2010-2011 Staff Publications

Briscoe, A. D., S. M. Bybee, G. D. Bernard, F. Yuan, M. P. Sison-Mangus, R. D. Reed, A. D. Warren, J. Llorente-Bousquets and C. C. Chiao. 2010. Reply to Nozawa et al.: Complementary statistical methods support positive selection of a duplicated UV opsin gene in Heliconius. Proceedings of the National Academy of Sciences, 107: E97.

Brown, J. W., T. A. Heard, R. Segura, and J. Rota. 2011. Leaf-roller moths (Tortricidae) reared from the invasive weed Mexican palo verde (Parkinsonia aculeata), with comments on their host specificity, biology, and geographic distribution. Journal of Insect Science, 11(7): 1-17.

Cho, S., Zwick, A., Regier, J.C, Mitter, C., Cummings, M.P., Yao, J., Du, Z., Zhao, H., Kawahara, A.Y., Weller, S., Davis, D.R., Baixeras, J., Brown, J.W., Parr, S. Can deliberately incomplete gene sample augmentation improve a phylogeny estimate for the advanced moths and butterflies (Hexapoda: Lepidoptera)? Systematic Biology 60(6): 782–796. Download PDF

Cox J. H., and T. C. Emmel. 2010. Ecological surveys of the Lepidoptera fauna of the Hunstein Range, East Sepik Province, Papua New Guinea, emphasizing butterfly populations and habitat in the Mt. Samsai area. Tropical Lepidoptera Research, 20(2): 88-99.

de-Silva, D. L., J. J. Day, M. Elias, K. R. Willmott, A. Whinnett, and J. Mallet. 2010. Molecular phylogenetics of the neotropical butterfly subtribe Oleriina (Nymphalidae: Danainae: Ithomiini). Molecular Phylogenetics and Evolution, 55: 1032-1041 (May).

Fontes, E., H. Frank, J. Gillmore, and D. Matthews. 2010. Dale H. Habeck, 1931-2010. Florida Entomologist, 93(3): 478-479.

Hall, J. P. W., and K. R. Willmott. 2010. Discovery of a new Lucillella species (Riodinidae: Symmachiini) in the eastern Andes of Ecuador using the single rope canopy access technique. Journal of the Lepidopterists’ Society, 64(3): 139-146 (November).

Hardy, D., M. A. Rickard, A. D. Warren and N. V. Grishin. 2011. Achalarus tehuacana (Hesperiidae: Eudaminae): a new United States record from southern Texas. News of the Lepidopterists’ Society, 52(4): 107-111, 127.

Hayden, J. E. 2010. Phylogeny, distribution, and description of a Caribbean species of Dicepolia (Lepidoptera: Crambidae). Tropical Lepidoptera Research, 20(2): 77–84.

Heppner, J. B. 2010. Zeller and his “Exotische Microlepidoptera.” Lepidoptera Novae (Gainesville), 3(1):1-28.

Heppner, J. B. 2010. Review of the genus Alinguata from Venezuela (Lepidoptera: Alucitidae). Lepidoptera Novae (Gainesville), 3(1): 57-59.

Heppner, J. B.. 2010. A new species of Paranthozela from Peru (Lepidoptera: Tortricidae: Olethreutinae). Lepidoptera Novae (Gainesville), 3(1): 61-63.

Heppner, J. B. 2010. Loxotoma elegans in Peru (Lepidoptera: Oecophoridae: Stenomatinae). Lepidoptera Novae (Gainesville), 3(1): 64.

Heppner, J. B. 2010. A new species of Scythris from western Peru (Lepidoptera: Scythrididae). Lepidoptera Novae (Gainesville), 3(1): 65-68.

Heppner, J. B. 2010. Riley’s metalmark moth, Hemerophila diva: a beauty in Florida (Lepidoptera: Choreutidae). Lepidoptera Novae (Gainesville), 3(2): 111-112.

Heppner, J. B. 2010. Syngamia florella and its variations in Florida and the Neotropics (Lepidoptera: Pyralidae: Pyraustinae). Lepidoptera Novae (Gainesville), 3(2): 113-118.

Heppner, J. B. 2010. Nearctic metalmark moths, 1. Genus Phormoestes (Lepidoptera: Choreutidae: Millieriinae). Lepidoptera Novae (Gainesville), 3(2): 119-124.

