MELASTOMATACEAE
of the world



Photos and illustrations of selected species
Delta description of the family
Our current understanding of relationships within the family
Current and ongoing research around the world
A comprehensive listing of publications on  the family
Melastomatologists around the world


Information on the largest tribe including a synonymy database
Various databases that include references to the Melastomataceae
"Weedy" under normal circumstances, some species are especially problematic when introduced
Notes on the cultivation of melastomes, plus suggestions of species that merit attention
Web sites worth exploring

 
 

Blakeeae morphological cladistic analysis character list
Note that this is a provisional list and will be thoroughly revised before formal publication.

1. internode quadrate (0); quadrangular (1); elliptical (2); terete (3); quadrangulo-sulcate (4); bifacial elliptic (5)
2. internode formicaria absent (0); present (1)
3. node pubescence present (0); absent (1)
4. nodal formicaria absent (0); present (1)
5. nodal swelling absent (0); somewhat (1); marked (2)
6. node width (mm) 2.5-5 (0); 5.4-10.4 (1); 12-18 (2)
7. stipules absent (0); present (1)
8. stipules coriaceous, solitary (0); thin, multi-layered (1)
9. petiole length (mm) 6-15 (0); 18-48 (1); 72-152 (2)
10. petiole base not enlarged (0); slightly (1); stem-clasping (2)
11. petiole insertion basal (0); slightly peltate (1)
12. leaf width (cm) 1 (0); 2.5-5.5 (1); 5.8-7.8 (2); 8.5-16.6 (3); 18.6-25 (4)
13. leaf length cm <9 (0); 10-17 (1); 18-26 (2); 28-34 (3)
14. foliar acarodomatia absent (0); between major veins (1); primary-axillary (2); marginal flaps (3)
15. acarodomatia laminar revolute (0); lebetiform (1); hair tuft (2); marsupiform (3); 16. pubescent-marsupiform (4); flanged (5); multiple flanges (6); absent (7)
16. plinervy (mm) 1.8-2.9 (0); 3.7-9.2 (1)
17. secondary vein pairs 2 (0); 3 (1); 4 (2); 5 (3)
18. secondary veins basal (0); pseudoplinerved (1); plinerved (2)
19. distance between tertiary veins (mm) 0.36-0.8 (0); 1.25-2 (1); 2.16-2.7 (2); 3.3-5.7 (3)
20. tertiary veins obscure (0); +/- obscure (1); distinct (2)
21. weak inter-tertiary veins present (0); absent (1)
22. lamina membranaceous (0); subcoriaceous (1); coriaceous (2)
23. margin entire (0); callose-serrulate (1); crenate-serrulate (2); dentate (3); serrate (4)
24. leaf length / width 1-1.4 (0); 1.5-1.9 (1); 2-2.3 (2); 2.4-2.9 (3); 3.9 (4)
25. leaf length / petiole length 1.6-3.1 (0); 3.5-5.4 (1); 6-7.5 (2); 10.6-13.3 (3)
26. inflorescence terminal (0); axillary (1)
27. flowers per axil usually 1 (0); 2-7 (1); 10+ (2)
28. flowers not pendant (0); pendant (1)
29. flowers regular (0); irregular by androecium (1)
30. peduncle quadrate (0); quadrangular (1); elliptic (2); terete (3); qudrangulo-sulcate (4); bifacial elliptic (5)
31. peduncle smooth (0); lenticellate (1)
32. peduncle insertion vertically stacked (0); scattered (1)
33. peduncle length (mm) 3-8.3 (0); > 16 (1)
34. peduncle length / hypanthium length 0.22-0.84 (0); 1.28-2.63 (1); 3.4-5.3 (2);>7 (3)
35. outer floral bract pairs number 0 (0); 1 (1); 2 (2)
36. outer floral bracts lenticels absent (0); present (1)
37. outer floral bracts membranaceous (0); subcoriaceous (1); coriaceous (2)
38. outer floral bracts free (0); fused (1)
39. outer floral bracts length fused / length total 0.14-0.27 (0); 0.38-0.9 (1)
40. outer floral bracts fused portion length (mm) 0.2-2.5 (0); 9-14 (1)
41. outer floral bracts basally overlapping absent (0); present (1)
42. outer floral bracts margins entire (0); undulate (1); obscurely callose denticulate (2)
43. outer floral bracts rounded (0); slightly angled (1)
