Omiltemi [Omilteme], Guerrero.  Holotype USNM 543582.

 

  DISTRIBUTION RECORDS.-  GUERRERO: Omilteme.

 

 

Eucalodium (Oligostylus) moussonianum Crosse & Fischer, 1872

 

Eucalodium moussonianum Crosse & Fischer, 1872;  Jour. Conchyl., 20: 225.-  Fischer & Crosse, 1873; Miss. Sci. Mex., I: 375; pl. 14, figs. 11, 11a.-  Strebel, 1880; Beitrag, IV: 67; pl. 5, fig. 16; pl. 13; fig. 14.—  Pilsbry, 1902; Man. Conch., 15: 16-17; pl. 3, figs. 15-18. 

 

  TYPE LOCALITY.-  Not stated.

 

  DISTRIBUTION RECORDS.-  Unknown.

 

 

Eucalodium (Oligostylus) neglectum Crosse & Fischer, 1872

 

Eucalodium neglectum Crosse & Fischer, 1872; Jour. Conchyl., 20: 302.-  Fischer & Crosse, 1873; Miss. Sci. Mex., I: 373; pl. 14, figs. 8, 8a.-  Strebel, 1880; Beitrag, IV: 67; pl. 13, fig. 15.-  Pilsbry, 1902; Man. Conch., 15: 17-18; pl. 3, figs. 19-20. 

 

  TYPE LOCALITY.-  Oaxaca.

 

  DISTRIBUTION RECORDS.-  OAXACA: known only from the type locality.

 

 

Eucalodium (Oligostylus) splendidum (Pfeiffer, 1860)

 

Cylindrella splendidum Pfeiffer, 1860; Proc. Zool. Soc. London: 139; pl. 50, fig. 1.

Eucalodium splendidum (Pfeiffer).  Fischer & Crosse, 1873; Miss. Sci. Mex., I: 372; pl. 15, figs. 3, 3a.-  Strebel, 1880; Beitrag, IV: 69; pl. 5, fig. 17.-  Pilsbry, 1902; Man. Conch., 15: 14; pl. 6, figs. 33-39; pl. 8, figs. 19, 23.

 

  TYPE LOCALITY.-  Zacatepec, Oaxaca.

 

  DISTRIBUTION RECORDS.-  OAXACA: known only from the type locality.

 

 

Eucalodium (Oligostylus) sumichrasti Crosse & Fischer, 1878

 

Eucalodium sumichrasti Crosse & Fischer, 1878; Jour. Conchyl., 26: 250.-  Crosse & Fischer, 1879, 27: 46; pl. 2, fig. 2.-  Fischer & Crosse, 1890; Miss. Sci. Mex., II: 665; pl. 72, figs. 1-1a.-  Pilsbry, 1902; Man. Conch., 15: 10-11; pl. 1, figs. 10-11.

 

  TYPE LOCALITY.-  Chiapas.

 

  DISTRIBUTION RECORDS.-  CHIAPAS: known only from the type locality.

 

 

Eucalodium (Oligostylus) walpoleanum Crosse & Fischer, 1872

 

Eucalodium walpoleanum Crosse & Fischer, 1872; Jour. Conchyl., 20: 75.-  Fischer & Crosse, 1873; Miss. Sci. Mex., I: 377; pl. 14, figs. 6-6a.-  Strebel, 1880; Beitrag, IV: 67; pl. 7, figs. 1a-1b.-  Pilsbry, 1902; Man. Conch., 15:11-12; pl. 4, figs. 1-4; pl. 8, fig. 22.-  Bartsch, 1906; Proc. U. S. Nat. Mus., 31: 111. 

 

  TYPE LOCALITY.-  Woods of Palenque, Chiapas.

 

  DISTRIBUTION RECORDS.-  CHIAPAS: Palenque.  GUATEMALA: Dept. Alta Verapaz.

 

 

Subgenus Ptychocentrum Bartsch, 1943

 

Coelocentrum (Ptychocentrum) Bartsch, 1943; Nautilus, 56: 91-92; pl. 7, figs. 8-9.

 

  TYPE SPECIES.-  Coelocentrum bourgeoisae Bartch, 1943 [=Coelocentrum (Ptychocentrum) marianum Bartsch, 1943].

 

  DISTRIBUTION.-  Central Chiapas, México.

 

  TAXONOMY.-  One species.