Heppner, J. B. 2010. Scythris medullata from western Peru, with notes on the family in the Neotropics (Lepidoptera: Scythrididae). Lepidoptera Novae (Gainesville), 3(2): 125-128.

Heppner, J. B. 2010. Profilinota phillita in Peru (Lepidoptera: Oecophoridae: Depressariinae). Lepidoptera Novae (Gainesville), 3(2): 129-132.

Heppner, J. B. 2010. Notes on the brachypterous moth, Pringleophaga kerguelensis, of the Subantarctic Kerguelen Islands (Lepidoptera: Tineidae). Lepidoptera Novae (Gainesville), 3(3): 133-143.

Heppner, J. B. 2010. The new genus Rindgeria and its species in North America and Central America (Lepidoptera: Geometridae: Ennominae). Lepidoptera Novae (Gainesville), 3(3): 149-153.

Heppner, J. B. 2010. Incawockia, a new genus and species from Peru (Lepidoptera: Urodidae: Galacticinae). Lepidoptera Novae (Gainesville), 3(3): 154-158.

Heppner, J. B. 2010. Nearctic metalmark moths, 2. Genus Brenthia (Lepidoptera: Choreutidae: Brenthiinae). Lepidoptera Novae (Gainesville), 3(3): 159-163.

Heppner, J. B. 2010. Notes on Vietnam moths, 9. Massing of Vitessa moths (Lepidoptera: Pyralidae: Pyralinae). Lepidoptera Novae (Gainesville), 3(3): 164.

Heppner, J. B. 2010. Notes on Vietnam moths, 10. Notes on Wockia in Vietnam (Lepidoptera: Urodidae: Galacticinae). Lepidoptera Novae (Gainesville), 3(3): 165-168.

Heppner, J. B. 2010. Notes on Vietnam moths, 11. Propachys nigrivena (Lepidoptera: Pyralidae: Pyralinae). Lepidoptera Novae (Gainesville), 3(3): 169-171.

Heppner, J. B. 2010. Notes on Vietnam moths, 12. Ashinaga longimana in Vietnam (Lepidoptera: Oecophoridae: Oecophorinae). Lepidoptera Novae (Gainesville), 3(3): 172-174.

Heppner, J. B. 2010. Notes on Vietnam moths, 13. Phycodes taonopa in Vietnam (Lepidoptera: Brachodidae: Phycodinae). Lepidoptera Novae (Gainesville), 3(3): 175-178.

Heppner, J. B. 2010. Notes on Vietnam moths, 14. Vespisesia, a new clearwing genus and species from Vietnam (Lepidoptera: Sesiidae: Sesiinae). Lepidoptera Novae (Gainesville), 3(3): 179-181.

Heppner, J. B. 2010. Notes on Vietnam moths, 15. A new Cerace species for Vietnam (Lepidoptera: Tortricidae: Tortricinae). Lepidoptera Novae (Gainesville), 3(3): 182-186.

Heppner, J. B. 2010. Notes on Vietnam moths, 16. A new Darantasia from Vietnam (Lepidoptera: Arctiidae: Lithosiinae). Lepidoptera Novae (Gainesville), 3(4): 255-258.

Heppner, J. B. 2010. Notes on Vietnam moths, 17. Trebania muricolor in Vietnam (Lepidoptera: Pyralidae: Pyralinae). Lepidoptera Novae (Gainesville), 3(4): 259-261.

Heppner, J. B. 2010. Panama moth notes, 3. Cotaena plenella (Lepidoptera: Glyphipterigidae). Lepidoptera Novae (Gainesville), 3(3): 187-190.

Heppner, J. B. 2010. Panama moth notes, 4. A new species of Taeniostolella (Lepidoptera: Glyphipterigidae). Lepidoptera Novae (Gainesville), 3(3): 191-193.

Heppner, J. B. 2010. Panama moth notes, 5. Notes on Darceta falcata (Lepidoptera: Noctuidae: Agaristinae). Lepidoptera Novae (Gainesville), 3(4): 225-227.

Heppner, J. B. 2010. Fauna of British India, Sphingidae larvae, by Bell and Scott, 1937. Lepidoptera Novae (Gainesville), 3(4): 199-216.

Heppner, J. B. 2010. Notes on the Moluccan and Papuan genus Cocytia (Lepidoptera: Noctuidae: Cocytiinae). Lepidoptera Novae (Gainesville), 3(4): 217-221.

