Selected Publications by PIRE Participants


Steadman, D.W. & B.J. MacFadden. In press. An early Miocene eagle (Aves: Accipitridae) from Panama. Journal of Paleontology.

De Baets, K., Antonelli, A., Donoghue, P. Tectonic blocks and molecular clocks. (2016, in press) Philosophical Transactions B.

Bacon, C.D., Molnar, P., Antonelli, A., Crawford, A.J., Montes, C., Vallejo-Pareja, M.C. 2016. Quaternary glaciation and the Great American Biotic Interchange. Geology 44(5): 375–378.

Ramírez, D. A., Foster, D. A., Min, K., Montes, C., Cardona, A., Sadove, G. 2016. Exhumation of the Panama basement complex and basins: Implications for the closure of the Central American seaway. Geochemistry, Geophysics, Geosystems. doi:10.1002/2016GC006289.

Bloch, J. I., Woodruff, E. D., Wood, A. R., Rincon, A. F., Harrington, A. R., Morgan, G. S., Foster, D. A., Montes, C., Jaramillo, C. A., Jud, N. A., Jones, D. S., and MacFadden, B. J. 2016. First North American Fossil Monkey and Early Miocene Tropical Biotic Interchange. Nature. doi:10.1038/nature17415

Pimiento, C., MacFadden, B. J., Clements, C. F., Varela, S., Jaramillos, C., Velez-Juarbe, J., Silliman, B. R. 2016. Geographical distribution patterns of Carcharocles megalodon over time reveal clues about extinction mechanisms. Journal of Biogeography. doi: 10.1111/jbi.12754

Bokor, J., Broo, J., Mahoney, J. 2016. Using Fossil Teeth to Study the Evolution of Horses in Response to a Changing Climate. The American Biology Teacher 78(2): 166-169. doi: 10.1525/abt.2016.78.2.166

Jud, N. A., Nelson, C. W., Herrera, F. 2016. Fruits and wood of Parinari from the early Miocene of Panama and the fossil record of Chrysobalanaceae. American Journal of Botany. doi: 10.3732/ajb.1500425


Bacon, C. D., Silvestro, D., Jaramillo, C. A., Smith, B. T., Chakrabarty, P., and Antonelli, A. 2015. Reply to Lessios and Marko et al.: Early and progressive migration across the Isthmus of Panama is robust to missing data and biases. Proceedings of the National Academy of Sciences. doi:10.1073/pnas.1515451112

MacFadden, B. J., Symister, C., Cannarozzi, N., Pimiento, C., DeGracia, C. 2015. Comparative diagenesis and rare earth element variation in Miocene invertebrate and vertebrate fossils from Panama. The Journal of Geology. doi:10.1086/684006

Pyenson, N. D., Vélez-Juarbe, J., Gutstein, C. S., Little, H., Vigil, D., O'Dea, A. 2015. Isthminia panamensis, a new fossil inioid (Mammalia, Cetacea) from the Chagres Formation of Panama and the evolution of ‘river dolphins’ in the Americas. PeerJ. doi:10.7717/peerj.1227

Rincon, A. F., Bloch, J. I., MacFadden, B. J., Jaramillo, C. A. 2015. New early Miocene protoceratids (Mammalia, Artiodactyla) from Panama. Journal of Vertebrate Paleontology. doi:10.1080/02724634.2015.970688. New early Miocene protoceratids (Mammalia, Artiodactyla) from Panama.

Hendy, A.J.W., Jones, D.S., Moreno, F., Zapata V., and Jaramillo, C. 2015. Neogene molluscs, shallow marine paleoenvironments, and chronostratigraphy of the Guajira Peninsula, Colombia. Swiss Journal of Palaeontology 134(1): 45-75. doi: 10.1007/s13358-015-0074-1

Moreno, F., Hendy, A.J.W., Quiroz, L., Hoyos, N., Jones, D.S., Zapata, V., Zapata, S., Ballen, G.A., Cadena, E., Cardenas, A.L., Carrillo-Briceño, J.D. Carrillo, J.D., Delgado-Sierra, D. Escobar, J., Martinez, J.I., Martinez, C. Montes, C., Moreno, J., Perez, N.,Sanchez, R., Suarez, C.,Vallejo-Pareja, M.C., and Jaramillo, C. 2015. Revised stratigraphy of Neogene strata in the Cocinetas Basin, La Guajira, Colombia. Swiss Journal of Palaeontology 134(1): 5-43. doi: 10.1007/s13358-015-0071-4.

Pimiento, C., Balk, M. A. 2015. Body-size trends of the extinct giant shark Carcharocles megalodon: a deep-time perspective on marine apex predators. Paleobiology, 41: 479-490. doi:10.1017/pab.2015.16.