 

 

Eucalodium (Ptychocentrum) marianum (Bartsch, 1943)

 

Coelocentrum (Ptychocentrum) bourgeoisae Bartsch, 1943; Nautilus, 56: 91-92; pl. 7, figs. 8-9. (Not Coelocentrum bourgeoisae Pilsbry, 1939).

Coelocentrum (Ptychocentrum) marianum Bartsch, 1943; Nautilus, 56: 144.-  Solem, 1957; Notulae Naturae, (298): 8.

 

  TYPE LOCALITY.-  Tecpatán, Chiapas.  Holotype USNM 536900.

 

  DISTRIBUTION RECORDS.—  CHIAPAS: known only from the type locality.

 

 

 

Subgenus Resupinata Von Martens, 1897

 

Resupinata Von Martens, 1897: 255.

 

  TYPE SPECIES.-  Cylindrella speciosa Dunker, 1844 (Pilsbry, 1953: 134).

 

  DISTRIBUTION.-  Central Veracruz, adjacent northern Oaxaca and San Luís Potosí, México.

 

  TAXONOMY.-  Six species and five subspecies.

 

 

Eucalodium (Resupinata) cereum Strebel, 1880

 

Eucalodium cereum Strebel, 1880; Beitrag, IV: 72; pl. 6, fig. 9.-  Pilsbry, 1902; Man. Conch., 15: 22-23; pl. 7, figs. 4-7.

 

  TYPE LOCALITY.-  San Antonio del Norte, 3 leagues east from Neolingo, on road to Misantla, Veracruz.

 

  DISTRIBUTION RECORDS.-  VERACRUZ: known only from the type locality.

 

 

Eucalodium (Resupinata) densecostatum Strebel, 1880

 

Eucalodium densecostatum Strebel, 1880; Beitrag, IV: 71; pl. 6, fig. 10.-  Pilsbry, 1902; Man. Conch., 15: 20; pl. 1, fig. 4. 

 

  TYPE LOCALITY.-  Orizaba, Veracruz.

 

  DISTRIBUTION RECORDS.-  VERACRUZ: Orizaba; Misantla (Bartsch, 1906: 111).

 

 

 

Eucalodium (Resupinata) deshayesianum Crosse & Fischer, 1873

 

Eucalodium deshayesianum Crosse & Fischer, 1872; Jour. de Conchyl., 20: 223.-  Fischer & Crosse, 1873; Miss. Sci. Mex., I: 384; pl. 14, figs. 9-9a.-  Pilsbry, 1902; Man. Conch., 15: 24; pl. 2, figs. 13-14. 

 

  TYPE LOCALITY.-  “Southern Mexico”.

 

  DISTRIBUTION RECORDS.-  Unknown.

 

 

Eucalodium (Resupinata) edwardsianum Crosse & Fischer, 1872

 

Eucalodium edwardsianum Crosse & Fischer, 1872.  Jour. de Conchyl., 20: 224.-  Fischer & Crosse, 1873;  Miss. Sci. Mex., I: 383; pl. 14, figs. 10-10a.-  Strebel, 1880; Beitrag, IV: 69; pl. 6, fig. 11; pl. 11, figs. 6, 7, 13; pl. 12, fig. 1 (anatomy).-  Pilsbry, 1902; Man. Conch.; 15: 23-24; pl. 2, figs. 8-9, 12. 

 

  TYPE LOCALITY.-  Cordova, Veracruz.

 

  DISTRIBUTION RECORDS.-  VERACRUZ: Córdoba.

 

 

Eucalodium (Resupinata) ischnostele (Pilsbry, 1909)

 

Coelocentrum ischnostele Pilsbry, 1909; Nautilus, 22: 139.

Eucalodium (Resupinata) ischnostele (Pilsbry).  Pilsbry, 1953; Proc. Acad. Nat. Sci. Phila., 105: 133-134; pl. 3, figs. 1, 1a.

 

  TYPE LOCALITY.-  Mecos Falls, San Luís Potosí, México. Lectotype ANSP 99487A (H. B. Baker, 1963: 223).

 

  DISTRIBUTION RECORDS.-  SAN LUÍS POTOSÍ: Mecos Falls; near Tomasopo.

 

 

Eucalodium (Resupinata) speciosum speciosum (Dunker, 1844)

 

Cylindrella speciosum Dunker in Phillipi, 1844; Abbild. Beschreib., I: 86.