Heppner, J. B. 2010. Notes on the African genus Ommatothelxis (Lepidoptera: Oecophoridae: Xyloryctinae). Lepidoptera Novae (Gainesville), 3(4): 222-224.

Heppner, J. B. 2010. The genus Eumarozia and a new species in Peru (Lepidoptera: Tortricidae: Olethreutinae). Lepidoptera Novae (Gainesville), 3(4): 228-231.

Heppner, J. B. 2010. Guatemala moth notes, 3. Eumarozia beckeri in Guatemala (Lepidoptera: Tortricidae: Olethreutinae). Lepidoptera Novae (Gainesville), 3(4): 232-234.

Heppner, J. B. 2010. Guatemala moth notes, 4. Caripeta hyperythrata in Guatemala (Lepidoptera: Geoemtridae: Ennominae). Lepidoptera Novae (Gainesville), 3(4): 235-236.

Heppner, J. B., and J. Brambila. 2010. Florida Lepidoptera notes, 9. Duponchelia fovealis in Florida. (Lepidoptera: Pyralidae: Pyraustinae). Lepidoptera Novae (Gainesville), 3(4): 237-241.

Heppner, J. B. 2010. Florida Lepidoptera notes, 10. A new Tischeria from Florida (Lepidoptera: Tischeriidae). Lepidoptera Novae (Gainesville), 3(4): 242-244.

Heppner, J. B. 2010. Florida Lepidoptera notes, 11. Digitivalva clarkei in Florida (Lepidoptera: Acrolepiidae). Lepidoptera Novae (Gainesville), 3(4): 245-248.

Heppner, J. B. 2010. Florida Lepidoptera notes, 12. Bucculatrix needhami (Lepidoptera: Bucculatricidae). Lepidoptera Novae (Gainesville), 3(4): 249-250

Heppner, J. B. 2010. Florida Lepidoptera notes, 13. Perimede erransella (Lepidoptera: Cosmopterigidae). Lepidoptera Novae (Gainesville), 3(4): 251-254.

Heppner, J. B. 2010. Review of the Neotropical genus Hexeretmis (Lepidoptera: Alucitidae). Lepidoptera Novae (Gainesville), 3(4): 263-268.

Heppner, J. B., and Y. S. Bae. 2010. Notes on Taiwan Moths, 6. Cerace stipatana (Lepidoptera: Tortricidae: Tortrici­nae). Lepidoptera Novae (Gainesville), 3(3): 195-198.

Heppner, J. B., and T. S. Dickel. 2010. The European yellow underwing moth, Noctua pronuba (Lepidoptera: Noctuidae), in Florida. Southern Lepidopterists’ News (Gainesville), 32: 58-59.

Heppner, J. B., T. S. Dickel and V. A. Brou, Jr. 2010. New North American records of the Asian species, Simplicia cornicalis, in Florida and Louisiana (Lepidoptera: Noctuidae: Herminiinae). Lepidoptera Novae (Gainesville), 3(1): 53-56.

Judd, R. E. and D. L. Matthews. 2010. Report on the capture of a gravid Samia cynthia female in Alachua County, Florida. Southern Lepidopterists’ News, 32(4): 182-184.

Kawahara, A.Y.,J. C. Sohn, J. De Prins, and S. Cho. 2010. Taxonomic report of five leaf-mining moths new to Korea (Lepidoptera: Gracillariidae). Entomological Research, 40: 131-135.

Kawahara, A.Y., Ohshima, I., Kawakita, A., Mitter, C., Regier, J.C., Davis, D., Wagner, D.L., De Prins, J., Lopez-Vaamonde, C. Increased gene sampling strengthens support for higher-level groups within leaf mining moths and relatives (Lepidoptera: Gracillariidae). BMC Evolutionary Biology 11:182, DOI:10.1186/1471-2148-11-182.

Kawahara, A.Y., Tangalin, N., Rubinoff, D. Life-history notes on the fern-mining endemic Hyposmocoma (Euperissus) trivitella Swezey 1913 (Lepidoptera: Cosmopterigidae) from Kauai and a report of associated parasitoids (Hymenoptera: Bethylidae). Proceedings of the Hawaiian Entomological Society 43: 9-12.