44. outer floral bracts length (mm) 1.4-3 (0); 4.7-10.9 (1); 12.3-18.4 (2); 20-40 (3)
45. outer floral bracts width (mm) 2-3.4 (0); 5.7-18 (1); 20-36 (2)
46. outer floral bracts width / inner floral bracts width 0.23-0.85 (0); 0.93-1.28 (1); 1.45-2.77 (2)
47. outer floral bracts length / width 0.42-0.91 (0); 0.97-1.65 (1); 2.2-5.9 (2)
48. outer floral bracts length / inner floral bract length 0.7-0.97 (0); 1.04-1.42 (1); 1.57-2.15 (2)
49. outer floral bracts length / hypanthium length 0.3-0.7 (0); 1-1.7 (1); 2-2.8 (2); 3.5-6.8 (3)
50. outer floral bracts length / calyx lobe length <0.8 (0); 1.2-3.8 (1); 5.2-10 (2); 12-20 (3); 25-40 (4); 142 (5)
51. inner floral bracts green (0); yellow-green (1); magenta marginally (2); brown (3)
52. inner floral bracts basally overlapping absent (0); present (1)
53. inner floral bracts entire (0); toothed (1); fimbriate (2); undulate (3)
54. inner floral bracts length (mm) 0.6-2.7 (0); 3-5 (1); 8-13.5 (2); 14.1-17.4 (3); 26-33 (4)
55. inner floral bracts width (mm) 0.22-1.7 (0); 3.2-5.8 (1); 7.4-9.9 (2); 12.5-24 (3); 32-40 (4)
56. inner floral bracts free (0); fused (1)
57. inner floral bracts fused portion length (mm) <0.5 (0); >10 (1)
58. inner floral bracts rounded (0); slightly angled (1); distinctly angled (2)
59. inner floral bracts nerves 1 (0); 3 (1); 5 (2); 7 (3); 9 (4); 11 (5); many (6)
60. inner floral bracts length / hypanthium length 0.18-0.58 (0); 0.87-2.32 (1); 3.39-5.3 (2)
61. inner floral bracts length / calyx lobe length 0.3-1.5 (0); 1.9-3.9 (1); 4.7-7.5 (2); 9.2-19 (3); 34-37 (4); 71(5)
62. hypanthium pubescence absent (0); mainly basal (1); mainly apical (2); throughout (3)
63. hypanthium lenticels absent (0); present (1)
64. hypanthium ribbed (0); veined (1); obscurely angled (2); terete (3)
65. hypanthium length (mm) 1.8-2.9 (0); 3.8-5.2 (1); 6.1-8.4 (2); 9.6-13 (3); >17 (4)
66. hypanthium width (mm) 1.6-2.3 (0); 3-5.4 (1); 6.2-8.4 (2); 9.4-12.7 (3)
67. hypanthium length / width 0.6-0.95 (0); 1.02-1.74 (1)
68. hypanthium length / inner hypanthium length 0.4-1.8 (0); 1.9-5 (1); >8 (2)
69. external calyx tooth absent (0); present (1)
70. calyx lobes 4 (0); 5 (1); 6 (2); varying (3)
71. calyx lobe veins arising from center (0); sides (1)
72. calyx lobes imbricate no (0); yes (1)
calyx lobes widest at base (0); above (1)
73. calyx lobes subtended by a prominent ridge no (0); yes (1)
74. calyx lobe length (mm) 0.2-0.8 (0); 1-5 (1); 6.2-23.7 (2)
75. calyx lobe length + calyx tube length (mm) 0.8-2.8 (0); 3.2-4.5 (1); >/=5.9 (2)
76. calyx lobe length / width 0.08-0.84 (0); 1.41-1.82 (1)
77. calyx lobe adaxial callose thickening absent (0); present (1)
78. calyx median thickening absent (0); present (1)
79. calyx tube prominent vein network absent (0); present (1)
80. calyx tube length (mm) 0 (0); 0.36-1.34 (1); 1.66-4.73 (2); >/=5.4 (3)
81. calyx tube deflexed below torus absent (0); present (1)
82. calyx abaxial pubescence absent (0); apical 1/2 (1); median (2); throughout (3); basal (4)
83. calyx adaxial pubescence absent (0); apical 1/2 (1); median (2); throughout (3)
84. calyx foliose apical appendage absent (0); present (1)
85. petal number 4 (0); 5 (1); 6 (2)
86. petal surface smooth (0); tuberculate (1)
87. petal pubescence marginal (0); absent (1); adaxial (2)
88. corolla rotate to ascending (0); pseudocampanulate (1)
89. petal insertion narrow (0); somewhat broad (1); broad (2)
90. petals white to pink-tinged (0); pink (1); lavender to purple (2); red (3); green(4); cream (5)
91. petal length (mm) 3.2-11.7 (0); 13.8-21.7 (1); 25-55 (2)
92. petal width (mm) 2-4.6 (0); 5.7-15 (1); 17-29 (2)
93. petal length / width 0.85-1.08 (0); 1.17-1.47 (1); 1.6-3.6 (2)