Velez-Juarbe, J., Wood, A. R., De Gracia, C., Hendy, A. J. W. 2015. Evolutionary Patterns among Living and Fossil Kogiid Sperm Whales: Evidence from the Neogene of Central America. PLoS ONE 10(4): e0123909. doi:10.1371/journal.pone.0123909

Wood, A. R., Ridgwell, N. M. 2015. The first Central American chalicothere (Mammalia, Perissodactyla) and the paleobiogeographic implications for small-bodied schizotheriines. Journal of Vertebrate Paleontology. DOI: 10.1080/02724634.2014.923893

Klompmaker, A.A., R.W. Portell, S.E. Lad, and M. Kowalewski. 2015. The fossil record of drilling predation on barnacles. Palaeogeography, Palaeoclimatology, Palaeoecology 426: 95–111. doi:10.1016/j.palaeo.2015.02.035

Bacon, C. D., Silvestro, D., Jaramillo, C., Tilston Smith, B. T., Chakrabarty, P., Antonelli, A. 2015. Biological evidence supports an early and complex emergence of the Isthmus of Panama. Proceedings of the National Academy of Sciences, published ahead of print. doi:10.1073/pnas.1423853112.

Klompmaker, A. A., Hyžný, M., Portell, R. W. 2015. Growth, inter- and intraspecific variation, palaeobiogeography, taphonomy and systematics of the Cenozoic ghost shrimp Glypturus. Journal of Systematic Palaeontology. DOI:10.1080/14772019.2015.1009505.

Carrillo-Briceño, J., de Gracia, C., Pimiento, C., Aguilera, O., Kindlimann, R., Santamarin, P., and Jaramillo, C. 2015. A new Late Miocene Chondrichthyan Assemblage from the Chagres Formation, Panama. Journal of South American Earth Sciences, v. 60, p. 56-70.

Carrillo, J. D., Forasiepi, A., Jaramillo, C., and Sánchez-Villagra, M. R. 2015. Neotropical mammal diversity and the Great American Biotic Interchange: spatial and temporal variation in South America's fossil record. Frontiers in Genetics, v. 5, p. 451.



Herrera, F., Manchester, S. R., Koll, R., and Jaramillo, C. 2014. Fruits of Oreomunnea (Juglandaceae) in the early Miocene of Panama. Pages 124-133 in W. D. Stevens, O. M. Montiel, and P. Raven, editors. Paleobotany and Biogeography: A Festschrift for Alan Graham in His 80th Year. Missouri Botanical Garden Press, St Louis, MO. Fruits of Oreomunnea (Juglandaceae) in the early Miocene of Panama

Herrera, F., Manchester, S., Vélez-Juarbe, J., Jaramillo, C. 2014. Phytogeographic History of the Humiriaceae (Part 2). International Journal of Plant Sciences 175(7): 828-840. doi:10.1086/676818

Vasquez, S., Pimiento, C. 2014. Sharks and Rays from the Tonosi Formation (Eocene of Panamá). Revista Geológica de América Central 51: 165-169. (in Spanish)

Pimiento, C., Clements, C.F. 2014. When Did Carcharocles megalodon Become Extinct? A New Analysis of the Fossil Record. PLoS ONE 9(10).

Sepulchre, P., Arsouze, T., Donnadieu, Y., Dutay, J.-C., Jaramillo, C., Le Bras, J., Martin, E., Montes, C., and Waite, A. J., 2014. Consequences of shoaling of the Central American Seaway determined from modeling Nd isotopes. Paleoceanography 29.

MacFadden, B. J., Bloch, J. I., Evans, H., Foster, D. A., Morgan, G. S., Rincon, A., Wood, A. R. 2014. Temporal Calibration and Biochronology of the Centenario Fauna, Early Miocene of Panama. Journal of Geology 122(2): 113-135. doi:10.1086/675244


Hastings, A., Bloch, J., Jaramillo, C., Rincon, A., MacFadden, B. 2013. Systematics and Biogeography of Crocodylians from the Miocene of Panama. Journal of Vertebrate Paleontology 33: 239-263.

Hendy, A. J. W. 2013. Spatial and stratigraphic variation of marine paleoenvironments in the middle-upper Miocene Gatun Formation, Isthmus of Panama. Palaios 28: 210-227.

Jaramillo, C. and Cardenas, A. 2013 Global Warming and Neotropical Rainforests: A historical perspective. Annual Reviews of Earth and Planetary Sciences 41: 741-766.

Pimiento C., Gonzalez G., Hendy A., Jaramillo C., MacFadden B.J., Montes C., Suarez S.C., Shippritt M. 2013. Early Miocene chondrichthyans from the Culebra Formation, Panama: A window into marine vertebrate faunas before closure of the Central American Seaway. Journal of South American Earth Sciences 42: 159-170

Pimiento C., Gonzalez G., Ehret D.J., MacFadden B.J., Hendy A., Jaramillo C. (2013) Sharks and Rays (Chondrichthyes, Elasmobranchii) from the late Miocene Gatun Formation of Panama. Journal of Paleontology 87: 755-774.

Rincon, A. F., J. I. Bloch, B. J. MacFadden, and C. A. Jaramillo. 2013. First Central American record of Anthracotheriidae (Mammalia, Bothriodontinae) from the early Miocene of Panama, Journal of Vertebrate Paleontology, 33:2, 421–433.