Kawahara, A.Y., Nishida, K., Rubinoff, D. The behavior of the Hawaiian dancing moth, Dryadaula terpischorella (Lepidoptera: Tineidae). Journal of the Lepidopterists’ Society 65(2): 133-135

Kawahara, A. Y. Review of Martin, A., Soares, A. Bizarro, J. A Guide to the hawkmoths of the Serra dos Órgãos, South-eastern Brazil. Tropical Lepidoptera Research 21(2): 105.

Kawahara, A.Y. 2011. Review of Goldsmith, M.R. and Marec F. (eds) “Genetics and Molecular Biology of Lepidoptera.” Florida Entomologist, 94(1): 119-120.

Kim S. R., B. K. Byun, K. T. Park, and S. H. Lee. 2010. A taxonomic review of the genus Nippoptilia (Lepidoptera: Pterophoridae) from Korea, with description of a new species. Journal of Natural History, 44: 601-613.

Kim S. R., B. K. Byun, K. T. Park, and S. H. Lee. 2010. Taxonomic study of the tribe Oidaematophorini (Lepidoptera: Pterophoridae) from Korea, with descriptions of two new species. Journal of Natural History, 44: 1377-1399.

Kim S. R., B. K. Byun, K. T. Park, and S. H. Lee. 2010. Genus Promalactis Meyrick (Lepidoptera, Oecophyoridae) in the northern Vietnam, Part 1: Descriptions of six new species. Florida Entomologist, 93(4): 546-557.

Kwon, Y. D., E. M. Ji, and K. T. Park. 2010. A new record of Disparia diluta from Korea, with note on Disparia nihonica (Lepidoptera: Notodontidae). Korean Journal of Systematic Zoology, 26(1): 35-37.

Lukhtanov, V. A. 2010. Dobzhansky’s rule and reinforcement of pre-zygotic reproductive isolation in zones of secondary contact. Zhurnal Obshchei Biologii, 71(5): 372–385.

Lukhtanov, V. A. 2010. From Haeckel’s phylogenetics and Hennig’s cladistics to the method of maximum likelihood: advantages and limitations of modern and traditional approaches to phylogeny reconstruction. Entomological Review, 90(3): 299–310.

Lukhtanov, V. A. 2010. Role of natural selection in speciation: reinforcement of pre-zygotic reproductive isolation in Agrodiaetus blue butterflies. In Charles Darwin and Current Biology. Ed. E.I. Kolchinsky. St. Petersburg. Pp. 268-276.

Lukhtanov, V. A., and V. G. Kuznetsova. 2010. What genes and chromosomes say about the origin and evolution of insects and other arthropods. Russian Journal of Genetics, 46(9): 1115–1121.

Matthews, D. L. 2010. A new species of Hellinsia from the Southeastern United States (Lepidoptera: Pterophoridae). Bulletin of the Allyn Museum, 161: 1-13.

Matthews, D. L. 2010. Mississippi plume moths from the Bryant Mather collection (Lepidoptera: Pterophoridae). Southern Lepidopterists’ News, 32(2): 50-55.

Matthews, D. L. and G. Gielis. 2011. Adaina ipomoeae Bigot and Etienne, 2009, new records for Florida and the West Indies (Lepidoptera: Pterophoridae). Insecta Mundi, 0156: 1-3.

Matthews, D. L., and J. Y. Miller. 2010. Notes on the Cacao Plume Moth in Honduras and description of the larvae and pupae (Lepidoptera: Pterophoridae). Tropical Lepidoptera Research, 20(1): 28-34.

Matthews, D. L., and J. Y. Miller. 2010. Remembering Dale H. Habeck, October 21, 1931 – May 17, 2010. Southern Lepidopterists’ News, 32(3): 141-142.

Miller, J. Y., D. L. Matthews, and J. Brambila. 2011. Southern Lepidopterists at Butterflyfest 2010. Southern Lepidopterists’ News, 33(1): 28-30.

Mitter, K. T., T. B. Larsen, S. Collins, G. Vande Weghe, J. De Prins, W. De Prins, S. Sfian, E. Zakharov, D. J. Hawthorne, A. Y. Kawahara, and J. C. Regier. 2011. Genetic evidence for multiple cryptic species of Pseudopontia (Pieridae: Pseudopontiinae). Systematic Entomology, 36(1): 139-163.

Mullen, S. P., W. Savage, and K. R. Willmott. 2010. Rapid diversification and not clade age explains high diversity in neotropical Adelpha butterflies. Proceedings of the Royal Society of London B doi:10.1098/rspb.2010.2140 (November).