94. stamens 8 (0); 10 (1); 12 (2)
95. stamens isomorphic (0); dimorphic (1)
96. anthers free (0); laterally connate (1)
97. filament adaxial keel absent (0); present (1)
98. filament lateral flanges absent (0); present (1)
99. filaments terete (0); +/- flat (1); strap-like (2); triangular (3); D-shaped (4)
100. filament length (mm) 2-3.6 (0); 4.2-5.8 (1); 6.6-8.7 (2); 9.1-10.9 (3); >17 (4)
101. anther thecae surface smooth (0); slightly rugose (1)
102. theca base sagitate (0); decurrent (1); rounded (2)
103. thecae yellow (0); cream (1); white (2); purple (3)
104. anther length (mm) 2.4-3.7 (0); 4.7-8.8 (1); 9.8-12.5 (2)
105. anther width (mm) 0.4-0.6 (0); 1.2-1.4 (1); 1.8-2.8 (2); 3-3.5 (3); >4 (4)
106. anther width 0.03-0.24 (0); 0.3-0.64 (1)
107. anther length / width 1.02-2.4 (0); 3.03-3.29 (1); 4-4.5 (2); 5.3-9.4 (3)
108. connective sculpturing absent (0); present (1)
109. connective decurrent along filament absent (0); present (1)
110. pores well separated (0); approximate (1); confluent (2)
111. pores circular (0); elliptic (1); teardrop (2); triangular (3); diamond (4); semicircular 112. (5)
113. pore length / width 0.33-0.54 (0); 0.63-0.91 (1); 1-1.57 (2); 1.66-2.57 (3)
114. anther pore length (mm) 0.04-0.4 (0); 0.6 (1)
115. anther pore(s) dorsal (0); apical (1); ventral (2)
116. thecae dorsal thickening absent (0); slight (1); pronounced (2)
117. connective ventral (0); dorsal (1)
118. connective invagination absent (0); dorsal (1); between filament-tip (2)
119. connective callose (0); bilobed callose (1); triangular (2); elongated triangular (3); 120. two long ridges (4); flattened-caudate (5); caudate (6)
121. stamen length / petal length 0.34-0.57 (0); 0.61-0.94 (1); 1.02-2.2 (2)
122. stamen length / style length 0.54-0.82 (0); 0.86-1 (1); 1.06-1.33 (2); 1.4-2.2 (3)
123. stamen length / hypanthium length 1-1.75 (0); 1.96-2.51 (1); 2.88-3.79 (2)
124. filament length / anther length 0.31-0.98 (0); 1.06-1.75 (1)
125. torus glabrous (0); pubescent (1)
126. inner hypanthium ridges absent (0); present (1)
127. inner hypanthium pubescence absent (0); present (1)
128. inner hypanthium thickening absent (0); present (1)
129. inner hypanthium length (mm) 0.4-1.3 (0); 1.5-3.7 (1); 4.8-5.1 (2)
130. style pubescence absent (0); present (1)
131. style straight (0); shallowly curved (1); distinctly curved (2)
132. stigma truncate (0); slightly expanded (1); rounded-capitate (2); elongated-capitate (3)
133. ovary pubescence apical 1/4 (0); apical 1/2 (1); absent (2)
134. ovary inferior (0); 4/5 inferior (1); 3/5 inferior (2); 1/2 inferior (3); 2/5 inferior (4); 1/5 inferior (5); superior (6)
135.locule number 2 (0); 3 (1); 4 (2); 5 (3); 6 (4)
136.locule width (mm) 0.8-2.25 (0); 3.2-4.4 (1); 5.1-9.5 (2)
137.locule length / width 0.36-0.55 (0); 0.6-0.89 (1); 0.96-1.65 (2)
138.locule length (mm) 0.9-1.3 (0); 2-2.7 (1); 2.9-4.4 (2); 7 (3); 8 (4)
placentation intruded axile (0); axile (1)
139. ovary apex setose (0); glabrous (1); glandular (2); ciliate (3); pubescent (4)
140. style + stigma length (mm) 4.4-7.5 (0); 9-16.7 (1); 17.2-25.5 (2); 37 (3)
141. stigma width (mm) 0.12-1 (0); 1-3.9 (1)
142. stigma length (mm) 0.06-1.2 (0); 2-4.2 (1)
143. ovary apical appendages absent (0); simple (1); bifid (2)
144. ovary apex ridges absent (0); obscure (1); prominent (2)
145. ovary apex concave (0); flat (1); round (2); cone (3); cyclindrical (4)
146. fruit a capsule (0); berry (1)


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Page developed and maintained by Darin S. Penneys.
Please send your comments and suggestions to the webmaster: dpenneys@ufl.edu
Page last updated: 8 January, 2005