Cadena, E., Bourque, J., Rincon, A., Bloch, J.I., Jaramillo, C., and MacFadden, B. 2012 New Turtles (Chelonia) from the Late Eocene through Late Miocene of the Panama Canal Basin. Journal of Paleontology 86: 539-557.

Head, J., Rincon, A., Suarez, C., Montes, C., and Jaramillo, C. 2012 Fossil evidence for earliest Neogene American faunal interchange: Boa (Serpentes, Boinae) from the early Miocene of Panama. Journal of Vertebrate Paleontology 32: 1328-1334.

Herrera, F., Manchester, S., and Jaramillo, C. 2012 Permineralized fruits from the late Eocene of Panama give clues of the composition of forests established early in the uplift of Central America. Review of Palaeobotany and Palynology 175:10-24.

Jaramillo, C. 2012 Historia Geológica del Bosque Húmedo Neotropical. Revista de la Academia Colombiana de Ciencias Exactas, Físicas y Naturales 36: 59-80.

MacFadden, B. J., L. H. Oviedo, G. M. Seymour, and S. Ellis. 2012. Fossil horses, orthogenesis, and communicating evolution in museums. Evo Edu Outreach 5:29-37.

Montes, C., Bayona, G., Cardona, A., Buchs, D.M., Silva, C.A., Moron, S.E., Hoyos, N., Ramirez, D.A., Jaramillo, C., and Valencia, V. 2012 Arc-Continent Collision and Orocline Formation: Closing of the Central American Seaway. Journal of Geophysical Research 117, B04105, doi:10.1029/2011JB008959.

Montes, C., Cardona, A., MacFadden, R., Moron, S.E., Silva, C.A., Restrepo-Moreno, S., Ramirez, D.A., Wilson, J., Farris, D., Bayona, G.A., Jaramillo, C., Valencia, V., and Flores, J.A. 2012 Evidence for middle Eocene and younger emergence in Central Panama: implications for Isthmus closure. Geological Society of America Bulletin, DOI: 10.1130/B30528.1.

Morgan, G. S. and N. J. Czaplewski. 2012. "Evolutionary history of the Neotropical Chiroptera: The fossil record", In Evolutionary history of bats: Fossils, molecules, and morphology, G. F. Gunnell and N. B. Simmons, editors. Cambridge University Press, Cambridge.

Rincon, A., Bloch, J.I., Suarez, C., MacFadden, B.J., and Jaramillo, C. 2012 New Floridatragulines (Mammalia, Camelidae) From The Early Miocene Las Cascadas Formation, Panama. Journal of Vertebrate Paleontology, 32: 456-475.

Whattam, S.A, Montes, C., McFadden, R., Cardona, A., Ramirez, D. and Valencia, V. 2012. "Age and origin of earliest adakitic-like magmatism in Panama: implications for the tectonic evolution of the Panamanian magmatic arc system", Lithos 142-143: 226-244.

Hendy, Austin. 2012. Key to the Fossils of the Gatun Formations. Field_Key_Fossils_Gatun.pdf


Farris, D.W., Jaramillo, C., Bayona, G., Restrepo-Moreno, S., Montes, C., Cardona, A., Mora, A., Speakman, R.J., Glasscock, M.D., Reiners, P., Valencia, V. 2011 Fracturing of the Panamanian Isthmus during initial collision with South America. Geology 39: 1007-1010.

Pimiento C. and Pringle R. 2011. Fossil sharks: Learning from and about the past. Science Scope 34 (6) 52-59. 


Herrera, F., Manchester, S., Jaramillo, C., MacFadden, B., and Silva, S. 2010 Phytogeographic History and Phylogeny of the Humiriaceae. International Journal of Plant Sciences 171: 392-408.

MacFadden, B.J., Kirby, M.X., Rincon, A., Montes, C., Moron, S., Strong, N., and Jaramillo, C. 2010 Extinct Peccary "Cynorca" occidentale (Tayasuidae) from the Miocene of Panama and Correlations to North America. Journal of Vertebrate Paleontology 84: 288-298.

Pimiento C., Ehret D.J., MacFadden B.J., Hubbell G. 2010. Ancient Nursery Area for the Extinct Giant Shark Megalodon from the Miocene of Panama. PLoS ONE 5(5): e10552, DOI: 10.1371/journal.pone.0010552

Uhen, M., A. Coates, C. Jaramillo, C. Montes, C. Pimeinto, A. Rincon, N. Strong, and J. Velez-Juarbe. 2010. Marine mammals from the Miocene ofPanama. Journal of South American Earth Sciences, p. 167, vol. 30.

Vega, F., Nyborg, T., Kovalchuk, G., Etayo, F., Luque, J., Rojas-Briceño, A., Patarroyo, P., Porras-Muzquiz, H., Armstrong, A., Bemudez, H., and Garibay, L. 2010. On some Panamerican Cretaceous Crabs (Decapoda: Raninoida). Boletín de la Sociedad Geológica Méxicana 62: 263-279.


MacFadden, B. J. 2009. Three-toed browsing horse Anchitherium (Equidae) from the Miocene of Panama. Journal of Paleontology 83(3):489-492.

Additional Publications