Eucalodium speciosum speciosum (Dunker).  Fischer & Crosse. 1873;  Miss. Sci. Mex., I: 381; pl. 15, figs. 6, 6a.-  Strebel, 1880; Beitrag, IV: 71; pl. 6, fig. 6.-  Pilsbry, 1902; Man. Conch.; 15: 20-21; pl. 2, figs. 1-3.-  Bartsch, 1906; Proc. U. S. Nat. Mus., 31: 110.

Eucalodium boucardi var. minor Fischer & Crosse, 1873; Miss. Sci. Mex., I: 381; pl. 15, figs. 6-6a (shell). 

 

   TYPE LOCALITY.-  Unknown.

 

   DISTRIBUTION RECORDS.-  VERACRUZ: Córdoba; Coatpec; Cuesta de Misantla, Jalapa.

 

 

Eucalodium (Resupinata) speciosum boucardi (Pfeiffer, 1856)

 

Cylindrella boucardi Pfeiffer, 1856; Proc. Zool. Soc. Lond.: 321; pl. 35, fig. 1.

Eucalodium boucardi (Peiffer). Fischer & Crosse, 1873; Miss. Sci. Mex., I: 381.-  Strebel, 1880; Beitrag, IV: 70; pl. 5, fig. 15.

Eucalodium speciosum boucardi (Pfeiffer).  Von Martens, 1897; Biol. Cent. Amer.: 262.-  Pilsbry, 1902; Man. Conch., 15: 21-22; pl. 2, figs. 6-7; pl. 8, fig. 18. 

 

  TYPE LOCALITY.-  Cordova, Veracruz, México.

 

  DISTRIBUTION RECORDS.-  VERACRUZ: Córdoba; Orizaba.

 

 

Eucalodium (Resupinata) speciosum fischeri Von Martens, 1897

 

Eucalodium speciosum (Dunker). Fischer & Crosse, 1873; Miss. Sci. Mex., I:: 379; pl. 15, figs. 7-7a (shell).

Eucalodium speciosum fischeri Von Martens, 1897.  Biol. Cent. Amer.: 262.-  Pilsbry, 1902; Man. Conch.; 15: 22; pl. 2, figs. 10-11. 

 

  TYPE LOCALITY.-  Chiquihuitl, Veracruz.

 

  DISTRIBUTION RECORDS.-  VERACRUZ: Chiquihuitl.

 

 

Eucalodium (Resupinata) speciosum minimum Von Martens, 1897

 

Eucalodium speciosum g Fischer & Crosse, 1873; Miss. Sci. Mex., I: 381.

Eucalodium speciosum minimum Von Martens, 1897; Biol. Cent. Amer.: 262.-  Pilsbry, 1902; Man. Conch. 15: 22. 

 

  TYPE LOCALITY.-  Cordova, Veracruz, México.

 

  DISTRIBUTION RECORDS.-  VERACRUZ: Córdoba; Atoyac.

 

 

Eucalodium (Resupinata) speciosum strebeli Von Martens, 1897

 

Eucalodium boucardi form B, Strebel, 1860; Beitrag, IV: 71; pl. 5, figs. 8-9.

Eucalodium speciosum strebeli Von Martens, 1897.  Biol. Cent. Amer., 262.-  Pilsbry, 1902; Man. Conch., 15: 22; pl. 2, figs. 4-5. 

 

  TYPE LOCALITY.-  Environs of Jalapa, especially about the brooks on the road to Neolingo, and Chirimoya, Veracruz.

 

  DISTRIBUTION RECORDS.-  VERACRUZ: environs of Jalapa.

 

 

 

Genus Anisospira Strebel, 1880

 

Anisospira Strebel, 1880: 77.-  Pilsbry, 1902; Man. Conch., 15: 24-25.-  Thompson, 1968; Bull. Fla. St. Mus., 12: 142.

 

  TYPE SPECIES.-  Cylindrella liebmanni Pfeiffer, 1846.

 

  DISTRIBUTION.-  Known from lowland regions of Colima, Michoacán and Oaxaca, México.

 

  TAXONOMY.-  Anisospira includes two subgenera, seven species and two subspecies.

 

 

Subgenus Anisospira Strebel, 1880

 

  DISTRIBUTION.-  Southern Oaxaca, México.

 

  TAXONOMY.-  Three species, one of which has two subspecies

 

 

Anisospira (Anisospira) dalli dalli (Von Martens, 1901)

 

Anisospira strebeli Dall, 1897; Proc. U. S. Nat. Mus., 19: 353; pl. 33, figs. 7-8 (not Anisospira strebeli Pfeffer, 1887).