Nieukerken, E.J.., Kaila, L., Kitching, I.J., Kristensen, N.P., Lees, D.C., Minet, J., Mitter, C., Mutanen, M., Regier, J.C., Simonsen, T.J., Wahlberg, N., Yen, S.-H., Zahiri, R., Adamski, D., Baixeras, J., Bartsch, D., Bengtsson, B.Å., Brown, J.W., Bucheli, S.R., Davis, D.R., De Prins, J., De Prins, W., Epstein, M.E., Gentili-Poole, P., Gielis, C., Hättenschwiler, P., Hausmann, A., Holloway, J.D., Kallies, A., Karsholt, O., Kawahara, A.Y., Koster, S.J.C., Kozlov, M., Lafontaine, J.D., Lamas, G., Landry, J.-F., Lee, S., Nuss, M., Park, K.T., Penz, C., Rota, J., Schmidt, B.C., Schintlmeister, A., Sohn, J.C., Solis, M.A., Tarmann, G.M., Warren, A.D., Weller, S., Yakovlev, R.V., Zolotuhin, V.V., Zwick, A. (2011) Order Lepidoptera Linnaeus, 1758. In: Zhang, Z.-Q. (Ed.), Animal biodiversity: An outline of higher-level classification and survey of taxonomic richness. Zootaxa 3148: 212–221.

Padrón P. S. 2010. Systematics and Biogeography of High Altitude Tropical Andean Satyrines (Lepidoptera, Nymphalidae: Satyrinae). M.Sc. Thesis, University of Florida, Gainesville, Florida, USA. 98 pp.

Park, K. T. 2011. A new species of Halolaguna Gozmány from Thailand (Lepidoptera, Lecithoceridae). Journal of Asia-Pacific Entomology, 14: (doi:10.1016/j.aspen.2010.12.006).

Park, K. T. 2011. Lecithoceridae (Lepidoptera, Gelechioidea) of New Guinea, Part II: Hamatina Park with description of two new species. Journal of Asia-Pacific Entomology, 14: (doi: 10.1016/j.aspen.2010.12.005).

Park, K. T. 2011. Lecithoceridae (Lepidoptera, Gelechioidea) of New Guinea, Part I: Onnuria Park with description of two new species. Proceedings of the Entomological Society of Washington, 113: (in press, no DOI number).

Park, K. T., and B. K. Byun. 2010. A new genus Neopectinimura (Lepidoptera, Lecithoceridae) with descriptions of five new species. Florida Entomologist, 93(2): 298-307.

Park, K. T., and C. Wu. 2010. Genus Lecithocera of Thailand Part V. with reports of nine species including six new species (Lepidoptera: Lecithoceridae). Tropical Lepidoptera Research, 20: 62-70.

Park, K. T. 2010. A new genus Woonpaikia Park, gen. nov. (Lepidoptera, Lecithoceridae), with descriptions of two new species Syntetarca Gozmány. Journal of Asia-Pacific Entomology, 13(3): 239-242.

Park, K. T. 2010. First record of Torodora species from Papua New Guinea, with a description of a new species (Lepidoptera, Lecithoceridae). Proceedings of the Entomological Society of Washington, 112: 404-409.

Park, K. T. 2010. Four new species of the subfamily Torodorinae from Thailand (Lepidoptera: Lecithoceridae). Lepidoptera Novae, 3: 145-148.

Park, K. T. 2010. Two new genera, Caveana gen. nov. and Triviola gen. nov., and two new Torodora species from Thailand (Lepidoptera, Lecithoceridae). Entomological Science, 13(2): 250.

Park, K. T. 2010. Two new species of the genus Telephata Meyrick (Lepidoptera, Lecith oceridae) from Papua New Guinea with notes on T. nitens (Daikonoff), comb. nov. Entomological Science, 14(1): 82-86.

Rota, J. 2011. Data partitioning in Bayesian Analysis: Molecular phylogenetics of metalmark moths (Lepidoptera: Choreutidae). Systematic Entomology, 36: 317-329.

Salcedo, C. 2010. Environmental elements involved in communal roosting in Heliconius butterflies (Lepidoptera: Nymphalidae). Journal of Environmental Entomology, 39(3): 907-911.