Eucalodium dalli Von Martens, 1901; Biologia Centrali-Americana: 633 (new name for Anisospira strebeli Dall, 1897).

Anisospira dalli (Von Martens). Pilsbry, 1902; Man. Conch., 15: 26-27; pl. 1, fig. 12; pl. 10, figs. 28-29.—  Solem, 1957; Notulae naturae, (298): 5; pl. 1, fig. 6.

Anisospira dalli dalli (Von Martens).  Thompson, 1968; Bull. Fla. St. Mus., 12: 142-147; figs.10a-f, 11-12.

 

  TYPE LOCALITY.-  Huilotepéc, Oaxaca, México.  Holotype USNM 107366.

 

  DISTRIBUTION RECORDS.—  OAXACA: limestone hills west and north of Tehuantepec (Thompson, 1968).

 

 

Anisospira (Anisospira) dalli stringens Thompson, 1968

 

Anisospira dalli stringens Thompson, 1968; Bull. Fla. St. Mus., 12: 147-148; figs. 10g-l.  

 

  TYPE LOCALITY.-  Hill about 0.4 mi. S of the Presa Benito Juarez on the Rio Tehuantepec, about 18 mi. NW of Tehuantepec, Oaxaca.  Holotype FLMNH 19079.

 

  DISTRIBUTION RECORDS.-  OAXACA: known only from the type locality.

 

 

Anisospira (Anisospira) liebmanni (Pfeiffer, 1846)

 

Cylindrella liebmanni Pfeiffer, 1846; Zeitschrift fur Malak.:1 59.

Anisospira liebmani (Pfeiffer).  Strebel, 1880; Beitrag, IV:  79; pl. 5, figs. 12-13; pl. 14, fig. 2.-  Pilsbry, 1902; Man. Conch., 15: 28-29; pl. 10, figs. 22-27; pl. 11, figs. 1-3.-  Solem, 1957; Notulae Naturae, (298): 5-6; pl. 1, fig. 5.-  Thompson, 1968; Bull. Fla. St. Mus., 12: 133-142; figs. 1-9 (shells, anatomy, distribution).

Cylindrella trochaeformis Sowerby, in Reeve, 1843; Conch. Icon., XX, pl. 9, fig. 80.

Cylindrella hyalina Pfeiffer in Philippi, 1847: Abild., II; pl. 2, fig. 2.

Eucalodium (Anisospira) orcutti Dall, 1910; Nautilus, 24: 34-36. 

Anisospira orcutti (Dall).  Solem, 1957; Notulae naturae, (298): 6, pl. 1.

Liocentrum wilmoti Bartsch, 1948; Proc. Biol. Soc. Wash., 38: 52; fig. 2. 

 

  TYPE LOCALITIES.-  Cylindrella liebmanni: unknown.  Cylindrella trochaeformis: unknown.  Cylindrella hyaline: unknown.  Eucalodium (Anisospira) orcutti: Rio Verde, Oaxaca; holotype USNM 212319.  Liocentrum wilmoti: along the Pan American Highway between Oaxaca and Tehuantepec, Oaxaca; holotype USNM 589051.

 

  DISTRIBUTION RECORDS.-  Southeastern OAXACA.

 

 

Anisospira (Anisospira) vonmartensi (Strebel, 1880)

 

Eucalodium vonmartensi Strebel, 1880; Beitrag, IV: 73-74; pl. 11, figs. 8, 14; pl. 12, fig. 3; pl. 13, fig. 13.

Anisospira vonmartensi (Strebel).  Bartsch, 1947; Jour. Wash. Acad. Sci., 37: 284-285.

 

  TYPE LOCALITY.-  "México".

 

  DISTRIBUTION RECORDS.-  Unknown.

 

 

 

Subgenus Trachycion Thompson, 1968

 

Trachycion Thompson, 1968; Bull. Fla. St. Mus., 12: 148.

 

  TYPE SPECIES.-  Anisospira recticosta townsendi Pilsbry & Cocherell, 1903.

 

  DISTRIBUTION.—  Coastal areas of Colima, Michoacán and Oaxaca, México.

 

  TAXONOMY.-  Four species, one of which has two subspecies.

 

 

Anisospira (Trachycion) hadromylla Thompson, 1968

 

Anisospira hadromylla Thompson, 1968; Bull. Fla. St. Mus., 12: 150-152, figs. 13a-d. 

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