Salcedo, C. 2010. Evidence of pollen digestion at nocturnal aggregations of Heliconius sara in Costa Rica (Lepidoptera: Nymphalidae). Tropical Lepidoptera, 20(1): 35-37.

Salcedo, C. 2010. The Biology of Heliconius Night Roosting: a Foundation. Ph.D. dissertation. University of Florida./p>

Salcedo, C. 2011. Evidence of predation and disturbance events at Heliconius butterflies roosting aggregations in Panama and Costa Rica. Journal of Natural History, in press.

Salcedo, C. 2011. Pollen preference for Psychotria sp. is not learned in Heliconius erato (Lepidoptera: Nymphalidae). Journal of Insect Science. In press.

Scott, J., A. Y. Kawahara, J. Skevington, S.H. Yen, A. Sami, M. Smith, and J. Yack. 2010. The evolutionary origins of ritualized acoustic signals in caterpillars. Nature Communications, 1: 4.

Simonsen, T. J., N. Wahlberg, A. D. Warren, and F. A. H. Sperling. 2010. The evolutionary history of Boloria (Lepidoptera: Nymphalidae): phylogeny, zoogeography and larval foodplant relationships. Systematics and Biodiversity, 8(4): 513-529.

Sohn, J. C., and K. T. Park. 2010. Three new species of Lycophantis Meyrick (Lepidoptera: Yponemeutidae) from Vietnam with keys of the world species. Oriental Insects, 44: 225-233.

Sohn, J. C., S. W. Cho, and K. T. Park. 2010. New records of three Yponomeutinae (Lepidoptera: Yponomeutidae) from Korea. Korean Journal of Systematic Zoology, 26: 225-233.

Sourakov, A. 2010. Book Review: Moths of Western North America (2009). Powell J. and P. Opler. Florida Entomologist, 93(2): 330.

Sourakov, A. 2010. Natural and sexual selection in satyrine wing patterns: a complex story. News of Lepidopterists’ Society, 52(1): 6-7, 15.

Sourakov, A. 2010. New hostplant records for two noctuid species in Florida. Notes of the Association for Tropical Lepidoptera, June 2010 issue, p. 2.

Sourakov, A., and T. Paris. 2010. Fall webworm, Hyphantria cunea (Lepidoptera: Arctiidae). Featured Creatures Website, http://entomology.ifas.ufl.edu/creatures/, University of Florida. Publication Number: EENY-486.

Tennent, W. John, J. Y. Miller, and A. Rawlins. 2010. Distribution of Acrophtalmia chione Felder & Felder, 1867, with descriptions of two new subspecies from Eastern Indonesia (Lepidoptera: Nymphalidae: Satyrinae). Bulletin of the Allyn Museum, 162: 1-7.

Vershinina, A. O., and V. A. Lukhtanov. 2010. Geographical distribution of the cryptic species Agrodiaetus alcestis alcestis, A. alcestis karacetinae and A. demavendi (Lepidoptera, Lycaenidae) revealed by cytogenetic analysis. Comparative Cytogenetics, 4(1): 1-11.

Vila, R., V. A. Lukhtanov, G. Talavera, T. F. Gil, N. E. Pierce. 2010. How common are dot-like distribution ranges? Taxonomical oversplitting in Western European Agrodiaetus (Lepidoptera, Lycaenidae) revealed by chromosomal and molecular markers. Biological Journal of the Linnean Society, 101: 130-154.

Ware, J., J. Thomas, and A. Y. Kawahara. 2011. Instant symposium: When entomologists date. American Entomologist, 57(1): 46.

Warren, A. D., J. C. Whelan, and T. C. Emmel. 2010. Notes on mate-locating behavior by the skipper Phocides polybius lilea (Reakirt, [1867]) (Lepidoptera: Hesperiidae: Eudaminae). Tropical Lepidoptera Research, 20(1): 38-40.

Willmott, K. R., and J. P. W. Hall. 2010. A new species of Dynamine Hübner, [1819] from northwestern Ecuador (Lepidoptera: Nymphalidae: Biblidinae). Tropical Lepidoptera Research, 20: 23-27.

Willmott, K. R., M. Elias, and A. Sourakov. 2011. Two possible caterpillar mimicry complexes in neotropical danaine butterflies (Nymphalidae, Danainae: Danaini, Ithomiini). Annals of Entomological Society of America, 104(6):1108-1118.